D. villosus Szymczakowski, 1961: 152,

1963: 678 (description correction), 1968: 21 (redescription of male genitalia); Gnaspini, 1991a: 328; Salgado, 2005d: 980, 2010b: 162 (assignment to group).

Holotype male in ZMPA.

Type locality: “Bocaina region”, São Paulo State, Brazil.

Distribution: Brazil (in caves and forest): Minas Gerais, Paraná, Santa Catarina, and São Paulo States. Larvae: Gnaspini, 1993b: 551 .