Onthophagus (Onthophagus) coscineus Bates, 1887 Plate 40A

Onthophagus coscineus Bates, 1887: 79 (original description. Type locality: PANAMA, volcán de Chiriqui).

Onthophagus coscineus: Gillet 1911a: 205 (complete list of species); Boucomont and Gillet 1927: 205 (catalog of species); Boucomont 1932: 323 (distribution); Blackwelder 1944: 211 (list of species from Latin America); Howden and Young 1981: 95 (characters in key), 111 (redescription); Kohlmann and Solís 2001: 163 (characters in key), 187 (redescription); Medina et al. 2001: 139 (list of species for Colombia); Ratcliffe 2002: 17 (cited for Panama); Pulido-Herrera and Zunino 2007: 102 (cited as synonym of Onthophagus digitifer Boucomont, 1932); Carvajal et al. 2011: 322-323 (cited for Ecuador); Krajcik 2012: 177 (complete list of species); Solís and Kohlmann 2012: 8 (list of species from Costa Rica); Ratcliffe et al. 2015: 195 (cited for Peru); Delgado and Curoe 2014: 64 (characters in key, cited for Panama).

Onthophagus (Onthophagus) coscineus: Chamorro et al. 2018: 96 (cited for Ecuador).

Type specimens.

Onthophagus coscineus Bates, 1887. The lectotype is deposited at the NHML (see Kohlmann and Solís 2001: 187). Locality: V de Chiriquí 25-4000 feet [= 760-1220 m], not examined.

Distribution.

Colombia, Costa Rica, Ecuador, and Panama.

Records examined.

CARCHI: Tobar Donoso, 300 m (2 specimens MECN); ESMERALDAS: Gualpi (1 specimen CEMT); SANTO DOMINGO DE LOS TSÁCHILAS: Santo Domingo, Puerto Limón, 395 m (1 specimen CEMT).

Temporal data.

Collected in March, April, September, and October.

Remarks.

Inhabits coastal lowland evergreen forests from 300-395 m a.s.l. Collected with pitfall traps baited with human feces.