Eleutherodactylus verrucosus (Reinhardt and Lütken, 1862)

(Figs. 2–3)

Leiuperus verrucosus Reinhardt and Lütken, 1862 .

Ischnocnema verrucosa ­ Reinhardt and Lütken, 1862; Lynch, 1971, 1972; Sazima and Cardoso, 1978; Frost, 1985.

Paludicola verrucosa ­ Boulenger, 1882; Nieden, 1923.

Pleurodema verrucosa ­ Parker, 1927; Cochran, 1955; Bokermann, 1966; Gorham, 1966.

Epsophus verrucosus Miranda­Ribeiro, 1937 .

Eleutherodactylus verrucosus (Miranda­Ribeiro, 1937) ­ Bokermann, 1966. Eupsophus versus Gorham, 1966 [substitute name for Eupsophus verrucosus Miranda­Ribeiro, 1937, preoccupied in Eupsophus by Borborocoetes verrucosus Philippi (= Eupsophus nodosus Duméril and Bibron)].

Additionally, we examined and compared the specimens of Eleutherodactylus verrucosus with Eleutherodactylus juipoca Sazima and Cardoso (MNRJ 4103, holotype) and Eleutherodactylus octavioi Bokermann (MZUSP 73670, holotype), both species from Southeastern Brazil and thought to be related to E. verrucosus (Lynch, 1972; Sazima and Cardoso, 1978). The three species are perfectly distinguishable, and we consider them valid taxa.