Hemiarius Bleeker, 1862

MP 94, ML 91

(Figs 1–3, 30)

Type species: Cephalocassis stormii Bleeker, 1858 .

Diagnosis

Parasphenoid very wide in ventral view (66, 0> 1); dorsal crest on premaxilla beginning between lateral one-third or one-half of anterior margin (124, 0> 1); dorsal crest of hyomandibula short and high (141, 0> 1); third basibranchial chalice shaped (168, 0> 1); transverse crest associated with neural spine of fourth vertebra very high (200, 0> 1); median crest associated with neural spine of fourth vertebra very high (202, 0> 1).

Ambiguous optimization: Mesethmoid median portion moderately wide (2, 0> 1); gas bladder lateral diverticula shallow, rounded bulges (244, 0> 1).

Included species

Hemiarius bleekeri Popta, 1900 * sedis mutabilis

Hemiarius hardenbergi Kailola, 2000 * sedis mutabilis

Hemiarius harmandi Sauvage, 1880 * sedis mutabilis

Hemiarius manillensis Valenciennes, 1840 * sedis mutabilis Hemiarius stormii Bleeker, 1858

Hemiarius sumatranus Anonymous, 1830

Hemiarius verrucosus Ng, 2003 * sedis mutabilis.

Habitat and distribution: Predominantly brackish waters, South and Southeast Asia (Fig. 24).

Remarks

The total-evidence analysis supports the monophyly of Hemiarius as observed in previous morphological studies (Marceniuk and Menezes 2007, Marceniuk et al. 2012). The inclusion of Hemiarius in the subtribe Ariina is supported by the MP and ML analyses.