taxonID	type	description	language	source
FB54535042732A23F2BB2B0CE2E3FCE1.taxon	diagnosis	Diagnosis Lateral pronotal collar usually dentate. Wings hyaline. Timbal covers well developed, with the exception of Sinosemia Matsumura, 1927 stat. rev., which has narrowed timbal covers widely exposing timbals medially. Male abdominal sternites translucent if not coated with dark-coloured pigmentation (as also seen in sister tribes like Cicadini Batsch, 1789, Leptopsaltriini Moulton, 1923 and Psithyristriini Distant 1905). Male abdominal sternites without bumps or tubercle-like projections. Uncus usually bifurcate, but in a few cases, fused and developed to a large uncus covering ventral side of pygofer. Aedeagus thin. Basal lobe of pygofer absent or indistinct, with a few exceptions in Biura, Songga and Cantata. See also Lee and Emery (2014) and Pham and Lee (2021) for the tribal diagnosis. Included subtribes (4)	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042732A22F2B02948E0CCFEFB.taxon	diagnosis	Diagnosis Pronotal collar narrow posteriorly. Fore wing usually with infuscations on radial and radiomedial crossveins, with the exception of Meimuna sensu stricto which has none. Male operculum basically obconical or up-triangular, with length about or less than two times the width, with exceptions of Meimuna sensu stricto, a species group in Meimuna and Cantata. Male abdomen obconical. Uncus short and bifurcate. See also Lee and Emery (2014) and Pham and Lee (2021) for subtribal diagnosis. Previous inclusions By the actions of Wang et al. (2021), Aolina included nine genera: Biura Lee and Sanborn, 2015, Cantata Lee and Pham, 2021, Haphsa Distant, 1905, Kaphsa Lee, 2012, Khimbya Distant, 1905, Meimuna Distant, 1905, Sinapsaltria Kato, 1940, Songga Lee, 2016 and Zaphsa Lee and Emery, 2014. Taxonomic changes in the present paper	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042732A22F2B02948E0CCFEFB.taxon	description	Five out of the above 13 genera, Aola, Changa, Haphsa, Sinosemia and Sinotympana are referred to here as the ‘ Haphsa - like genera’, which commonly have the following characteristics, different from other aoline genera: uncus short and broad, with uncal lobes fused at base, bifurcate distally; and basal lobes of pygofer absent.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042722A22F2CA2BB4E700FA49.taxon	diagnosis	Diagnosis Body medium-sized (body length around 29 mm). Head including compound eyes wider than mesonotum. Postclypeus well prominent, with symmetrical series of fuscous fasciae along transverse grooves. Pronotum about as long as head and distinctly shorter than mesonotum excluding cruciform elevation. Lateral pronotal collar dentate. Mesonotum greenish or ochraceous with black marks. Wings hyaline, with eight and six apical cells on fore wing and hind wing, respectively. Fore wing with apical cells in short length. Fore wing with infuscations on radial and radiomedial crossveins. Male operculum short and wide, wider than long, extending slightly beyond posterior margin of abdominal sternite II. Male opercula slightly separated. Male abdomen about as long as head and thorax together. Male abdominal tergite 3 wider than mesonotum. Timbal cover wider than long, completely concealing timbal in dorsal view, with roundly extended posterolateral corner. Male pygofer obovate in ventral view. Clasper and upper lobes of pygofer absent. Uncus thick and short, with uncal lobes fused at base, bifurcate distally; distally hooked inwardly in lateral view. Basal lobe of pygofer absent, but a pair of basal-lobe-like structures developed, like thin pieces of paper. Aedeagus thin, protruding from underside of roof-like structure of uncus. Species included	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042722A22F2CA2BB4E700FA49.taxon	description	Aola bindusara (Distant, 1881) comb. rev.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042722A22F2CA2BB4E700FA49.taxon	discussion	Remarks Aola can be distinguished from other Haphsa - like genera by the following characteristics: postclypeus well prominent, making pronotum about as long as head; male operculum short, extending slightly beyond posterior margin of abdominal sternite II; male abdomen short; timbal cover completely concealing timbal, with roundly extended posterolateral corner; and a pair of basal-lobe-like structures protruding like thin pieces of paper.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042722A21F2042C19E166FC43.taxon	diagnosis	Diagnosis Body medium-sized (body length around 28 mm). Head including compound eyes wider than mesonotum. Postclypeus well prominent, with symmetrical series of fuscous fasciae along transverse grooves, sometimes indistinct centrally. Pronotum slightly longer than head and distinctly shorter than mesonotum excluding cruciform elevation. Lateral pronotal collar dentate. Mesonotum greenish or ochraceous with black marks. Wings hyaline, with eight and six apical cells on fore wing and hind wing, respectively. Fore wing with apical cells in short length. Fore wing with infuscations on radial and radiomedial crossveins. Male operculum short and comparatively narrow, longer than wide, extending slightly beyond posterior margin of abdominal sternite II. Male opercula widely separated. Male abdomen about as long as head and thorax together. Male abdominal tergite 3 wider than mesonotum. Timbal cover widely open dorsally. Male pygofer obovate in ventral view. Clasper and upper lobes of pygofer absent. Uncus thick and short, with uncal lobes fused at base, bifurcate distally; curved inwardly in lateral view. Basal lobe of pygofer absent, but a pair of basal-lobe-like structures developed, like ridges. Aedeagus thin, protruding between bases of uncal lobes. Species included	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042722A21F2042C19E166FC43.taxon	discussion	Remarks This genus can be distinguished from other Haphsa - like genera by the male opercula, which are widely separated, the timbal cover, which is widely open dorsally, and the ridge-like basal-lobe-like structure.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042712A20F2E92E28E75FFDF8.taxon	diagnosis	Diagnosis Body medium-sized (body length around 36 mm), but larger than other Haphsa - like genera. Head including compound eyes wider than mesonotum. Postclypeus moderately prominent, with symmetrical series of fuscous fasciae along transverse grooves. Pronotum much longer than head and distinctly shorter than mesonotum excluding cruciform elevation. Lateral pronotal collar dentate. Mesonotum mostly black with two pairs of thin longitudinal fasciae. Wings hyaline, with eight and six apical cells on fore wing and hind wing, respectively. Fore wing with apical cells in short length. Fore wing with infuscations on radial and radiomedial crossveins, and sometimes with roundish infuscation on each hind submargin of radius anterior 2, radius posterior, median 1 to 4 and cubitus anterior 1 veins. Male operculum short and wide, roughly triangular, about as long as wide, extending slightly beyond posterior margin of abdominal sternite III at longest. Male opercula slightly overlapped or slightly separated. Male abdomen longer than head and thorax together. Male abdominal tergite 3 distinctly wider than mesonotum. Timbal cover about as long as wide, fully concealing timbal in dorsal view. Male pygofer obovate in ventral view. Clasper and upper lobes of pygofer absent. Uncus large, with uncal lobes fused at base, bifurcate distally, protruding ventrally; hooked or curved inwardly in lateral view. Basal lobe of pygofer absent. Aedeagus thin, protruding between bases of uncal lobes. Species included	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042712A20F2E92E28E75FFDF8.taxon	discussion	Remarks This genus can be distinguished from other Haphsa - like genera by the following characteristics: body much larger; uncus in same breadth from base to apex and hooked at apex in lateral view; and timbal cover entirely concealing timbal in dorsal view.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042702A2FF2C428B8E283FF64.taxon	diagnosis	Diagnosis Body medium-sized (body length around 24 mm), but smaller than other Haphsa - like genera. Head including compound eyes wider than mesonotum. Postclypeus moderately prominent, without symmetrical series of fuscous fasciae along transverse grooves but with marks forming the shape of the letter X in general. Pronotum longer than head and distinctly shorter than mesonotum excluding cruciform elevation. Lateral pronotal collar dentate. Mesonotum greenish or ochraceous with black marks. Wings hyaline, with eight and six apical cells on fore wing and hind wing, respectively. Fore wing with apical cells in comparatively long. Fore wing with infuscations on radial and radiomedial crossveins. Male operculum longer than wide, about 1.5 times to twice as long as wide, reaching about middle of abdominal sternite IV. Male opercula separated. Male abdomen much longer than head and thorax together. Male abdominal tergite 3 slightly wider than mesonotum. Timbal cover about as long as wide, mostly concealing timbal but exposing some portion of inner and anterior parts of timbal in dorsal view. Male pygofer obovate in ventral view. Clasper and upper lobes of pygofer absent. Uncus thick and short, with uncal lobes fused at base, bifurcate and widely separated from each other distally in ventral view; distally hooked inwardly in lateral view. Basal lobe of pygofer absent. Aedeagus thin, protruding between bases of uncal lobes. Species included	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042702A2FF2C428B8E283FF64.taxon	description	Changa sita (Distant, 1881) comb. rev. and Changa jsguillotsi (Boulard, 2005) comb. rev.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB54535042702A2FF2C428B8E283FF64.taxon	discussion	Remarks This genus is distinguished from Haphsa by the small body size, more slender body (ratio of body length to head width about 3.20), long male abdomen, black to fuscous marks on the postclypeus, forming the shape of the letter X, long fore wing apical cells, small timbal cover and the shape of uncal lobes which are widely separated from each other at apices and are hooked inwardly.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB545350427F2A2FF29D2B3EE2F1F907.taxon	diagnosis	Diagnosis Body medium-sized (body length around 28 mm). Head including compound eyes slightly wider than mesonotum. Postclypeus moderately prominent, with symmetrical series of fuscous fasciae along transverse grooves. Pronotum much longer than head and distinctly shorter than mesonotum excluding cruciform elevation. Lateral pronotal collar dentate. Mesonotum mostly black with some greenish or ochraceous marks. Wings hyaline, with eight and six apical cells on fore wing and hind wing, respectively. Fore wing with apical cells in short lengths. Fore wing with infuscations only on radial and radiomedial crossveins. Male operculum long and wide, about twice as long as wide, almost reaching posterior margin of abdominal sternite V. Male opercula separated medially. Male abdomen slightly longer than head and thorax together. Male abdominal tergite 3 wider than mesonotum. Timbal cover about as long as wide, mostly concealing timbal but exposing a small portion of inner part of timbal in dorsal view. Male pygofer obovate in ventral view. Clasper and upper lobes of pygofer absent. Uncus large, with uncal lobes fused at base, bifurcate distally. Uncus seldom curved (not hooked), almost parallel to pygofer distally in lateral view, in nicomache. Basal lobe of pygofer absent. Aedeagus thin, protruding from underside of roof-like structure of uncus. Species included	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB545350427F2A2FF29D2B3EE2F1F907.taxon	description	Haphsa nicomache (Walker, 1850). Haphsa conformis Distant, 1917, Haphsa dianensis Chou et al., 1997 [‘ nomen dubium’, as indicated by Wang et al. (2021)], Haphsa durga (Distant, 1881) (Figure 2), Haphsa fratercula Distant, 1917, Haphsa karenensis Ollenbach, 1928, Haphsa lueta Wang et al., 2015, Haphsa nana Distant, 1913, Haphsa opercularis Distant, 1917 and the new species (Figure 1) described below are not regarded as congeneric in morphological as well as molecular phylogenetical views, but I refrain from erecting a new genus here, awaiting a further comprehensive morphological review with sufficient material.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB545350427F2A2FF29D2B3EE2F1F907.taxon	discussion	Remarks Haphsa sensu stricto can be distinguished from other Haphsa - like genera by the following characteristics: male operculum very long, almost reaching posterior margin of abdominal sternite V; uncus with basal roof-like structure; and uncal lobes not hooked inwardly in lateral view. One of the species placed in Haphsa, Haphsa sulaiyai (Boulard, 2005), is now transferred to Cantata to become Cantata sulaiyai (Boulard, 2005) comb. n., which has the peculiar characteristics of Cantata: basal lobes of pygofer present; uncal lobes long and slender; and male opercula long.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB545350427E2A2CF2892E8BE289FBEA.taxon	description	(Figure 1)	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB545350427E2A2CF2892E8BE289FBEA.taxon	materials_examined	Type material Holotype. Male, [THAILAND:] Thaïlande Nord / Cordillère du / Doi Mon Kia / Mars 2002 / Michel Boulard, rec. ’, ‘ Meimuna / durga (D.) / Michel Boulard det 2008 ’, ‘ 02. TH. CR. DMK. 01 ’, ‘ Haphsa new spp., FlaEnt / Young June LEE, 2009 / Voucher 1 / 4 ’.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB545350427E2A2CF2892E8BE289FBEA.taxon	etymology	Etymology The specific name is the Latin feminine adjective meaning ‘ short’, in reference to the comparatively short male abdomen.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB545350427E2A2CF2892E8BE289FBEA.taxon	description	Measurements of the holotype in millimetres (n = 1 male) Length of body: 28.3. Length of head and thorax together: 13.4. Length of abdomen: 14.9. Width of head including compound eyes: 9.3. Width of pronotum: 9.6. Width of mesonotum: 8.1. Width of abdominal tergite 3: 9.3. Length of fore wing: 37.9. Width of fore wing: 11.2. Fore wing span: 83.4.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB545350427E2A2CF2892E8BE289FBEA.taxon	diagnosis	Diagnosis This new species is similar to and has often been misidentified as Haphsa durga (Figure 2) but can be distinguished by the following characteristics: postclypeus with fasciae along a few anterior transverse grooves only (with fasciae along transverse grooves all the way down to posterior margin in durga); male operculum slightly convex at inner posterior margin (slightly concave in durga); male abdomen mostly ochraceous (mostly black in durga); male abdomen slightly longer than head and thorax together (much longer in durga); and uncal lobes broad, touched each other in about basal two-thirds, with apex triangular, angled laterad (more or less slender, touched each other in about basal one-third, with apex round, angled posteriad in durga), in ventral view. This new species can be distinguished from other species currently in Haphsa (H. conformis, H. fratercula, H. karenensis, H. lueta, H. nana and H. opercularis) by the postclypeus having fasciae along a few anterior transverse grooves only and mostly unmarked centrally and the peculiar shape of the uncal lobes as listed below.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
FB545350427E2A2CF2892E8BE289FBEA.taxon	description	Description of male (Figure 1) Head. Head including compound eyes wider than mesonotum. Head ochraceous (probably due to discolouration) with black to fuscous marks. Vertex with axe blade-shaped median mark enclosing ocelli, a pair of rectangular marks, with narrow extensions towards posterior margin of vertex, between compound eyes and postclypeus (or the axe blade-shaped median mark), and a pair of small spots on anterior corners of vertex (close to postclypeus). Postclypeus with fasciae along a few anterior transverse grooves and with mountain-shaped mark posteriorly. Anteclypeus black with median longitudinal ochraceous fascia being extended to both lateral sides. Lorum fuscous. Gena mostly brown with transverse black fascia between compound eye and base of antenna. Rostrum ochraceous but fuscous on apex; apex passing posterior margin of hind coxae. Thorax. Thorax ochraceous (probably due to discolouration) with black to fuscous marks. Inner area of pronotum with the following black to fuscous marks: a pair of median longitudinal fasciae, broadened at anterior ends; a pair of curved narrow fasciae along lateral margins of inner area of pronotum; and rather indistinct spots (near paramedian and lateral fissures) inside of inner area of pronotum. Pronotal collar with indistinct two pairs of spots at lateral inner corners. Anterolateral pronotal collar dentate. Mesonotum with the following marks: long longitudinal median fascia extending posteriad but not reaching anterior margin of cruciform elevation; a pair of small roundish spots enclosing scutal depressions; a pair of fasciae along parapsidal sutures; and a pair of longitudinally arranged three spots on lateral sigilla. Cruciform elevation with a pair of tiny spots near tip of anterior arms. Thoracic sternites including legs without distinct marks but with pigmented portions at tibiae, tarsi and claws. Wings. Wings hyaline but slightly tinged with brown. Fore wing with long apical cells compared to Haphsa nicomache. Fore wing venation orange basally but fuscous apically. Fore wing with infuscations on radial and radio-medial crossveins. Basal cell hyaline and slightly tinged. Basal membrane and hind wing jugum whitish grey. Operculum. Male operculum ochraceous with no marks; longer than wide, roughly downward-pointing triangular with inner margin concave basally and with rounded apex, reaching posterior margin of abdominal sternite V. Opercula slightly separated medially. Abdomen. Male abdomen ochraceous to brown, slightly longer than head and thorax together. Tergite 3 slightly narrower than pronotum. Tergite 1 with narrow black posterior margin. Tergite 2 and timbal covers without distinct marks. Tergite 3 with narrow black anterior margin, not extending to lateral margin. Tergites 4, 5, 6 and 7, each with a pair of median fuscous spots anteriorly. Tergite 8 with narrow black anterior margin. Timbal cover about as long as wide, with apex widely rounded, mostly concealing timbal but exposing a small portion of inner part of timbal in dorsal view. Abdominal sternites ochraceous to brown without marks. Genitalia. (Figure 1 (C, D )) Pygofer obovate in ventral view. Distal shoulder not developed. Uncus with bifurcate, broad uncal lobes in ventral view, apex triangular, angled laterad; curved inwardly in lateral view. Uncal lobes touching each other in about basal two-thirds. Aedeagus thin.	en	Lee, Young June (2025): A new species of Haphsa Distant (Hemiptera: Cicadidae) from Northern Thailand, with a taxonomic review of Haphsa and its allied genera. Journal of Natural History 59 (13 - 16): 1049-1061, DOI: 10.1080/00222933.2025.2474200, URL: https://doi.org/10.1080/00222933.2025.2474200
