3.2.3. Rumilara obscura sp. nov.

Figs 2a, 3a, 4b-i, 5a, 6a, 7a, 8a, 9a, 9e, 10a-d, 11

Material examined.

Type material: Holotype ♂ (PUCE) " Ecuador, Cotopaxi prov., Otonga env., Rio Esmeralda, 00°25′10.4″ S, 78°59′46.9″ W, 1760m a.s.l., 8.8.2013, stream ca 3m wide, in primary forest, with gravel, stones, submerged wood, Čiampor Jr & Čiamporová-Zaťovičová lgt." ; Paratypes 4♂♂, 2♀♀ (PUCE, CCB), 2 ex with the same data as holotype; 4♂♂, 1♀, " Ecuador, Cotopaxi prov., Artos env., above confluence with Rio Esmeralda, 00°25′09.6″ S, 78°59′40.2″ W, 1735m a.s.l., 8.8.2013, stream ca 2m wide, fast flowing from steep slope, with gravel, stones with moss, boulders, Čiampor Jr & Čiamporová-Zaťovičová lgt." Other material: 4 larvae with the same data as holotype; 5 larvae " Ecuador, Cotopaxi prov., Artos env., above confluence with Rio Esmeralda, 00°25′09.6″ S, 78°59′40.2″ W, 1735m a.s.l., 8.8.2013, stream ca 2m wide, fast flowing from steep slope, with gravel, stones with moss, boulders, Čiampor Jr & Čiamporová-Zaťovičová lgt." .

Adult diagnosis.

The species can be distinguished by a combination of following characters: 1) head and clypeus fully clothed by pubescence; 2) femora bare or with a row of dark, long hairs on dorsal margin; 3) elytra without dense pubescence; 4) aedeagus with rod-like sclerite; 5) parameres only feebly asymmetrical.

Adult diagnostic description.

Male. Colour: Body (Figs 2a, 3a) black; lateral sides of femora and tibiae brown to pale brown; tarsal claws pale brown with reddish tincture. Pubescence: Head and clypeus fully clothed with short, prone pubescence, most densely set behind eyes. Pronotum (Fig. 6a) partly covered by fine, short, moderately dense, prone pubescence. Elytra at lateral and apical edges with short, recumbent pubescence, and with numerous moderately long, curved, semi-erect, hair-like setae. Femora with long, dark, densely set, hair-like setae near dorsal margin; pro- and metatibiae apically with a fringe of long, dark, hair-like setae on inner margin; mesotibiae without pubescence. Metaventrite densely clothed with short, prone pubescence. Surface: Head, clypeus and pronotum sparsely microreticulate, partly concealed by pubescence. Elytra with rows of punctures deeply impressed; intervals nearly flat. Prosternal process (Fig. 7a) moderately wide, concave between procoxae, then subtriangular with rounded apex. Aedeagus (Figs 11a-c) with long, dorsal, rod-like sclerite on penis; parameres asymmetrical, left one slightly longer. - Female externally similar to male, except slightly larger.

Measurements.

♂ - CL: 2.70-2.93 mm; PL: 0.67-0.70 mm; PW: 0.76-0.79 mm; EL: 2.02-2.24 mm; EW: 0.99-1.15 mm. ♀ - CL: 2.98-3.00 mm; PL: 0.68-0.72 mm; PW: 0.80-0.83 mm; EL: 2.25-2.31 mm; EW: 1.11-1.14 mm.

Variation.

The greatest differences were observed in distribution of pubescence (mainly on legs, around eyes, partly on pronotum and elytra) due to its often removal.

Etymology.

From Latin obscūrus (dim, dark, obscure) due to its dark colour.

Larva

(Fig. 9a, e) with length 3.86-4.04 mm, greatest width 1.04-1.16 mm. Pleurites on abdominal segments 1-5. Abdominal segments 1-5 explanate. Surface around the frontal pair of pronotal gibbosities of the same colour as the rest of pronotum. Tergum of abdominal segments 1-7 with a pair of distinct, longitudinally arranged small gibbosities.

Distribution.

Known from two localities in Cotopaxi Province (Figs 16a-b) and one locality in Santo Domingo de los Tsáchilas Province (Fig. 16c).