Trypodendron signatum (Fabricius, 1792)

Distribution in Iran. Caspian Sea area and other northern provinces (Modarres Awal 1997 as Xyloterus signatus), Guilan (Borumand 1998), Iran (no locality cited) (Knížek 2011).

General distribution. Europe, Turkey, through Russia to Mongolia, China, Korea, Japan.

Biology. This species breeds in angiosperm trees in the families Betulaceae, Fagaceae, Oleaceae, Salicaceae, Sapindaceae (Pfeffer 1995) . Recorded in Iran from Betula pendula (Betulaceae), Fagus orientalis, Quercus sp. ( Fagaceae), Ulmus pumila (Ulmaceae) (Modarres Awal 1997). The biology is similar to T. lineatum (see above). The species usually attacks dead or dying trees. However, attacks on living Fagus sylvatica (Fagaceae) have been recorded in Belgium (Gaubicher et al. 2003, Kühnholz et al. 2001), possibly because the trees were stressed by frost damage (Henin et al. 2003).