Bia actorion (Linnaeus)

(Figs 1a, 3, 11 a–b, 12)

Type species.

Diagnostic re-description. General characters as described for the genus. (1) MF DFW white apical ocelli small to large. (2) MF DFW orange band varies from thin (Venezuela, Alto Orinoco) to medium-wide; orange scales extended proximally along veins, including the anterior tip of discal cell in some specimens (Brazil, Amazonas, Manaus, but not all specimens). (3) M DFW iridescent band from anal margin to approximately half of the CuA2 cell (one specimen from Venezuela with DFW iridescence clearly reaching CuA2); F DFW iridescent band well developed but variable; usually broad below CuA2, and more extended towards tornus than those of female decaerulea decaerulea and rebeli . (4) M DFW androconial organ on the CuA-CuA1-CuA2 intersection dark brown, matching scale color of surrounding area. This character is shared by B. actorion and B. decaerulea . (5) M DHW discal androconial pad varies from being nearly black and darker than associated hairpencil (Venezuela; Guyana; Suriname; Brazil, Manaus) to dark brown and of similar color to hairpencil (photos from Colombian specimens, Ecuador, northern Peru; Brazil, northern Pará) . (6) M DHW discal hairpencil dark brown. (7) F VFW ripple pattern of the postmedial area slightly less dense than that of M in some localities (Brazil and Guyana) and clearly less dense in others (Venezuela, Ecuador). Male genitalia (Figs 2 and 11 a): tip of valva usually narrow, spiny projection usually of medium height, infrequently tall, spines larger in specimens from the Guyana region; gnathos wide (Guyana region) to narrow (other localities).

Type material. The iconotype designated by Hemming (1964) from Clerck’s Icones (Fig. 1a) does not show the color of androconial organs and also lacks locality data, both of which are important attributes for definitive identification of actorion . We therefore recognize that a neotype should be selected for this species, which will be done separately from this study. Note that specimens in the Linnean Collection are not types (male http://linneanonline.org/14787/ and female http://linnean-online.org/14788/ last accessed 30 August 2016; see also http:// butterfliesofamerica.com/L/t/ Bia _a.htm last accessed 30 August 2016).

Remarks. Geographical variation in some diagnostic characters serves as the basis for the subspecific classification proposed below.