Gastrotheca nicefori Gaige, 1933

Holotype: UMMZ 73242.

Type locality: Pensilvania, Departamento Caldas, Colombia.

Distribution: Region 1. Panama, Colombia, and Venezuela. In Venezuela, distributed throughout the Cordillera de Mérida. Very likely present in Sierra de Perijá.

Remarks: Duellman (1989b) synonymized Gastrotheca yacambuensis Yústiz, 1976 with G. nicefori, without explanation. La Marca (1992) recognized the synonymy, but Barrio-Amorós (1998, 2004, 2009) was reluctant to do so as it was done without providing any comparison. Duellman (2015) presented arguments for the synonymy (see below), but molecular evidence (Castroviejo et al. 2015) indicated G. yacambuensis could be a valid species, different from G. nicefori . Further evidence is necessary to assess this issue adequately. The species is included in the subgenus Duellmania (Duellman 2015) and in the Gastrotheca marsupiata species group sensu Castroviejo-Fisher et al. (2015).

Selected references: Yústiz (1976a,b, 1996); Duellman (1980, 1989b, 2015); Duellman et al. (1988); La Marca (1991b “1994,” 1992); Castroviejo-Fisher et al. (2015).