Pachyula Ogilby, 1898

(Figs 1–3, 38F, 49)

Type species: Hemipimelodus crassilabris Ramsay & Ogilby, 1886 .

Diagnosis

Mesethmoid median portion very narrow (2, 2> 0); epioccipital visible dorsally (41, 0> 1); vomer anterior margin pronounced and acute (50, 0> 1); basioccipital lateral process absent with anterior and posterior portions equally developed (83, 1> 0); anteroventral portion of opercle subtrapezoidal, very short (127, 1> 2); metapterygoid one and a one-half times longer than deep in perpendicular section (135, 1> 2); posterolateral processes of urohyal short (160, 1> 0); third basibranchial chalice shaped (168, 0> 1); posterior margin of first epibranchial straight (181, 1> 0); first pharyngobranchial large and depressed (190, 0> 1); median crest associated with neural spine of fourth vertebra very high (202, 0> 1); Müllerian ramus robust (207, 0> 1); ventral surfaces of parapophyses of fifth and sixth vertebrae conspicuously concave (212, 0> 1); adipose fin-base long (222, 2> 1).

Ambiguous optimization: Mesethmoid medial notch narrow and deep (1, 0> 1); lateral horn of lateral ethmoid compressed and spatulate (11, 0> 1); Müllerianramusdistalone-thirdgentlycurved (208, 2> 1); posterior process of cleithrum very long (224, 0> 2).

Included species

Pachyula crassilabris Ramsay & Ogilby, 1886

Pachyula conorhynchus Weber, 1913 * sedis mutabilis.

Habitat and distribution: Freshwater, southern New Guinea (Fig. 38).

Remarks

Total-evidence analysis confirms the validity of this genus, in agreement with previous morphological and molecular studies(Marceniuk and Menezes 2007, Betancur-R.2009, Marceniuk et al. 2012).