Paracinetodus gen. nov.

(Figs 1–3, 38B, 50)

ZooBank registration: urn:lsid:zoobank.org:act: 01D775F1- CA33-42B6-AE05-AC1D3E9DBE6C.

Type species: Arius carinatus Weber, 1913 .

Diagnosis

Lateral horn of lateral ethmoid compressed and spatulate (11, 0> 1); extrascapular subtriangular (37, 1> 2); epioccipital contacting small narrow area of diagonal crest associated with neural spine of fourth vertebra (44, 2> 0); ventral tip of subvertebral process acute (80, 0> 2); basioccipital lateral process absent with anterior and posterior portions equally developed (83, 1> 0); transcapular process at right angle to body axis (86, 0> 1); transcapular process short and thick (87, 0> 2); space between transcapular process and otic capsule small (89, 1> 2); premaxilla narrow and long, length two to three times its width (120, 1> 2); bony blade connecting posterolateral processes of urohyal absent (159, 0> 1); posterolateral processes of urohyal two-thirds as long as distal portion of bone (162, 0> 2); transverse crest associated with neural spine of fourth vertebra high (200, 0> 1); median crest associated with neural spine of fourth vertebra high (202, 0> 1); Müllerian ramus bony blade evident in more than one-half length (206, 0> 2); ventral surfaces of parapophyses of fifth and sixth vertebrae conspicuously concave (212, 0> 1); seventh vertebra free from ventral superficial ossification (214, 0> 1); posterior process of cleithrum very long (224, 1> 2); second dorsal cleithral process dorsally directed and parallel to first dorsal process (226, 0> 1).

Etymology

Derived from the Greek ‘para’ for ‘near’, highlighting its morphological similarity with the genus Cinetodus . Gender: masculine.

Included species

Paracinetodus carinatus Weber, 1913 .

Habitat and distribution: Freshwater,southern New Guinea (Fig.38). Remarks

Arius carinatus was included in Cinetodus in previous morphological studies (Kailola 2004, Marceniuk and Menezes 2007, Marceniuk et al. 2012). The results of the total-evidence analysis place the species in a lineage distinct from that of Cinetodus, corroborating a previous molecular result (Betancur-R. 2009), and requiring the establishment of the new genus.