Atopida insularis sp. nov.

(Figs 1J–K, 3C–D, 5G–H, 8E, 9A, 10D, 12D, 17O)

Type material. Holotype, male (NZAC): “New Zealand FD \ South Arm \ L Manapouri \ 14-Dec-98”, “ General \ beating”, “ Ex John Nunn \ Collection \ Donated 2015”; paratypes, 5 exx. (NZAC): same data as holotype; paratype, 1 ex. (LUNZ): “ NEW ZEALAND FD \ Fiordland Nat Pk \ Murchison Mts \ Plateau Ck \ 990m ”, “ 3. XII.1980 \ R. M. Emberson \ C. A. Muir ”, “ Malaise trap \ N. menziesii \ forest edge”; paratypes, 7 exx. (NZAC): “ Upper Holly- \ ford Val. \ Homer ”, “12.xii.66 \ A.K. Walker ”, “ Olearia \ ilicifolia”; paratype, 1 ex. (NZAC): “ Hidden falls \ Tr.14.xii.66”, “ Lower \ Hollyford Val. \ A.K.Walker ”, “ General \ beating”; paratype, 1 ex. (NZAC): “Hollyford Val. \ 13.I.67 \ A. K. Walker ”, “at night \ off ferns”; paratypes, 2 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ Gut hut \ 22-28 Nov 1981 ”, “ C.F. Butcher \ Malaise trap ”; paratypes, 2 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 600m \ 30 Nov 1981 ”, “ C.F. Butcher \ sweeping \ at night”; paratype, 1 ex. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 29 Nov 1981 \ C.F. Butcher ”, “ ex pan trap \ in tussock”; paratypes, 13 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 850m \ 30 Nov 1981 \ C.F. Butcher ”, “sweeping \ at night”; paratypes, 12 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 850m \ 30 Nov 1981 \ C.F. Butcher ”, “sweeping \ tussock”; paratype, 1 ex. (NZAC): “ NEW ZEALAND FD \ Secretary I \ 850m \ 30 Nov 1981 \ C.F. Butcher ”, “ Insect spray \ rock face”; paratypes, 4 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ ridge towards \ Mt Grono \ 27 Nov 1981 ”, “ C.F. Butcher ”; paratypes, 3 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ ridge towards \ Mt Grono \ 27-30 Nov 1981 ”, “ C.F. Butcher \ Pan trap ”; paratypes, 4 exx. (NZAC): “ NEW ZEALAND FD \ Secretary I \ ridge towards \ Mt Grono, 853m \ 27 Nov 1981 ”, “ C.F. Butcher \ Mats and mosses \ 81/189”; paratype, 1 ex. (NZAC): “ NEW ZEALAND FD \ Secretary I \ ridge towards \ Mt Grono, 853m \ 30 Nov 1981 ”, “ C.F. Butcher \ mats, moss and \ tussock 81/186”; paratype, 1 ex. (NZAC): “NEW ZEALAN FD \ Camp 1020m \ Tutoko Bench \ Darran Mts ”, “ 13 Jan 1977 \ T.K. Crosby and \ J.S. Dugdale \ Malaise trap ”; paratype, 1 ex. (NZAC): “ Alpine \ Garden ”, “ Homer Tunnel \ to Milford Rd. ”, “12.xii.66 \ A.K. Walker ”, “ General beating”; paratype, 1 ex. (NZAC): “ Bold Peak, \ 27-12 to 1-1-45 \ E.S. Gourlay ”, “ Atopida \ lawsoni Shp. \ Id.by J. Armstrong ” paratype, 1 ex. (NZAC): “ Fox Glacier \ 14.12.60 \ S.Westland,”, “ J.I. Townsend \ & P.R.Kettle ”; paratype, 1 ex. (NZAC): “ NEW ZEALAND WD \ Mt Tuhua, 1067m \ E. side of \ L. Kaniere \ 20 Nov 1984 ”, “ B. A. Holloway \ Moss and \ lichen 84/75” ; paratype, 1 ex. (NZAC): “ NEW ZEALAND FD \ Bauza I \ Mar 1984 \ C.F. Butcher ”; paratype, 1 ex. (NZAC): “ Flat Top Mt. Hunter \ Mts. L. Manapouri \ 4000’ Clark.”, “ A.E. Brookes \ Collection ” ; paratype, 1 ex. (NZAC): “ S of Cascade \ Eglinton R \ 30.X.66 J. I. T”, “moss under \ beech trees”; paratype, 1 ex. (NZAC): “New Zealand OL \ S. Mavora Lake \ East side \ 16-xii-2013 ”, “ Sifted \ ground moss \ Nothofagus \ forest”, “ Ex John Nunn \ Collection \ Donated 2015” ; paratype, 1 ex. (NZAC): “ New Zealand SL \ Princhester Base \ Takitimu Forest \ 10-Dec-04”, “ On low \ vegetation by \ stream”, “ Ex John Nunn \ Collection \ Donated 2015” ; paratype, 1 ex. (NZAC): “13-I-66 \ Takitimus 2500’ \ A.C. Eyles ”, “ Celmisia \ (sticky)” ; paratype, 1 ex. (AMNZ): “ Routeburn \ R. \ 4-I-45 \ C.E. Clarke ”, “ C.E. Clarke \ Collection ” ; paratype, 1 ex. (AMNZ): “ Upper \ Routeburn R. \ 18-I-26 \ C.E. Clarke ”; paratypes, 2 exx. (NMPC): “ NEW ZEALAND: Fiordland (FD) \ Borland Road 24 km NWW of \ Monowal, 45°41.13’S \ 167°20.29’E; 320 m; 24-27.i.2016; \ Seidel, Sykora & Fikáček lgt. \ 2016-NZ024”, “sifting of leaf litter, moss and \ rotten logs in degraded remnants \ of Nothofagus forest with \ numerous rotten logs”.

Diagnosis. Body small to moderately large (TL 3.3–4.5 mm), colouration variable. Similar to some other Atopida from South Island, certain identification is possible only on the basis of genital structures: trigonium much shorter than parameroids, curved laterally, parameroids triangularly pointed at apices, bursal sclerite subtriangular in outline, with paired tubercles in central portion.

Description. Male. Body elongated, moderately large (TL 3.7–4.5 mm), moderately convex. Body colouration variable, from yellowish to brown, head and pronotum slightly darker, elytra yellowish with dark suture or uniformly yellowish to dark brown. Punctation on head and pronotum granulate, strong and dense, punctures touching each other, punctation on pronotum becoming more sparse towards the middle and base, surface of pronotum uneven, in some specimens forming two tubercles in basal portion of pronotum, punctation on elytra simple, coarse, punctures separated by 0.3–0.7× diameter of a puncture. Body covered with moderately long, protruding, yellowish setae, more numerous and ruffled on head and pronotum. Supraantennal ridges well-marked, elevated, forming triangular flap. Pronotum widest slightly before middle of its length, sides curved, lateral carinae crenulate, anterolateral angles produced, triangular, pointed, posterolateral angles right-angled. Base of pronotum with longitudinal depression along posterior margin. Elytra without carinae, humeri well-marked. Ventrite 5 rounded, apex subtly truncate. Penis (L 0.70 mm, W 0.36 mm) elongate, with explanate lateral lobes, trigonium very short, narrow, curved laterally at ~right angle; parameroids elongated, sinuate, outer edges subtly denticulate, apices pointed, covered with punctures; pala triangular, very short, significantly shorter than parameroids. Tegmen (L 0.67 mm, W 0.46 mm) with narrow, triangular parameres, covered with scarce punctures and numerous denticles.

Female. Body small to moderately big (TL 3.3–4.1 mm), brachypterous. Supraantennal ridges well-marked, elevated, forming triangular flaps in some specimens. Colouration and punctation like in males, but central portion of pronotal disc sparsely punctate, with shining interspaces. Ventrite 5 rounded, with very shallow emargination at apex. Lateral portions of ventrites 3 and 4 rather big, oblique setal patches, small oblique patches visible also on ventrite 5. Bursal sclerite big (L 0.38 mm, W 0.32 mm) consisting of two interconnected plates—one with pair of tubercles visible in central portion of sclerite, the second forming hook-like processes resembling long spines in dorsal view.

Measurements and ratios. Males (n = 10) TL 3.65–4.45 mm (4.05 mm), PL 0.65–0.75 mm (0.69 mm), PW 1.10–1.30 mm (1.19 mm), EL 3.00– 3.70 mm (3.36 mm), EW 1.70–2.00 mm (1.89 mm), TL/EW 1.92–2.25 (2.14), PW/PL 1.57–1.86 (1.73), EL/EW 1.59–1.89 (1.78), EL/PL 4.33–5.46 (4.88), EW/PW 1.50–1.68 (1.59). Females (n = 10) TL 3.30–4.05 mm (3.82 mm), PL 0.60–0.75 mm (0.67 mm), PW 1.10–1.40 mm (1.24 mm), EL 2.75–3.30 mm (3.14 mm), EW 1.70–2.05 mm (1.83 mm), TL/EW 1.94–2.31 (2.09), PW/PL 1.60–2.08 (1.86), EL/EW 1.61–1.89 (1.71), EL/PL 4.40–5.25 (4.72), EW/PW 1.40–1.58 (1.48).

Distribution. The species is distributed mostly in south-west areas of South Island—Fiordland and Otago Lakes, rarely occurs also in Westland area (Fig. 18D).

Etymology. Latin insularis = insular, an allusion to the fact that the species is common on Secretary Island.