Onthophagus (Onthophagus) transisthmius Howden & Young, 1981 Plate 44B

Onthophagus transisthmius Howden & Young, 1981: 106 (original description. Type locality: Panama. Canal Zone, Gamboa Limbo Hunt Club).

Onthophagus transisthmius: Zunino and Halffter 1997: 161 (list of species); Medina et al. 2001: 140 (list of species for Colombia); Ratcliffe 2002: 17 (cited for Panama); Rossini et al. 2018a: 547 (figures: 2a-d, 2h-i, 5b), 552-553 (characters in key), 563 (redescription), 564 (distribution); Delgado and Curoe 2014: 65 (characters in key, cited for Panama); Rossini et al. 2018b: 10 (list of species of the osculatii complex).

Onthophagus (Onthophagus) transisthmius: Chamorro et al. 2018: 97 (cited for Ecuador).

Types specimens.

Onthophagus transisthmius Howden & Young, 1981. The holotype is deposited at the USNM (see Howden and Young 1981: 107). Locality: Panama. Canal Zone, Gamboa Limbo Hunt Club (not examined).

Distribution.

Bolivia, Colombia, Ecuador, Panama, Peru, and Venezuela.

Records examined.

MORONA SANTIAGO: road Mendez-Paute km 8 (1 specimen CEMT). PASTAZA: plataforma Villano (2 specimens CEMT); Villano (4 specimens CEMT).

Literature records.

NAPO: Jatun Sacha Biological Station, 450 m (Rossini et al. 2018a: 566), same locality, 21 km E Puerto Napo, 400 m (Rossini et al. 2018a: 566); Tena, 400 m (Rossini et al. 2018a: 566). NAPO [= ORELLANA]: La Joya de los Sachis [= Joya de los Sachas], 290 m (Rossini et al. 2018a: 566). SUCUMBÍOS: Limoncocha, 250 m (Rossini et al. 2018a: 566). TUNGURAHUA: 6 km W de Río Negro, 1200 m (Rossini et al. 2018a: 566).

Temporal data.

Collected in January, June, July, and August.

Remarks.

Inhabits the lowland evergreen forests and evergreen foothill forests of the Amazon region from 290-1200 m a.s.l. Collected with pitfall traps baited with human feces.