Lesteva (s.str.) pubescens Mannerheim, 1830

(Figs. 31, 38–46)

Lesteva pubescens Mannerheim, 1830; Luze 1903: 196, Johansen 1914: 589, Portevin 1929: 437, Palm 1948: 120, Lohse 1956: 55, 1964: 60, Tóth 1982: 100

Lesteva (Lestevella) pubescens: Jeannel & Jarrige 1949: 315

Lesteva (s.str.) pubescens: Zanetti 1987: 367, 2012: 72, Tronquet 2006: 33

Lesteva pubescens var. subaptera Mulsant and Rey, 1880: 69

Lesteva pubescens reyi Jarrige, 1968

Lesteva fageli Lohse, 1960: 6 syn.n.

Type material examined. Paratypes of Lesteva fageli Lohse, 1960: 1 ♀: ‘ Espagne: Estremadura | Guadalupe | (canal scierie) | V—1958, G. Fagel’ <printed>, ‘fageli sp. n. [handwritten] | det. Dr. G. A. Lohse’ <printed>, ‘Para- | type’ <red, printed>, ‘Coll. | G.A. Lohse | MHNG-1994’ <printed>, ‘ PARATYPE | Lesteva | fageli sp. n. | Det. Lohse 1960 ’ <yellow, printed>, ‘MHNG | ENTO | 00087326’ <printed> (MHNG); 1 ♂ [dissected]: same labels, but: ‘MHNG | ENTO | 00087327’ <printed> (MHNG); 1 ♂: same labels, but: ‘MHNG | ENTO | 00087328’ <printed> (MHNG). All paratypes with additional printed label: ‘ Lesteva (str.) pubescens Mannerheim, 1830 | Shavrin A.V. det. 2021’.

Material examined. PORTUGAL: CASTELO BRANCO: 3 ♂♂: Serra de Estrela. H. Franz leg. (NHMW); 1 ♂: Estrela (MHNG); 3 ♀♀: same data. H. Franz leg. (MHNG); SPAIN: GALICIA: 1 ♂, 1 ♀: Pontevedra Galice. 18- 30.07.1959. H. Coiffait leg. (MHNH) ; 1 ♂: Lugo, Linares. 1000 m a.s.l. 27.05.1996. W. Starke leg. (cA) ; ASTURIAS: 1 ♂ [identified by G.A. Lohse as fageli] , 2 ♀♀: ‘ Hispania la Granja’ (BMNH) ; SORIA: 2 ♂♂: Sierra de Urbión, forest W of Santa Inés. 1700 m a.s.l. 23.07.1996. D. Wrase & J.P. Zaballos leg. (cSch) ; CANTABRIA: 1 ♀: Espinama, 27.07.1972 (MHNG) ; 1 ♂, 4 ♀♀: same locality, Valle de Salvoron. 30.07.1972. (MHNG) ; 3 ♂♂: Poblagion (BMNH); CASTILLA Y LEÓN: 1 ♀: Sierra de Gredos. 16.07.1959. H. Coiffait leg. (MHNH) ; 1 ♂: Sierra de Gredos, Novacepeda. 01.06.1984. (cZan) ; 3 ♀♀: Puerto de Béjar. 800-1400 m a.s.l. 05-06.1957. G. Fagel leg. (MHNG) ; ARAGÓN: 1 ♂: Teruel (IRSN); CATALUñA: 1 ♂: Col de Aras ( Span. Pyren.). 31.05.1971. G. Lohse leg. (MHNG) ; 1 ♂, 4 ♀♀: Barcelona (MHNG); 1 ♂: Sierra de Montseny, Tordera River, 5 km E San-Bernat. 02- 12.5.1981. P. Zwick leg. (MHNG) ; 4 ♂, 6 ♀♀: Montseny (MHNG); MADRID: 1 ♀: Escorial (NMPC); 1 ♂: same locality. 30.07.1879. L. Bleuse leg. (NMPC); 1 ♂: ‘ Guadarrama | Eppelsh. ’, ‘ L. pandellei J. Bohač det. 1980’ (NMPC) ; 2 ♂♂, 2 ♀♀: Sistema Central Mts., Pinilla de Valle, 40°91’N -3°82’W. 28.04.2014. A.V. Shavrin & A. Anichtchenko leg. (cSh) ; 2 ♀♀: Puerto de la Morcuera. 11.10.2004. de la Rosa leg. (cSh) ; 1 ♂: Sierra de Estrela, Manteigas. 1000-1100 m a.s.l. H. Franz leg. (MHNH) ; CASTILLA LA MANCHA: 2 ♂♂: Sierra de Alcaraz, 15 km NNE Riópar, 38°34’01N 02°20’03W 1120 m a.s.l. 10.04.2003. P. Wunderle leg. (cW) ; VALENCIA: 1 ex.: ‘ Buñol Valencia Moróder’ (MHNH); ANDALUSÍA: 5 ♂♂, 7 ♀♀: Despenaperros, Arroyo de los Arcos, 38°38‘N - 3°51‘W. 01- 02.05.2014. A.V. Shavrin leg. (cSh) ; 1 ♂: ‘ Pnte. Las Herrerias S. Cazorla (Jaén) Mateau-Cobos coll.’ (MHNH); 1 ♀: ‘ Fte. Bermejo. Jaén’, ‘ruela S. Cazorla Mateau-Cobos coll.’ (MHNH) ; 1 ♂: Sierra Nevada, Capileira. 1600 m a.s.l. 22.03.1994. V. Assing leg. (cSh) ; 9 ♂♂, 4 ♀♀: same data, but P. Wunderle leg. (cW, cSh); 1♂: Mt. Mulhacen. 2000 m. a.s.l. 21.05.1967. M.E. Bacchus & B. Levey leg. (MHNG) . LOCALITY NOT SPECIFIED: 1 ♂: ‘ Miguelon’ (MHNH) .

Additional material. ANDORRA: 1 ♂: Puerto de Envalira. 2200-2300 m a.s.l. 30.05.1994. M. Schülke & Grünberg leg. (cSch) ; FRANCE: 1 ♂: ‘ Gavarnie 25.5.[19]31’ (MHNH) ; 2 ♂♂: ‘ Gavarnie, H. Pyr. 12-31, vii. 14. G.C.C. ’ (BMNH) ; 2 ♂♂: ‘ Lourdes’ (IRSN); 8 ♂♂, 7 ♀♀: Por d`Boucharo. 30.06.1976. (MHNG) ; 14 ♂♂, 6 ♀♀: Ht. Canigou. 02.07.1976. (MHNG) ; 4 ♂♂, 3 ♀♀: Sahorre d. Vernet. 04.07.1976. (MHNG) ; 1♀: Ariège, Auzat. 19.07.1972 (MHNG) ; 1 ex.: ‘ Pyr. Or. La Vall. H. Coiffait 24.3.[19]48’ (MHNH) ; 1 ♂, 1 ♀, 1 ex.: Banyuls-sur-Mer (IRSN, MHNH); 1 ♂: Gard Fressac. 28.10.1982. I. Löbl leg. (MHNG) ; 8 ♂♂, 2 ♀♀: ‘ Col de Aras Pyren. or.’ (MHNG) .

Remarks. Lesteva fageli was originally described based on several type specimens from Cantabria (“Puente del Cerezo”) and central Spain (“Rio Guadalupejo”). Lohse compared it with L. fontinalis and L. pubescens . He distinguished it from the latter by the darker body and smaller aedeagus (“0,64 mm gegenüber 0,75 mm bei pubescens ”). During the study of types of L. fageli, we found that these specimens are morphologically conspecific with L. pubescens, a variable species regarding coloration, body size, and the proportions of the forebody and aedeagus (Figs. 38–46). Thus, we synonymize L. fageli with L. pubescens . Regarding the presence of stout pubescence on the median lobe, this species is similar to L. pourtoyi Jarrige, 1972, a species originally described from “Cirque de Mahourat (Pyr[énées].-Atl[antiues].)”, but the aedeagus of this species is evidently narrower (Fig. 5 in Jarrige (1972)) and is furnished with longer pubescence. According to Tronquet (2014), L. pourtoyi is also known from Spain (“ pourtoyi Jarrige, 1972: 158 . - Pyr. occ. et cent. Esp. (chaîne Cantabrique”)), but this record needs confirmation. Habitus of L. pubescens as in Fig. 31; aedeagus as in Figs. 38–46. Lesteva pubescens is widely distributed in Europe (Schülke & Smetana 2015) and had already been reported from Spain (Lohse 1960).

It is here recorded from Portugal for the first time.