Strigiphilus heterocerus (Grube, 1851)

(Figs 2–6)

Docophorus heterocerus Grube, 1851: 469 .

Type host: Strix uralensis liturata (Tengmalm, 1795) .

Strigiphilus fukuro Uchida, 1948: 307, fig.1; Hopkins & Clay 1952: 339.

Type host: Strix uralensis hondoensis (Clark, 1907) .

Neodocophorus uralensis Eichler, 1949: 15, figs 33–34; Hopkins & Clay 1952: 340.

Type host: Strix uralensis liturata (Tengmalm, 1795) .

Strigiphilus heterocerus (Grube, 1851): Hopkins & Clay, 1952: 339.

“ Strigiphilus laticephalus Złotorzycka, 1974: 324, fig. 48”. Not Philopterus cursor var laticephalus Uchida, 1949 . “ Strigiphilus laticephalus Adam & Daróczi, 2006: 157, figs 11c,d”. Not Philopterus cursor var laticephalus Uchida, 1949 . Strigiphilus heterocerus (Grube, 1851): Price et al. 2003: 239.

Diagnosis. Among the Japanese species of Strigiphilus, this species is easily identified by (1) a short preconal area of the head with anterior margin slightly concave (Figs 2, 5), (2) sexually dimorphic antennae (Figs 2, 5), (3) the shape of dorsal anterior plates (Figs 3, 6), and (4) the morphology of the male genitalia (Fig. 4).

Material examined. Type specimens. Ex Strix uralensis hondoensis: Syntypes of Strigiphilus fukuro: 1♂ (slide-mounted), Fukuro [= Strix uralensis hondoensis, in Katakana], Nagano Prefecture [in Kanji], Japan, 24 Dec. 1916 (ELKU); 2♂, 5♀, 9N (preserved in alcohol) same data (ELKU). Slide-mounted syntypes are labeled as “(1) Syntype / Strigiphilus fukuro / Uchida, 1948 / det. Shimada & Yoshi, 2020; (2) Nagano Pref. / (24. xii. 1916)/ ex Strix uralensis ” (ELKU); 1♀ (slide-mounted #010693510) (1) “ Strigiphilus fukuro Uchida, 1948 / T. Clay det./ PARA- TYPE ” [no holotype designation was made in the original description by Uchida (1948), therefore this specimen is actually a syntype]. (2) “ Strigiphilus fukuro n. sp. / Fukuro [in Katakana]/ Shinshu [in Kanji = Nagano Prefecture] 33” (NMHL). Data taken from NHML online database accessed on 23 Apr. 2020 —https://data.nhm.ac.uk

Non-type specimens. Ex Strix uralensis spp .: 23♂, 48♀, 1N (slide-mounted) Amakubo, Tsukuba City, Ibaraki Prefecture, Japan, 9 Feb. 2009, Haruhiko Asuka & Miyako Tsurumi (SEHU) ; 26♂, 89♀, 40N (preserved in alcohol) same data (YIO); 1♂, 1♀ (slide-mounted), Bibi, Chitose City, Hokkaido, Japan, 10 Mar. 2006, Miyako Tsurumi (SEHU) ; 4♂, 18♀, 23N (preserved in alcohol) same data (YIO) .

Japanese host: Strix uralensis hondoensis (Clark, 1907) .

Other hosts. Strix uralensis uralensis Pallas, 1771, Strix uralensis liturata (Tengmalm, 1795) .