Leptodactylus rugosus Noble, 1923

Holotype: AMNH 1169.

Type locality: “near Kaieteur Falls, British Guiana.”

Distribution: Region 5. Venezuela and Guyana. In Venezuela, in lowlands and uplands of eastern Guiana shield (Bolívar State) including summits of some tepuis.

Remarks: In Leptodactylus pentadactylus species group of De Sá et al. (2014).

Selected references: Ginés (1959); Rivero (1961, 1964b,d, 1968d); Heatwole et al. (1965); Heyer (1979, 1995); Hoogmoed and Gorzula (1979); Donnelly and Myers (1991); Magdefrau et al. (1991); Duellman (1997); Gorzula and Señaris (1998); Heyer and Thompson (2000); Barrio-Amorós and Duellman (2009); Heyer and Barrio-Amorós (2009); De Sá et al. (2014); Señaris et al. (2014).