Mantidactylus betsileanus clade

A rather species-rich clade of small to medium sized species (21.1–40.0 mm adult SVL), most of which are characterized by comparatively slender body shape with comparatively long limbs, relative to most other Brygoomantis species, and a more or less strongly expressed white tip on the snout. Contains: M. betsileanus, M. noralottae, M. tripunctatus (revalidated herein) and five new species (named based on holotypes depicted in Fig. 33). Note that several morphologically or bioacoustically similar species previously considered to be related to M. betsileanus (e.g. Glaw & Vences 2007) are here placed in a different clade ( M. fergusoni clade, see below)