Tomarus roigjunenti Neita & Ratcliffe, 2017

(Figs. 9K, 16O, 20F, 28F; 47)

Tomarus roigjunenti Neita & Ratcliffe, 2017: 8 . Original combination.

Male holotype not examined. Male paratype (USNM) “ Neuquen / Capital Federal / 1.iii.1961 / Dafuern // TOMARUS / ROIGJUNENTI / NEITA & RATCLIFFE / PARATYPE ”. Female paratype (USNM) “ Neuquen / Capital / 1.iii.1961 // TOMARUS / ROIGJUNENTI / NEITA & RATCLIFFE / PARATYPE ”. Two female paratypes (USNM) “ Mendoza / C.S. Reed // TOMARUS / ROIGJUNENTI / NEITA & RATCLIFFE / PARATYPE ” Type locality: Neuquén, Argentina .

Description. Habitus as in Fig. 28F. Length 26.3–27.6 mm; humeral width 12.6–13.9 mm. Color dark reddish brown to black. Head: Frons coarsely rugose. Frontoclypeal region with 2 transverse tubercles separated by about 3 tubercle diameters. Clypeus narrowed towards apex, base 3.8 wider than apex. Clypeal teeth triangular, separated by less than a tooth diameter. Mandible with 2 apical teeth and a lateral, narrowly rounded tooth. Mentum abruptly constricted at apical 3rd. Galea of maxilla with teeth 5 and 6 well developed. Interocular distance 4 times an eye width. Pronotum: Surface with dense, large punctures. Apical tubercle small, rounded. Fovea shallow, striate, narrow (1/5 as wide as interocular distance). Scutellum: Surface with minute, sparse punctures. Elytra: First interval punctate, punctures similar in size to other intervals; sutural stria complete. Inner surface of apex with rounded, small tubercles forming 38–40 parallel lines. Abdomen: Apex of tergite IV with a triangular wide area with large, elongate tubercles forming about 18 diagonal, parallel lines. Pygidium with dense rugosity on basal 3rd (Fig. 9K), apex regularly rounded. Venter: Apex of prosternal process flat, longitudinally oval. Metasternum with short, sparse setae on anterior angles. Legs: Protibia tridentate with an additional inconspicuous, basal convexity. Metatibia triangular; apex slightly crenulate, with 21–25 spinules. Male genitalia: Spiculum gastrale without basal plates. Phallobase 1.1 times longer than parameres. Parameres with lateral teeth large, acute; apical half narrowed, apices outwards, rounded (Figs. 16O, 20F). Internal sac with copulatory lamella and short lamellar spiny belt; with a complex of 1–3 spine-like, accessory lamellae, with dense granules at base.

Diagnosis. Tomarus roigjunenti can be recognized by the following character combination: frontal tubercles transverse; pronotal fovea and tubercle inconspicuous (Fig. 28F); pronotum deeply and densely punctate; metasternum with sparse, minute setae on anterior angles; internal sac with a complex of 1–3 spine-like, accessory lamellae; and parameres with lateral teeth large and long, arcuate; apical half narrowed, apices outwards, rounded (Figs. 16O, 20F).

Distribution. Argentina (Neita-Moreno & Ratcliffe 2017).

Locality records (Fig. 47). 4 examined specimens from USNM. Records from Neita-Moreno & Ratcliffe (2017). ARGENTINA (20). Chubut (1): Chubut. Buenos Aires (1). Buenos Aires. Mendoza (4). Tunuyan; Uspallata. Neuquén (9). Capital Federal; Barrancas; Collón Cura; Covunco; Isla Victoria; Laguna Blanca; Vatra Lauquen. Río Negro (5). Allen; Cipolletti; General Roca; Río Negro; Villa Regina .

Natural history. Some specimens were found in sand dunes and adults are attracted to lights at night (NeitaMoreno & Ratcliffe 2017).