12. Tylodinus variabilis Luna-Cozar, sp. nov.

(Figures 2, 50–51, 107, 132)

Diagnosis. Length male, 3.4–4.0 mm, female, 3.2–4.0 mm. Width male, 1.6–1.7 mm, female 1.5–2.0 mm. As for T. buchanani with the follow exceptions: body clothed with short scales, light scales on vertex and pronotum frequently whitish, but in some specimens light brown. Head with overlapping scales on vertex; front moderately concave. Elytra with basal margin straight (Figure 2), stria 10 complete, surface granulate, tubercles similar to T. buchanani . Metasternal tooth absent. Abdomen squamous, scales setigerous, ventrite 1 convex; ventrite 2 longer than 3–4 combined, convex. Hind femora length almost reaching elytral apex, front tibiae sinuate, middle and hind tibiae subparallel, premucro very small, hind tibiae with apical ¼ curved. Male genitalia (Fig. 107) in lateral view slender; in ventral view with lateral sides concave, subparallel; apical process in lateral view weakly curved. Female genitalia with tergite eight trapezoidal, apex slightly curved, sternite eight oblong, median membranous area subapical; spermatheca shallowly curved, ramus long and slender, nodulus slender.

Geographic distribution. México (Chiapas).

Habitat and elevation. Specimens were collected from leaf litter in high elevation cloud forest between 2300–2750 m.

Derivation of specific name. Name derived from the Latin word variabilis = variable.

Material examined. Total 95 males, 90 females. Holotype male (CMNC): México: Chiapas, San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16°44'33''N, 92°41'38.4''W), 15.ix.1991, coll. R . S. Anderson. Paratypes: México: Chiapas, Huixtán, 15 km E San Cristóbal, elev. 2500 m (16°44'49.6''N, 92°29'24.22''W), 28.v.2008, coll. LLAMA (Wa-A-05-1-24) — 1 female (CMNC); same locality (16.74717°N 92.49017°W), 28.v.2008, coll. LLAMA (Wa-A-05-1-48) — 1 female (CMNC); same locality (16.74698°N 92.49013°W), 28.v.2008, coll. LLAMA (Wa-A-05-1-16) — 1 male (CMNC); Bazom, elev. 2450 m (16°44'18.96''N, 92°29'18.24''W), 9.vii.2003, coll. R . S. Anderson — 1 male (CMNC); San Cristobal de las Casas Airport, elev. 2350 m (16°41'25.08''N, 92°31'48''W), 25.vii.2005, coll. R . S. Anderson — 1 female (CMNC); San Cristóbal de las Casas, Cerro Huitepec (pico) 5 km W San Cristóbal, elev. 2750 m (16°44'33''N, 92°41'38.4''W), 15.ix.1991, coll. R . S. Anderson — 6 males, 6 females (CMNC); same locality, 18.ix.1991, coll. R . S. Anderson — 4 males, 2 females (CMNC); Cerro Huitepec 5 km W San Cristóbal, elev. 2650 m (16°44'37''N, 92°41'23.3''W), 17.ix.1992, coll. R . S. Anderson — 2 males, 5 females (BMNH, CMNC); Corazón de María, elev. 2449 m (16°41'7.4''N, 92°32'11.4''W), 8.vi.2006, coll. J. L. Cozar — 1 male (ECOS); Huitepec, elev. 2466 m (16°44'54.2''N, 92°41'6.4''W), 5.v.2006, coll. J. L. Cozar — 2 males, 2 females (ECOS); same locality, 15.vi.2006, coll. J. L. Cozar — 1 male (ECOS); same locality, 3.vii.2006, coll. J. L. Cozar — 2 males (ECOS); same locality (16.75181°N 92.68270°W), elev. 2480 m, 29.v.2008, coll. LLAMA (Wa-A-05-2-01) — 2 males, 1 female (CWOB); same locality (16.75104°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-18) — 1 male, 1 female (CMNC); same locality (16.75109°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-17) — 1 female (CMNC); same locality (16.75131°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-12) — 1 female (CMNC); same locality (16.75176°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-02) — 3 males, 1 female (CMNC); same locality (16.75163°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-05) — 3 males, 5 females (CMNC, ECOS); same locality (16.74689°N 92.48990°W), 29.v.2008, coll. LLAMA (Wa-A-05-1-01) — 1 male (CMNC); same locality (16.75018°N 92.68280°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-39) — 2 males (CMNC); same locality (16.75018°N 92.68284°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-40) — 1 male (CMNC); Huitepec Ocotal, Cerro La Cruz, elev. 2970 m (16°44'31.2''N, 92°41'37.7''W), 12.vi.2006, coll. J. L. Cozar — 3 males, 2 females (ECOS); Huitepec, 5 km W San Cristóbal, elev. 2750 m (16°44'33''N, 92°41'38.4''W), 14-25.ix.1992, coll. R . S. Anderson — 2 females (CMNC); same locality, 14.ix.1992, coll. R . S. Anderson — 5 males, 2 females (CMNC); Huitepec, S. Cristóbal, elev. 2480 m (16.75041°N 92.68270°W), 29.v.2001, coll. LLAMA (Wa-A-05-2-32) — 1 male, 1 female (CMNC); same locality (16.75032°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-34) — 2 females (ECOS); same locality (16.75045°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-31) — 1 male, 2 females (CMNC); same locality (16.75163°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-05) — 1 female (CMNC); same locality (16.75050°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-30) — 1 male, 1 female (ECOS); same locality (16.75032°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-34) — 1 female (CMNC); same locality (16.75077°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-24) — 1 female (CMNC); same locality (16.75018°N 92.68298°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-43) — 1 female (CMNC); same locality (16.75018°N 92.68303°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-44) — 1 female (CMNC); same locality (16.75018°N 92.68308°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-45) — 1 male (CWOB); same locality (16.75018°N 92.68317°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-47) — 1 male (CMNC); same locality (16.75018°N 92.68327°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-49) — 1 female (CMNC); same locality (16.75054°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-29) — 1 male (CMNC); same locality (16.75082°N 92.68270°W), 29.v.2008, coll. LLAMA (Wa-A-05-2-23) — 1 male (CMNC); Los Alcanfores, elev. 2450 m (16°44'12.47''N, 92°40'46.93''W), 25.iv.2006, coll. M. Girón — 1 male (ECOS); Reserva Huitepec, elev. 2400 m (16°44'12.5''N, 92°40'46.9''W), 18.xi.1993, coll. R . Jones — 4 males, 1 female (ECOS, UAQM); same locality, 18.xi.1993, coll. R . Jones — 1 male (ECOS); same locality, 16.ii.1994, coll. R . Jones — 1 male (ECOS); same locality, 16.ii.1994, coll. R . Jones — 2 males, 2 females (ECOS); same locality, 7.iv.1994, coll. R . Jones — 1 females (ECOS); same locality, 27.v.1994, coll. B. Gómez-G y O. L. Gómez-N— 2 males (ECOS); same locality, 25.viii.1994, coll. B. Gómez-G— 2 males (ECOS); same locality, 26.x.1994, coll. B. Gómez-G— 1 male, 1 female (ECOS); same locality, 24.i.1995, coll. R . Jones y B. Gómez-G— 1 female (ECOS); same locality, 24.i.1995, coll. R . Jones — 1 male (ECOS); same locality, 24.i.1995, coll. R . Jones-W y B. Gómez-G— 1 male (ECOS); same locality, 27.iv.1995, coll. B. Gómez-G y M. Girón-I— 1 female (ECOS); same locality, 14.vi.1995, coll. B. Gómez- G, M. Girón-I y A. Méndoza-R— 1 male (ECOS); same locality, 27.vi.1995, coll. B. Gómez-G y M. Girón-I— 1 female (ECOS); same locality, 5.vii.1995, coll. M. Girón — 1 male (UAQM); same locality, 5.vii.1995, coll. R . Jones — 1 male (UAQM); same locality, 10.vii.1995, coll. R . Jones — 1 female (UAQM); same locality, 25.vii.1995, coll. R . W. Jones — 4 males, 3 females (ECOS, UAQM); same locality, 1.viii.1995, coll. A. Mendoza y M. Girón — 1 male (UAQM); same locality, 15.viii.1995, coll. M. Girón — 1 female (UAQM); same locality, 3.xi.1995, coll. B. Gómez y M. Girón — 1 male, 2 females (UAQM); same locality, 14.xi.1995, coll. B. Gómez, M. Girón — 1 male, 1 female (UAQM); same locality, 5.xii.1995, coll. B. Gómez y M. Girón — 1 male, 2 females (UAQM); same locality, 6.ii.1996, coll. L. Ruiz-M y M. Girón-I— 3 males, 4 females (ECOS, UAQM); same locality, 14.ii.1996, coll. L. Ruiz-M y M. Girón-I— 1 male (ECOS); same locality, 14.ii.1996, coll. L. Ruiz-M y M. Girón-I— 2 males, 2 females (UAQM); same locality, 7.vii.2003, coll. R . S. Anderson — 1 female (CMNC); same locality, 12.ii.2004, coll. J. L. Cozar, M. Girón — 1 female (ECOS); same locality, 9.vi.2004, coll. J. L. Cozar, M. Girón — 1 male, 1 female (ECOS); same locality, 11.vi.2004, coll. J. L. Cozar — 1 female (ECOS); same locality, 2.vii.2004, coll. J. L. Cozar — 1 female (ECOS); same locality, 12.vii.2004, coll. J. L. Cozar — 1 female (ECOS); same locality, 13.vii.2004, coll. M. Girón — 1 male (ECOS); same locality, 15.vii.2005, coll. R . S. Anderson — 1 male (CMNC); same locality, 9.vi.2006, coll. U. Caballero — 1 female (ECOS); same locality, 12.vi.2006, coll. J. L. Cozar — 1 female (ECOS); same locality, 17.vi.2006, coll. U. Caballero — 3 females (ECOS); same locality, 11.vii.2007, coll. J. Longino — 1 female (CMNC); same locality, 9.vii.2007, coll. R . S. Anderson — 4 males, 4 females (CMNC); same locality, 9.vii.2007, coll. M. G. Branstetter — 2 females, 1 male (CMNC); same locality, 11.vii.2007, coll. R . S. Anderson — 3 males, 2 females (CMNC); Zoológico, elev. 2379 m (16°43'5.5''N, 92°42'9.7''W), 6.vi.2006, coll. J. L. Cozar — 1 male (ECOS); San Juan Chamula, Cerro Tzontehuitz, elev. 2800 m (16°49'10.2''N, 92°35'3.8''W), 15.vii.1994, coll. R . Jones — 1 male (UAQM) .