Anacroneuria ypane sp. n.

(Figs. 11-15)

Material examined. Holotype ♂, PARAGUAY: Amambay Department, Rio Ypane, cerca Pedro Juan Caballero, 25 November 1956, C.J.D. Brown (USNM).

Adult habitus. General color brown, patterned with areas of pale pigment. Head with a large area of dark pigment extending from ocellar region over central frons (Fig. 11); dark pigment area expanded laterally in central frons where a small oval pale area is located; lappets and antennae dark brown. Pronotum mostly dark but with a narrow pale median stripe and pale lateral areas. Hind femora dark brown in apical third but pale basally; hind tibia with small dark bands basally and apically but pale for most of length. Wing membrane brown, veins (particularly R vein) dark brown.

Male. Forewing length 10 mm. Hammer thimble shaped, apical diameter less than height (Fig. 12). Aedeagal apex a short, simple, truncate scoop bearing a ventral pair of prominent membranous lobes (Fig. 13); dorsal margin of apical area expanded into a pair of small rounded lateral lobes near point where hooks cross (Fig. 15); dorsal keel lines absent, apex bent strongly dorsad, giving a foot-like shape in lateral aspect (Fig. 14). Hooks slender.

Female. Unknown.

Larva. Unknown.

Etymology. The species name, used as a noun in apposition, is based on the type locality of Rio Ypane.

Diagnosis. This species is related to the Anacroneuria atrifrons Klap á lek group of species (Froehlich 2008) which includes at least A. azul Rojas & Baena (Stark et al. 1999), A. canchi Stark & Sivec (Stark & Sivec 1998) and A. pastaza Stark (Stark 2001) . Of these species, the aedeagus of A. ypane is most similar to that of the Colombian species, A. azul, but the membranous lobes in that species are smaller and a small transverse dorsal keel is present. The latter species also differs from A. ypane in having a much smaller area of dark head pigment.

Comments. Froehlich (2002) reported A. debilis (Pictet) from a single male collected in Parque Nacional Ybycui and A. trimacula Jewett from a single female collected at the same locality. Thus, Anacroneuria ypane becomes only the third member of the genus to be confirmed for Paraguay, and the first to be described from a Paraguayan specimen.