Doiichthys Weber, 1913

(Figs 1–3, 44)

Type species: Doiichthys novaeguineae Weber, 1913 .

Diagnosis

Lateral horn of lateral ethmoid long and posteriorly oriented (12, 1> 2); nasal not well defined (19, 0> 1); ventral crest of parieto-supraoccipital process weakly developed, restricted to base of process (48, 1> 0); three infraorbitals (95, 3> 1); articulation of autopalatine with lateral ethmoid slightly anterior to middle of bone (111, 2> 0); metapterygoid twice as deep as long in perpendicular section (135, 1> 0); first external branchiostegal ray proximally narrow and distally broad (147, 1> 0); posterior ceratohyal very long (153, 0> 2); urohyal very long (157, 1> 2); posterior end of urohyal bifurcate (158, 0> 1); urohyal posterolateral processes one-third as long as distal portion of bone (162, 2> 4); first and second epibranchials straight for entire length (178, 0> 1); mesial portion of first epibranchial very large and depressed (180, 0> 2); first pharyngobranchial on mesialendoffirstepibranchial (191, 1> 0); thirdpharyngobranchial funnel shaped (192, 1> 0); upper (pharyngeal) tooth plate long and narrow, its length three times its width (195,?> 2); transverse crest associated with neural spine of fourth vertebra low (200, 1> 0); median crest associated with neural spine of third vertebra low or absent (201, 1> 0).

Ambiguous optimization: Accessory tooth plates present (58, 0> 1).

Included species

Doiichthys novaeguineae Weber, 1913 .

Habitat and distribution: Brackish waters, southern New Guinea (Fig. 38).

Remarks

The total-evidence analysis supports a sister group relationship of Doiichthys plus Nedystoma, which was uncovered previously in independent molecular and morphological studies (Kailola 2004, Marceniuk and Menezes 2007, Betancur-R. 2009, Marceniuk et al. 2012). The synonymy of Doiichthys with Nedystoma proposed by Kailola (2004) is rejected and Doiichthys is recognized as a valid genus following Marceniuk and Menezes (2007) and Betancur-R. (2009).