Patu jidanweishi Miller, Griswold & Yin, sp. n.

urn:lsid:zoobank.org:act: 1C8AF042-4D42-48BA-A072-1058266329FD

Figs 65 A-E, 66, 67 A-D, 68-71, 100

Patu sp., Griswold and Yan, 2003: 356-360.

Material Examined. Holotype: CHINA: Yunnan: Lushui Co., Pianma Township, Chang Yan He, 9.3 km ESE Pianma, 25.99363°N, 98.66651°E, 2470 m, 13-15 May 2005, mixed broadleaf deciduous and evergreen forest, dusting small webs near ground in forest understory, C. Griswold, CGY104 (CASENT 9029293, HNU), 1 ♁.

Paratypes: [same data as holotype] (CASENT 9022356, CAS), 2 ♁, 4 ♀, 4 juvs; [same data as holotype] (CASENT 9029294, PV 0713-0716, CAS), 1 ♀; [same data as holotype] (CASENT 9029295, PV 0723-0728, CAS), 1 ♀; Longling Co., Longjiang Township, Xiao Hei Shan Nature Reserve, 1.2 km SSE of Route S317 at km 23.5, 24.82888°N, 98.76001°E, 2020 m, 27-28 May 2005, good primary broadleaf forest, dusting webs in understory, C. Griswold, CGY128 (CASENT 9029296, PV, CAS), 1 ♀ ; Nujiang Prefecture, native forest in Gaoligongshan at 9.5 road km ESE Pianma, 25.98333°N, 98.66667°E, 2500 m, 15-18 October 1998, C. Griswold, D. Kavanaugh, C.L. Long (CASENT 9029303, CAS), 1 ♁, 1 ♀ ; [same data] (CASENT 9023961, CG 17 x98(2)7-15, CAS), 1 ♀; [same data] (CASENT 9023960, CG 17 x98(2)16- 22, CAS), 1 ♀; Lushui Co., Pianma Township, Chang Yan He, 9.3 km ESE Pianma, 25.99363°N, 98.66651°E, 2470 m, 12-21 May 2005, mixed broadleaf deciduous and evergreen forest, pitfall traps, C. Griswold, D. Kavanaugh, K. Guo, CGY103 (CASENT 9023102, CAS), 1♀, 4juvs ; Lushui Co., Pianma Township, Chang Yan He, 9.3 km ESE Pianma, 25.99363°N, 98.66651°E, 2470 m, 12 May 2005, mixed broadleaf deciduous and evergreen forest, winkler extraction of sifted leaf litter, C. Griswold, D. Kavanaugh, K. Guo, CGY102 (CASENT 9022328, HNU), 1♀ ; Fugong Co., S Fork, Yamu He, 1.51 km 150° SW of confluence [with N Fork], Gaoligongshan, 27.11905°N, 98.83108°E, 1723 m, 26 April 2004, moist shaded embankments, C. Griswold, CGY13 (CASENT 9020753, HNU), 1♀ ; Gaoligongshan, 0.4 km SSE Shibali forest station, 27.16337°N, 98.78208°E, 2475 m, 5 May 2004, dusting webs in understory of good forest, C. Griswold, CGY29 (CASENT 9019862, CAS), 1 ♁, 1 ♀ ; [same data] (CASENT 9019863, HNU), 1 ♁, 1 ♀; [same data] (CASENT 9019864, HNU), 1 ♀, 4 juvs; Gaoligongshan, 0.5 km radius of Shibali forest station, 27.16519°N, 98.77891°E, 2525 m, 1-9, May, 2004, dusting webs in forest, C. Griswold, CGY25 (CASENT 9019875, CAS), 1 ♁, 1 ♀ ; [same data] (CASENT 9019877, HNU), 1 ♀, 11 juvs; [same data] (CASENT 9020650, HNU), 1 ♁, 1 ♀; [same data] (CASENT 9019874, CAS), 1 ♁, 1 ♀; [same data] (CASENT 9019876, HNU), 1 ♁, 1 ♀; [same data] (CASENT 9019873, CAS), 1 ♁, 1 ♀; [same data] (CASENT 9024143, HNU), 1 ♁, 2 ♀, 1 juv; Gaoligongshan, Shibali forest station, 27.16636°N, 98.77667°E, 2563 m, 3-11 May 2004, good forest, pitfall traps, C. Griswold, D. Kavanaugh, CGY21 (CASENT 9020351, HNU), 2 ♁, 1 ♀ ; [same data] (CASENT 9020350, CAS), 1 ♁, 1 ♀; Gaoligong Shan, Nujiang Prefecture: Nujiang State Nature Reserve, QiQi He, 9.9 airkm W of Gongshan, 27.715°N, 98.565°E, 2000 m, 9-14, July, 2000, H.-M. Yan, D. Kavanaugh, C.E. Griswold, H.-B. Liang, D. Ubick, & D.-Z. Dong (CASENT 9000375, HNU), 4 ♁, 4 ♀ ; [same data] (CASENT 9000374, CAS), 2 ♁, 10 ♀, 1 juv; [same data] (CASENT 9000373, HNU), 14 ♀; [same data] (CASENT 9000339, CAS), 1 ♀; [same data] (CASENT 9000370, “DL photo 7/12: symphyt beneath 1f w/ mys”, CAS), 1 ♀; [same data] (CASENT 9000338, CAS), 1 ♀; [same data] (CASENT 9000343, CAS), 1 ♀; [same data] (CASENT 9000371, HNU) 2 ♁, 10 ♀, 1 juv; [same data] (CASENT 9000372, CAS), 14 ♀; [same data] (CASENT 9000342, “DL photo 1/12: indiv, symphyt w/eggs”, CAS), 1 ♀; [same data] (CASENT 9000369, HNU), 1 ♀; [same data] (CASENT 9000340, CAS), 1 ♀; [same data] (CASENT 9000341, CAS), 1 ♀; [same data] (CASENT 9023115, HNU), 1 ♀ .

Etymology. Formed from the Chinese words for egg (jī dàn) and guardian (wèi shì), referring to the egg-guarding behavior described by Griswold and Yan (2003).

Diagnosis. Distinguished from other Patu species except P. woodwardi Forster, 1959, P. marplesi Forster, 1959, P. qiqi and P. xiaoxiao by the presence of only a single cheliceral tooth (Figs 69 E-F, 70D, Forster 1959: fig. 122; two or more in other Patu species); male distinguished from P. woodwardi and P. marplesi by the presence of two ventral subdistal spines on tibia II (Fig. 65E); distinguished from P. qiqi by the much longer embolus in P. jidanweishi (Fig. 66; short, not flexible in P. qiqi, Fig. 73B; male unknown in P. xiaoxiao); female distinguished from P. qiqi and P. xiaoxiao by the presence of a scape (Figs 67 A-C, 70A-B).

Description. Carapace brown, male clypeus strongly raised with clusters of sulci (Fig 69 A-D). Six eyes in three doublets (Fig. 69B). Sternum dark brown. Chelicerae fused near the base, with single prolateral tooth, fang furrow absent (Fig. 69 E-F). Labrum not swollen, bearing single plumose seta, with ventrally rounded labral tongue (Fig. 70 E-F). Female palp absent. Legs brown. Abdomen brown (Fig. 65 A-D).

Male palp: Cymbium covers proximodorsal part of bulb (Fig. 66). Tegular region divided into numerous lobes (Fig. 68B). Embolus long, free, follows complex path (Fig. 66).

Vulva: Epigynum a lightly sclerotized scape (Fig. 67 A-B) with a socket near the tip (Fig. 67C). Ducts make many turns around spermathecae (Fig. 67C).

Male (CASENT 9022356): Total length 0.69, carapace 0.29 long, 0.30 wide, clypeus 0.12, sternum 0.20 long, 0.20 wide, coxa IV separated by 2.10 times their width. Leg I: patella d1, tibia d2; Leg II: patella d1, tibia d2, v2 on tubercle, metatarsus with ventral tubercle near center; Leg III: patella d1, tibia d1; Leg IV: patella d1, tibia d1. Metatarsal trichobothria: TmI: 0.43; TmII: 0.42; TmIII: 0.50. Leg measurements: see Appendix A. Epiandrous gland spigots absent (Fig. 68F). Posterior lateral spinnerets with sticky silk triplet, aggregate gland spigots on common base (Fig. 71F).

Female (CASENT 9022356): Total length 0.88, carapace 0.34 long, 0.31 wide, clypeus 0.05, sternum 0.22 long, 0.21 wide, coxa IV separated by 1.83 times their width. Leg I: patella d1, tibia d2; Leg II: patella d1, tibia d2; Leg III: patella d1, tibia d1; Leg IV: patella d1, tibia d1. Metatarsal trichobothria: TmI: 0.43; TmII: 0.44; TmI- II: 0.48. Leg measurements: see Appendix A. Spinnerets (Fig. 71A-D), posterior lateral spinnerets with aggregate gland spigots on common base (Fig. 71D).

Natural History. This species builds a horizontal orb web (Fig. 72 A-B; see also page 12). Eggs (Fig. 67D) are placed on the periphery of the web, each individually wrapped (see Griswold and Yan 2003).