Genus Carolia Cantraine, 1838

Remarks

The genus Carolia Cantraine, 1838, is thought to have appeared for the first time in lower Eocene strata of Egypt, India, and Pakistan and to have spread across the Tethyan region during the middle Eocene (El-Shazly et al. 2016). Specimens from Maastrichtian beds in Egypt identified as examples of Carolia (Quaas 1902) were subsequently assigned to a new genus, Tarturia Strougo, 1983 (Strougo 1983), thought to be ancestral to Carolia (El Shazly et al. 2016) . Species of Carolia have been described from lower and lower middle Eocene beds of India (Cox 1938) and Oligocene beds in Argentina (Ihering 1907).