identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
FE5387F7FFD6FFC500A220AB369FFA8F.text	FE5387F7FFD6FFC500A220AB369FFA8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alectorobius Pocock 1907	<div><p>Genus Alectorobius Pocock, 1907</p><p>Both literature surveys and the current study revealed the presence of a single species belonging to this genus, Alectorobius coniceps (Canestrini, 1890) (Fig. 1) (Ali et al. 2024a).</p></div>	https://treatment.plazi.org/id/FE5387F7FFD6FFC500A220AB369FFA8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFDCFFCF00A224963764FEFF.text	FE5387F7FFDCFFCF00A224963764FEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alectorobius coniceps (Canestrini 1890)	<div><p>Alectorobius coniceps (Canestrini, 1890)</p><p>Distribution</p><p>This tick has been found in the nests of swallows in Charsadda District (Ali et al. 2024a). In the current study this species was found in Mardan District.</p></div>	https://treatment.plazi.org/id/FE5387F7FFDCFFCF00A224963764FEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFDDFFCE00A2249637D0FE6F.text	FE5387F7FFDDFFCE00A2249637D0FE6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Argas	<div><p>Genus Argas</p><p>Argas spp. are found in many parts of Pakistan, including Punjab, Sindh, and KP. These ticks commonly infest domestic fowl and ducks, especially in sheds containing these birds. Four species of this genus, Argas abdussalami Hoogstraal and McCarthy, 1965, Argas persicus (Oken, 1818), Argas reflexus (Fabricius, 1794), and Argas sp. “ rousetti,” have been reported in Pakistan (Fig. 3) (Yasin &amp; Abdussalam 1958; Hoogstraal &amp; McCarthy 1965b; Buriro &amp; Akbar 1978; Hoogstraal 1985; Qamar et al. 2009; Karim et al. 2017; Ali et al. 2019, 2023c; Zahid et al. 2021). Morphological identification of our collections revealed the presence of Ar. persicus and yielded the first report of Argas hermanni Audouin, 1826 (Fig. 3).</p></div>	https://treatment.plazi.org/id/FE5387F7FFDDFFCE00A2249637D0FE6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFDDFFCE00A220BF3459FAD6.text	FE5387F7FFDDFFCE00A220BF3459FAD6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Argas hermanni Audouin 1826	<div><p>Argas hermanni Audouin, 1826</p><p>Distribution</p><p>Argas hermanni is here reported from Pakistan for the first time, our specimens having been collected in Buner District (Fig. 3).</p></div>	https://treatment.plazi.org/id/FE5387F7FFDDFFCE00A220BF3459FAD6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFDDFFCE00A223233761F84E.text	FE5387F7FFDDFFCE00A223233761F84E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Argas persicus (Oken 1818)	<div><p>Argas persicus (Oken, 1818)</p><p>Distribution</p><p>Argas persicus has been reported from Sindh, Punjab (Yasin &amp; Abdussalam 1958; Hoogstraal &amp; McCarthy 1965b; Buriro &amp; Akbar 1978; Ghosh et al. 2007; Qamar et al. 2009; Karim et al. 2017), and KP (Ali et al. 2019, 2023c; Zahid et al. 2021; Tsai et al. 2024).</p></div>	https://treatment.plazi.org/id/FE5387F7FFDDFFCE00A223233761F84E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFDFFFCC00A224963636FEDB.text	FE5387F7FFDFFFCC00A224963636FEDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carios Latreille 1796	<div><p>Genus Carios</p><p>Bat-associated Carios ticks have been reported from northern KP. In earlier studies, Carios ticks were reported infesting bats in Peshawar, Charsadda, Mardan, Hangu, Swabi, and Dir Lower District (Ullah H et al. 2019; Zahid et al. 2023), while in our investigation these ticks were found on bat hosts in Bajaur, Charsadda, and Mardan, based on morphological identification of our specimens (Fig. 6).</p></div>	https://treatment.plazi.org/id/FE5387F7FFDFFFCC00A224963636FEDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFDFFFCC00A220FF3163FA3A.text	FE5387F7FFDFFFCC00A220FF3163FA3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carios vespertilionis Latreille 1796	<div><p>Carios vespertilionis Latreille, 1796</p><p>Distribution This tick has been reported from Dir Lower (Ullah H et al. 2019; Zahid et al. 2023).</p></div>	https://treatment.plazi.org/id/FE5387F7FFDFFFCC00A220FF3163FA3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFD8FFCB00A227DE36EDFB48.text	FE5387F7FFD8FFCB00A227DE36EDFB48.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ornithodoros Koch 1837	<div><p>Genus Ornithodoros</p><p>Ornithodoros species have been reported from different localities in Shangla, Azad Jammu and Kashmir, Bajuar, and Orakzai, infesting Asian water buffaloes and their sheds in high mountainous regions. Three Ornithodoros species have been reported in Pakistan: Ornithodoros papillipes (Birula, 1895), Ornithodoros tholozani (Laboulbène and Mégnin, 1882), and possibly the new species Ornithodoros (Pavlovskyella) pakistanensis (Fig. 8) (Doss 1974; Hoogstraal 1985; Karim et al. 2017; Ali et al. 2022b, 2024c). Morphological identification of ticks collected during this survey confirmed the presence only of Or. (Pavlovskyella) pakistanensis .</p></div>	https://treatment.plazi.org/id/FE5387F7FFD8FFCB00A227DE36EDFB48	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFD9FFCA00A2232C31C2F84B.text	FE5387F7FFD9FFCA00A2232C31C2F84B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amblyomma	<div><p>Genus Amblyomma</p><p>Three species, Amblyomma gervaisi (Lucas, 1847), Amblyomma javanense (Supino, 1897) and Amblyomma pakhtunensis Ali, Khan, Chitimia–Dobler and Mans, 2024, have been reported infesting reptiles in Pakistan (Fig. 10) (McCarthy 1967; Auffenberg &amp; Auffenberg 1990; Ali et al. 2019, 2024b; Khan M et al. 2022). Morphological identification of our tick collections revealed the presence of Am. gervaisi and Am. pakhtunensis .</p></div>	https://treatment.plazi.org/id/FE5387F7FFD9FFCA00A2232C31C2F84B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFD9FFCA00A2249635F8FE85.text	FE5387F7FFD9FFCA00A2249635F8FE85.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ornithodoros pakistanensis Ali, Chitimia - Dobler, Munoz-Leal and Mans	<div><p>Ornithodoros pakistanensis Ali, Chitimia–Dobler, Muñoz-Leal and Mans</p><p>Distribution</p><p>This tick has been reported from Shangla, Dir Upper, Dir Lower, Orakzai, and Bajaur districts (Ali et al. 2022b, 2024c).</p></div>	https://treatment.plazi.org/id/FE5387F7FFD9FFCA00A2249635F8FE85	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFDAFFC900A227C537CEFBD1.text	FE5387F7FFDAFFC900A227C537CEFBD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amblyomma gervaisi (Lucas 1847)	<div><p>Amblyomma gervaisi (Lucas, 1847)</p><p>Distribution</p><p>This species is known from Lahore, Multan (McCarthy 1967; Auffenberg &amp; Auffenberg 1990), and central and northern KP (Ali et al. 2019; Khan, M. et al. 2022).</p></div>	https://treatment.plazi.org/id/FE5387F7FFDAFFC900A227C537CEFBD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFDBFFC800A227FA35FAFB1B.text	FE5387F7FFDBFFC800A227FA35FAFB1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amblyomma pakhtunensis Ali, Khan, Chitimia - Dobler and Mans 2024	<div><p>Amblyomma pakhtunensis Ali, Khan, Chitimia–Dobler and Mans, 2024</p><p>Distribution</p><p>This recently described species has been reported infesting Indian monitor lizards in Bajaur District (Ali et al. 2024b).</p></div>	https://treatment.plazi.org/id/FE5387F7FFDBFFC800A227FA35FAFB1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC5FFD600A22496349EFEFF.text	FE5387F7FFC5FFD600A22496349EFEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dermacentor montanus Filippova and Panova 1974	<div><p>Dermacentor montanus Filippova and Panova, 1974</p><p>Distribution</p><p>Dermacentor montanus is here reported for the first time from Pakistan, having been found infesting yaks ( Bos grunniens) in Chitral.</p></div>	https://treatment.plazi.org/id/FE5387F7FFC5FFD600A22496349EFEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC6FFD500A224963767FEFF.text	FE5387F7FFC6FFD500A224963767FEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dermacentor raskemensis Pomerantzev 1946	<div><p>Dermacentor raskemensis Pomerantzev, 1946</p><p>Distribution</p><p>Dermacentor raskemensis has been reported from Ziarat, Quetta (McCarthy 1967; Dhanda et al. 1971), Swat, and Sandman Tangi (Apanaskevich 2013).</p></div>	https://treatment.plazi.org/id/FE5387F7FFC6FFD500A224963767FEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC6FFD400A2234B34E4FD83.text	FE5387F7FFC6FFD400A2234B34E4FD83.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haemaphysalis C.L.Koch 1844	<div><p>Genus Haemaphysalis</p><p>Haemaphysalis ticks are distributed throughout the north and west mountainous regions of KP, Balochistan, and some parts of Punjab and Sindh. Previous studies have yielded a total of 12 haemaphysaline ticks reported from both wild and domestic animals in Pakistan: Haemaphysalis bispinosa Neumann, 1897, Haemaphysalis canestrinii (Supino, 1897), Haemaphysalis cornupunctata Hoogstraal and Varma, 1962, Haemaphysalis danieli Černý and Hoogstraal, 1977, Haemaphysalis erinacei Pavesi, 1884, Haemaphysalis howletti Warburton, 1913, Haemaphysalis indica Warburton, 1910, Haemaphysalis intermedia Warburton and Nuttall, 1909, Haemaphysalis kashmirensis Hoogstraal and Varma, 1962, Haemaphysalis kutchensis Hoogstraal and Trapido, 1963, Haemaphysalis montgomeryi Nuttall, 1912, and Haemaphysalis sulcata Canestrini and Fanzago, 1878 (Fig. 16) (Hoogstraal &amp; Varma 1962; Dhanda 1964; Hoogstraal &amp; McCarthy 1965a; Hoogstraal et al. 1966; McCarthy 1967; Hoogstraal 1970, 1971; Begum et al. 1970b; Robertson et al. 1970; Černý &amp; Hoogstraal 1977; Karim et al. 2017; Ali et al. 2019, 2023; Kamran et al. 2021; Khan Z et al. 2022; Alam et al. 2022; Hussain et al. 2023; Khan M et al. 2023). Both literature and the results of the present study indicate that haemaphysaline ticks chiefly infest small ruminants in Pakistan, while Kamran et al. (2021) rarely collected Ha. bispinosa from equine hosts in Balochistan. The present study revealed the presence of seven Haemaphysalis species: Ha. bispinosa, Ha. cornupunctata, Ha. danieli, Ha. indica, Ha. kashmirensis, Ha. montgomeryi, and Ha. sulcata .</p></div>	https://treatment.plazi.org/id/FE5387F7FFC6FFD400A2234B34E4FD83	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC7FFD400A2219C3701F92E.text	FE5387F7FFC7FFD400A2219C3701F92E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haemaphysalis bispinosa Neumann 1897	<div><p>Haemaphysalis bispinosa Neumann, 1897</p><p>Distribution</p><p>The reported territories for Ha. bispinosa are Azad Jammu and Kashmir (Karim et al. 2017), KP (Khan Z et al. 2022, 2024; Alam et al. 2022; Aneela et al. 2023; Tila et al. 2023; Ali et al. 2023c; Majid et al. 2023; Khan M et al. 2023), Punjab (Begum et al. 1970b; Hussain et al. 2023), Balochistan (Kamran et al. 2021), Khairpur and the Neelum Valley (McCarthy, 1967).</p></div>	https://treatment.plazi.org/id/FE5387F7FFC7FFD400A2219C3701F92E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC0FFD300A2214B3009F9A7.text	FE5387F7FFC0FFD300A2214B3009F9A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haemaphysalis cornupunctata Hoogstraal and Varma 1962	<div><p>Haemaphysalis cornupunctata Hoogstraal and Varma, 1962</p><p>Distribution</p><p>This species has been found in central KP, Saidu Sharif Swat, Gilgit, Kaghan valley and Azad Jammu and Kashmir (Hoogstraal &amp; Varma 1962; McCarthy 1967; Ghosh et al. 2007; Karim et al. 2017; Khan SM et al. 2023; Tila et al. 2023; Shehla et al. 2023; Ali et al. 2023c; Khan M et al. 2023), Punjab (Hussain et al. 2023).</p></div>	https://treatment.plazi.org/id/FE5387F7FFC0FFD300A2214B3009F9A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC1FFD200A2211D3701FA79.text	FE5387F7FFC1FFD200A2211D3701FA79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haemaphysalis danieli Cerny and Hoogstraal 1977	<div><p>Haemaphysalis danieli Černý and Hoogstraal, 1977</p><p>Distribution</p><p>This species has been reported from KP and Gilgit Baltistan (Černý &amp; Hoogstraal 1977; Ahmad et al. 2022; Ali et al. 2023c; Khan M et al. 2023).</p></div>	https://treatment.plazi.org/id/FE5387F7FFC1FFD200A2211D3701FA79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC2FFD100A220523476FB43.text	FE5387F7FFC2FFD100A220523476FB43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haemaphysalis indica Warburton 1910	<div><p>Haemaphysalis indica Warburton, 1910</p><p>Distribution</p><p>This species has been reported from KP (Ali et al. 2019, 2023a), Lahore and Hyderabad (McCarthy 1967; Hoogstraal 1970).</p></div>	https://treatment.plazi.org/id/FE5387F7FFC2FFD100A220523476FB43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC3FFD000A2206536A6FA8D.text	FE5387F7FFC3FFD000A2206536A6FA8D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haemaphysalis kashmirensis Hoogstraal and Varma 1962	<div><p>Haemaphysalis kashmirensis Hoogstraal and Varma, 1962</p><p>Distribution</p><p>Haemaphysalis kashmirensis has been reported from Azad Jammu and Kashmir (Karim et al. 2017), Naran, Shogran (Hoogstraal &amp; Varma 1962), Babusar, Saidu Sharif, Gilgit and central KP (McCarthy 1967; Begum et al. 1970b; Khan Z et al. 2022; Khan SM et al. 2023; Khan M et al. 2023).</p></div>	https://treatment.plazi.org/id/FE5387F7FFC3FFD000A2206536A6FA8D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFCCFFDF00A2204C37E0FA81.text	FE5387F7FFCCFFDF00A2204C37E0FA81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haemaphysalis montgomeryi Nuttall 1912	<div><p>Haemaphysalis montgomeryi Nuttall, 1912</p><p>Distribution</p><p>Haemaphysalis montgomeryi has been reported from Gilgit Baltistan (Karim et al. 2017), KP (Robertson et al. 1970; Ali et al. 2019, 2023c; Khan Z et al. 2022; Aneela et al. 2023; Tila et al. 2023; Khan SM et al. 2023; Majid et al. 2023; Khan M et al. 2023; Ullah S et al. 2024), Azad Kashmir (Hoogstraal et al. 1966), Punjab, Sindh, and northern areas (McCarthy 1967; Begum et al. 1970b).</p></div>	https://treatment.plazi.org/id/FE5387F7FFCCFFDF00A2204C37E0FA81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFCDFFDE00A2206636C3FA8B.text	FE5387F7FFCDFFDE00A2206636C3FA8B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Haemaphysalis sulcata Canestrini and Fanzago 1878	<div><p>Haemaphysalis sulcata Canestrini and Fanzago, 1878</p><p>Distribution</p><p>This species occurs in many regions, including KP (Ghafar et al. 2020a; Alam et al. 2022; Aneela et al. 2023; Tila et al. 2023; Shehla et al. 2023; Khan M et al. 2023; Khan Z et al. 2024), Azad Jammu and Kashmir (Karim et al. 2017), Balochistan, and Punjab (McCarthy 1967; Begum et al. 1970b).</p></div>	https://treatment.plazi.org/id/FE5387F7FFCDFFDE00A2206636C3FA8B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFCEFFDC00A227CA31FCFEFF.text	FE5387F7FFCEFFDC00A227CA31FCFEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalomma C.L.Koch 1844	<div><p>Genus Hyalomma</p><p>Hyalomma ticks are found in nearly all provinces of Pakistan. Among these ticks, Hy. anatolicum Koch, 1844, is the most prevalent species and has been reported from several hosts. Hyalomma schulzei Olenev, 1931, and Hyalomma turanicum Pomerantzev, 1946, were the least reported species in literature as well as in the present study. In Pakistan, 12 Hyalomma species have been reported on various hosts: Hyalomma aegyptium (Linnaeus, 1758), Hy. anatolicum, Hyalomma asiaticum Schulze and Schlottke, 1930, Hyalomma brevipunctata Sharif, 1928, Hyalomma dromedarii Koch, 1844, Hyalomma hussaini Sharif, 1928, Hyalomma impeltatum Schulze and Schlottke, 1930, Hyalomma isaaci Sharif, 1928, Hyalomma kumari Sharif, 1928, Hy. schulzei, Hyalomma scupense Schulze, 1919, and Hy. turanicum (Fig. 24) (Kaiser &amp; Hoogstraal 1964; McCarthy 1967; Begum et al. 1970b; Apanaskevich et al. 2008, 2009, 2010; Apanaskevich &amp; Horak 2008, 2009, 2010; Karim et al. 2017; Rehman et al. 2017; Ali et al. 2019, 2020, 2021; Kasi et al. 2020; Ghafar et al. 2020b; Kamran et al. 2021; Alam et al. 2022; Khan Z et al. 2022; Zeb et al. 2022; Hussain et al. 2023; Ullah S et al. 2023, 2024; Aneela et al. 2023; Tila et al. 2023; Shehla et al. 2023; Khan M et al. 2023). Our tick collections confirmed the presence of eight Hyalomma species: Hy. anatolicum, Hy. asiaticum, Hy. dromedarii, Hy. isaaci, Hy. kumari, Hy. schulzei, Hy. scupense, and Hy. turanicum .</p></div>	https://treatment.plazi.org/id/FE5387F7FFCEFFDC00A227CA31FCFEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFCFFFDC00A225933173FD17.text	FE5387F7FFCFFFDC00A225933173FD17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalomma anatolicum Koch 1844	<div><p>Hyalomma anatolicum Koch, 1844</p><p>Distribution</p><p>This species is found commonly throughout the country (Kaiser &amp; Hoogstral 1964; McCarthy 1967), especially in Punjab (Sajid et al. 2011; Rehman et al. 2017; Karim et al. 2017; Ghafar et al. 2020b; Hussain et al. 2023), KP (Zeb et al. 2019; Ali et al. 2019, 2021, 2023c; Khan Z et al. 2022; Alam et al. 2022; Zeb et al. 2022; Aneela et al. 2023; Tila et al. 2023; Shehla et al. 2023; Khan M et al. 2023; Khan Z et al. 2024; Ullah S et al. 2024), Sindh (Karim et al. 2017), and Balochistan (Karim et al. 2017; Kasi et al. 2020; Kamran et al. 2021).</p></div>	https://treatment.plazi.org/id/FE5387F7FFCFFFDC00A225933173FD17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC8FFDB00A225FF36AFFD17.text	FE5387F7FFC8FFDB00A225FF36AFFD17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalomma asiaticum Schulze and Schlottke 1929	<div><p>Hyalomma asiaticum Schulze and Schlottke, 1929</p><p>Distribution</p><p>Hyalomma asiaticum is known from Balochistan, KP, and Punjab (Kaiser &amp; Hoogstraal 1964; McCarthy 1967; Begum et al. 1970b; Apanaskevich &amp; Horak 2010; Ullah S et al. 2024).</p></div>	https://treatment.plazi.org/id/FE5387F7FFC8FFDB00A225FF36AFFD17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFC9FFDA00A22647319FFC87.text	FE5387F7FFC9FFDA00A22647319FFC87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalomma dromedarii Koch 1844	<div><p>Hyalomma dromedarii Koch, 1844</p><p>Distribution</p><p>This species is found in Azad Jammu and Kashmir (Kaiser &amp; Hoogstraal 1964; McCarthy 1967; Begum et al. 1970b; Karim et al. 2017), Punjab (Ghosh et al. 2007; Hussain et al. 2023), KP (Siddiqi &amp; Jan 1986; Khan Z et al. 2022; Alam et al. 2022; Tila et al. 2023; Shehla et al. 2023; Ali et al. 2023c; Khan M et al. 2023; Ullah S et al. 2024), Sindh (Gadahi et al. 2013), and Balochistan (Kasi et al. 2020; Kamran et al. 2021).</p></div>	https://treatment.plazi.org/id/FE5387F7FFC9FFDA00A22647319FFC87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFCAFFD900A227433422FBAF.text	FE5387F7FFCAFFD900A227433422FBAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalomma isaaci Sharif 1928	<div><p>Hyalomma isaaci Sharif, 1928</p><p>Distribution</p><p>This species has been found in several regions of the country, including Balochistan, Punjab, and KP (McCarthy 1967; Apanaskevich &amp; Horak 2008; Karim et al. 2017; Alam et al. 2022; Hussain et al. 2023; Tila et al. 2023; Ullah S et al. 2024).</p></div>	https://treatment.plazi.org/id/FE5387F7FFCAFFD900A227433422FBAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFCBFFD800A2277F34B3FBF3.text	FE5387F7FFCBFFD800A2277F34B3FBF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalomma kumari Sharif 1928	<div><p>Hyalomma kumari Sharif, 1928</p><p>Distribution</p><p>This species has been reported from several regions of Pakistan (Apanaskevich et al. 2009), including the northern areas (Karim et al. 2017), KP (Ullah S et al. 2023; Khan M et al. 2023; Khan Z et al. 2024) and Kirthar range (McCarthy 1967).</p></div>	https://treatment.plazi.org/id/FE5387F7FFCBFFD800A2277F34B3FBF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFF4FFE700A227C93635FBCC.text	FE5387F7FFF4FFE700A227C93635FBCC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalomma schulzei Olenev 1931	<div><p>Hyalomma schulzei Olenev, 1931</p><p>Distribution</p><p>Hyalomma schulzei has been reported from many parts of Balochistan, including Quetta, Chagi, Kalat, Nushki and Gulistan (McCarthy 1967; Apanaskevich et al. 2008).</p></div>	https://treatment.plazi.org/id/FE5387F7FFF4FFE700A227C93635FBCC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFF5FFE600A220C9319FFA29.text	FE5387F7FFF5FFE600A220C9319FFA29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalomma scupense Schulze 1919	<div><p>Hyalomma scupense Schulze, 1919</p><p>Distribution</p><p>This tick has been reported in several regions of the country: Balochistan, Punjab, KP (McCarthy 1967; Kasi et al. 2020; Ghafar et al. 2020b; Alam et al. 2022; Hussain et al. 2023; Tila et al. 2023; Khan M et al. 2023; Ullah S et al. 2024), northern areas (Begum et al. 1970b), and Gilgit Baltistan (Karim et al. 2017).</p></div>	https://treatment.plazi.org/id/FE5387F7FFF5FFE600A220C9319FFA29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFF6FFE500A220AC3715FAED.text	FE5387F7FFF6FFE500A220AC3715FAED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hyalomma turanicum Pomerantzev 1946	<div><p>Hyalomma turanicum Pomerantzev, 1946</p><p>Distribution</p><p>This species has been found in Punjab, Balochistan and KP (Kaiser &amp; Hoogstraal 1964; McCarthy 1967; Majid et al. 2023; Ullah S et al. 2024).</p></div>	https://treatment.plazi.org/id/FE5387F7FFF6FFE500A220AC3715FAED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFF7FFE400A2277C34DBFBD1.text	FE5387F7FFF7FFE400A2277C34DBFBD1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixodes	<div><p>Genus Ixodes</p><p>Four species of the genus Ixodes have been reported infesting rodents and small ruminants in Pakistan’s northern areas. These species are Ixodes hyatti Clifford, Hoogstraal and Kohls, 1971, Ixodes kashmiricus Pomerantzev, 1948, Ixodes redikorzevi Olenev, 1927, and Ixodes stromi Filippova, 1957 (Fig. 33) (McCarthy 1967; Begum et al. 1970a, b; Clifford et al. 1971; Numan et al. 2022, 2023; Hussain et al. 2023; Guglielmone et al. 2023). This tick survey yielded only I. kashmiricus .</p></div>	https://treatment.plazi.org/id/FE5387F7FFF7FFE400A2277C34DBFBD1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFF0FFE300A2249637BBFEFF.text	FE5387F7FFF0FFE300A2249637BBFEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ixodes kashmiricus Pomerantzev 1948	<div><p>Ixodes kashmiricus Pomerantzev, 1948</p><p>Distribution</p><p>This species has been reported infesting goats and sheep in Shangla District, KP and Murree District, Punjab (Numan et al. 2022, 2023; Hussain et al. 2023).</p></div>	https://treatment.plazi.org/id/FE5387F7FFF0FFE300A2249637BBFEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFF0FFE300A223123749F85C.text	FE5387F7FFF0FFE300A223123749F85C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nosomma Schulze 1919	<div><p>Genus Nosomma</p><p>Nosomma is represented in Pakistan by a single species, Nosomma monstrosum (Nuttall and Warburton, 1908), which infests Asian buffaloes ( Bubalus bubalis). In a previous study, this species was reported in Haripur District, KP, northern Pakistan (Fig. 35) (Aiman et al. 2022). Morphological identification of the current tick collection revealed the presence of N. monstrosum .</p></div>	https://treatment.plazi.org/id/FE5387F7FFF0FFE300A223123749F85C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFF1FFE200A2208335F4FA95.text	FE5387F7FFF1FFE200A2208335F4FA95.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nosomma monstrosum (Nuttall and Warburton 1908)	<div><p>Nosomma monstrosum (Nuttall and Warburton, 1908)</p><p>Distribution</p><p>This species was first reported in Pakistan on Asian water buffaloes in Haripur District, KP (Aiman et al. 2022).</p></div>	https://treatment.plazi.org/id/FE5387F7FFF1FFE200A2208335F4FA95	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFF2FFE000A227E03672FF6B.text	FE5387F7FFF2FFE000A227E03672FF6B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhipicephalus Koch 1844	<div><p>Genus Rhipicephalus</p><p>Rhipicephalus spp. are distributed in all regions of Pakistan. Rhipicephalus microplus (Canestrini, 1888) is the most frequently reported tick species at all localities. Five Rhipicephalus species have been found to infest various vertebrate hosts in Pakistan: Rhipicephalus annulatus (Say, 1821), Rhipicephalus haemaphysaloides Supino, 1897, Rh. microplus, Rhipicephalus ramachandrai Dhanda, 1966, and Rhipicephalus turanicus Pomerantzev, 1940 (Fig. 37) (McCarthy 1967; Begum et al. 1970b; Siddiqi &amp; Jan 1986; Karim et al. 2017; Rehman et al. 2017; Cabezas-Cruz et al. 2019; Kasi et al. 2020; Ghafar et al. 2020b; Kamran et al. 2021; Ali et al. 2021, 2022, 2023b, c,; Khan Z et al. 2022; Alam et al. 2022; Zeb et al. 2022; Hussain et al. 2023; Aneela et al. 2023; Tila et al. 2023; Shehla et al. 2023; Khan M et al. 2023; Ullah S et al. 2024). This survey revealed the presence of two Rhipicephalus species: Rh. haemaphysaloides and Rh. microplus . Additionally, Rh. sanguineus (Latreille, 1806) has been reported in several previous Pakistan studies (Cabezas-Cruz et al. 2019; Ali et al. 2019, 2021, 2022a; Alam et al. 2022; Khan Z. et al 2022; Tila et al. 2023; Shehla et al. 2023). However, in their analysis of the geographical distribution of the hard ticks of the world, Guglielmone et al. (2023) do not list Rh. sanguineus as a species that occurs in Pakistan, pointing out that there are many populations of Rh. sanguineus sensu lato worldwide, some of which surely belong to Rh. sanguineus sensu stricto, as defined by Nava et al. (2018), whose molecular data will be essential to diagnosing these populations. In an attempt to confirm the presence of Rh. sanguineus, extensive sequencing and phylogenetic analyses were carried out based on several mitochondrial genes, which revealed that our species exactly matches an undetermined species that was categorized as Rhipicephalus sp. III by Dantas-Torres et al. (2013), whose sequences closely match Rhipicephalus sp. III from our study, with 99.9% identity for 12S rRNA and 100% for 16S rRNA. However, an evaluation of sp. III’s phylogenetic status shows a separate clade distinct from R. turanicus and Rh. sanguineus . In Pakistan Rh. turanicus has been recorded from hosts in KP (Ali et al. 2019, 2021, 2022a; Khan Z. et al. 2022; Alam et al. 2022; Aneela et al. 2023; Tila et al. 2023; Shehla et al. 2023; Ali et al. 2023b; Khan M et al. 2023; Ullah S et al. 2024), Balochistan (Kasi et al. 2020), Punjab (Rehman et al. 2017; Hussain et al. 2023), Gilgit-Baltistan, and Azad Jammu and Kashmir (McCarthy 1967; Karim et al. 2017). Again, however, Guglielmone et al. (2023) do not list Rh. turanicus as a species that occurs in Pakistan, noting that this species is taxonomically complex, and that one lineage, named Rh. turanicus sensu stricto by Bakkes et al. (2020), was described from two specimens collected on a dog at an unknown location in Central Asia, whereas Filippova (2008) stated that the type host of Rh. turanicus is Ovis aries (sheep) and the type locality is Tashkent, Uzbekistan. Clearly, morphological and molecular data from the type host at the type locality will be needed before any definitive statement can be made concerning the distribution of Rh. turanicus outside Uzbekistan.</p></div>	https://treatment.plazi.org/id/FE5387F7FFF2FFE000A227E03672FF6B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFF3FFE000A227EC35F4FB61.text	FE5387F7FFF3FFE000A227EC35F4FB61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhipicephalus haemaphysaloides Supino 1897	<div><p>Rhipicephalus haemaphysaloides Supino, 1897</p><p>Distribution</p><p>This species has been reported in Sindh, Balochistan (McCarthy 1967; Kamran et al. 2021), Punjab (Hussain et al. 2023; Abbas et al. 2025), KP (Ali et al. 2019, 2021, 2022a, b, c; Khan Z et al. 2022; Alam et al. 2022; Tila et al. 2023; Shehla et al. 2023; Khan M et al. 2023), and Gilgit Baltistan, Azad Jammu and Kashmir (Karim et al. 2017).</p></div>	https://treatment.plazi.org/id/FE5387F7FFF3FFE000A227EC35F4FB61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
FE5387F7FFFCFFEF00A227C035F4FB69.text	FE5387F7FFFCFFEF00A227C035F4FB69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhipicephalus microplus (Canestrini 1888)	<div><p>Rhipicephalus microplus (Canestrini, 1888)</p><p>Distribution</p><p>This tick has been reported from Sindh, KP, Balochistan, Punjab, and Azad Jammu and Kashmir (McCarthy 1967; Siddiqi &amp; Jan 1986; Karim et al. 2017; Rehman et al. 2017; Zeb et al. 2019, 2022; Ali et al. 2019, 2021–22a, 2023b, c; Nasreen et al. 2020; Kasi et al. 2020; Ghafar et al. 2020b; Kamran et al. 2021; Khan Z. et al. 2022, 2024; Alam et al. 2022; Hussain et al. 2023; Aneela et al. 2023; Tila et al. 2023; Shehla et al. 2023; Khan M et al. 2023).</p></div>	https://treatment.plazi.org/id/FE5387F7FFFCFFEF00A227C035F4FB69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ali, Abid;Almutairi, Mashal M.;Robbins, Richard G.;Chitimia-Dobler, Lidia;Ullah, Shafi	Ali, Abid, Almutairi, Mashal M., Robbins, Richard G., Chitimia-Dobler, Lidia, Ullah, Shafi (2025): Updated checklist, morphological descriptions, hosts and vector potential of ticks (Acari: Argasidae, Ixodidae) in Pakistan. Zootaxa 5725 (2): 151-202, DOI: 10.11646/zootaxa.5725.2.1, URL: https://doi.org/10.11646/zootaxa.5725.2.1
