Oreophrynella nigra Señaris, Ayarzagüena and Gorzula, 1994

Holotype: MHNLS 10583.

Type locality: Kukenan-tepui I, Bolívar State, Venezuela (9°51’N, 60°48’W), 2,500 m asl .

Distribution: Region 5. Endemic to Kukenan and Yuruaní Tepuis in Bolívar State.

Remarks: Kok et al. (2012) implied that O. nigra could be a synonym of O. quelchii due to the low genetic divergence between these forms. However, inhabiting different tepuis means reproductive isolation (allopatry), and having a different color pattern could be sufficient reason at the moment to maintain it as a valid species.

Selected references: McDiarmid and Gorzula (1989); Solano (1989); Señaris et al. (1994, 2014); Gorzula and Señaris (1998); McDiarmid and Donnelly (2005).