Lysmus Navás, 1910

(Figs 34–35)

(= Eosmylus Krüger, 1915: 73. Type species: Spilosmylus nigricornis Nakahara, 1914: 509)

(= Neolysmus Nakahara, 1955: 13 . Type species: Neolysmus ogatai Nakahara, 1955: 13)

Type species. Osmylus harmandinus Navás, 1910: 190 (by original designation).

Diagnosis. Three ocelli present; antennae less than FW length; wings ovoid, hyaline, FW sometimes with dark markings; FW costal area broadened basally, subcostal veinlets simple, not arranged sub-radially at base (sometimes basal 2–3 arranged divergently); RP stem short; 10–13 RP branches present; end-twigging of radial and medial veins along wing margin regular and one level; FW with stem of RP relatively short, RP1 arising proximal to M fork; RP in both wings with two distinct gradate series, subparallel in orientation, more basal crossveins semi irregular in arrangement, sometimes with third poorly defined gradate series; FW M vein fork near base of wing, between first and second branches of RP; FW CuP with more branches than CuA.

Comments. Lysmus was previously placed in Spilosmylinae, underscoring the difficulties in differentiating and classifying members of both subfamilies. Winterton et al. (2017) recovered this genus as the sister to Gryposmylus, and Winterton & Wang (2016) noted that the genera are differentiated by features such as the orientation of the basal subcostal veinlets and gradate series crossveins, and by the position of the M fork relative to the origin of RP1. Considering the variability of some venational features in Lysmus, this genus should be a priority for revision to clarify its status relative to other genera in the subfamily, especially genera such as Gryposmylus, Osmylidia and Protosmylus .

Included species.

L. beccarii (Navás, 1929) (Indonesia, Japan)

L. canimaculatus Jia & Liu, 2005 (China)

L. harmandinus (Navás, 1910) (Japan, Russia) (= Lysmus kurilensis Kuwayama, 1956; = Spilosmylus nigricornis Nakahara, 1914)

L. oberthurinus (Navás, 1910) (China)

L. ogatai (Nakahara, 1955) (Japan, China)

L. pallidius Yang & Liu, 2001 (China)

L. qingyuanus Yang et al., 1995 (China)

L. victus Yang, 1997 (China)

L. zanganus Yang et al. in Huang et al., 1988 (China)