taxonID	type	description	language	source
F17D7D2D123FFF815584FDCFFBBFFA3F.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 3 C 4 F 617 C- 227 F- 4 A 85 - B 385 - 0 C 1 D 2324 E 5 BE (Figs 1 A – C)	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123FFF815584FDCFFBBFFA3F.taxon	materials_examined	Type material (2 ♂♂, 4 ♀♀). Holotype: BRAZIL: ♂, labeled: “ Brasil: AM, Itapiranga | Faz. Caribi - Madeira podre | 16. xii. 2019 | 2 ° 32 ’ 55,74 ’’ S / 58 ° 48 ’ 24,21 ’’ W | L. Klemann-Junior; R. S. Moura | CESIT - UEA ”, “ HOLOTYPE | Glyptoma | curupira sp. nov. | Desig. Klemann-Junior et al. 2024 ” (INPA). Paratypes (1 ♂, 4 ♀♀), labeled: “ Brasil: AM, Itacoatiara | Faz. Imperial – Manual | 02. xii. 2019 – Mad. Podre | 3 ° 00 ’ 46,27 ’’ S / 58 ° 28 ’ 44,50 ’’ W | L. Klemann-Junior; R. S. Moura | CESIT - UEA ” (1 ♀ UEA); “ Brasil: AM, Itapiranga | Faz. Caribi - Madeira podre | 16. xii. 2019 | 2 ° 32 ’ 55,74 ’’ S / 58 ° 48 ’ 24,21 ’’ W | L. Klemann-Junior; R. S. Moura | CESIT - UEA ” (1 ♂ UEA, 1 ♀ INPA); “ Brasil: AM, Silves | Faz. Caribi - Madeira podre | 16. xii. 2019 | 2 ° 32 ’ 55,74 ’’ S / 58 ° 48 ’ 24,21 ’’ W | R. S. Moura; L. F. B. Pereira | CESIT - UEA ” (1 ♀ UFPR); “ Brasil: AM, Itapiranga | Faz. Caribe - Col. Manual (Mad. podre) | 17. xii. 2019 | 2 ° 32 ’ 50,48 ’’ S / 58 ° 48 ’ 23, 16 ’’ W | L. Klemann-Junior; R. S. Moura | CESIT - UEA ” (1 ♀ CEMT). All paratypes with yellow label “ PARATYPE | Glyptoma | curupira sp. nov. | Desig. Klemann-Junior et al. 2024 ”. Labels abbreviations, translations, and explanations: AM = Amazonas state; CESIT - UEA = Centro de Estudos Superiores de Itacoatiara - Universidade do Estado do Amazonas = written in a vertical line on the left side; Col. Manual (Mad. podre) = Coleta Manual (Madeira Podre) = captured manually under bark in decaying trunks and branches; Faz. = Fazenda = farm; Mad. Podre = Madeira podre = captured manually under bark in decaying trunks and branches; Manual = captured manually.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123FFF815584FDCFFBBFFA3F.taxon	diagnosis	Diagnosis. Among species of the G. cicatricosa group (with size ≥ 3.5 mm and relatively long antenna) G. curupira sp. nov. is similar to G. opacum in size and the totally matt surface. However, G. curupira can be differentiated from G. opacum by the size of the punctures of the head, pronotum and elytra, size of the interstices between the punctures, by the shapes of the antennomeres, and by the distinctive shape of the aedeagus. In G. curupira the size of the punctures on the head and pronotum are subequal to punctures on the elytra (the largest punctures on the head and pronotum are the same size as the smallest punctures on the elytra); in G. opacum punctures on the elytra are much larger than those on the head and pronotum. In G. curupira punctures on clypeus smaller and sparser than on vertex; in G. opacum the clypeus is more sparsely punctate and the punctures equal in size to those of the vertex. In G. curupira the interstices between the punctures on the head, pronotum, and elytra vary from 1 / 2 to 1 / 5 as wide as the diameter of the punctures; in G. opacum the interstices between the punctures are less than 1 / 4 as wide as the diameter of the punctures on the head, the interstices are indistinct and irregular on the elytra. In G. curupira, antennomeres 5 – 10 are slightly wider than long, the length of the setae are about 0.9 the width of the antennomeres; in G. opacum, antennomeres 5 - 10 are quadrate or slightly longer than wide, and the setae are half as long as the width of the antennomeres. The apical part of the median lobe of the aedeagus of G. curupira broader (in lateral view) than the apical part of the median lobe of G. opacum. The median lobe of G. curupira has sensilla; sensilla are absent in G. opacum. The paremeres of the aedeagus of G. curupira are strongly curved in lateral view and without sensilla; the parameres are straight and has two sensilla at the base in G. opacum.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123FFF815584FDCFFBBFFA3F.taxon	description	Description. Holotype male, BL: 4.0 mm, FL: 2.01 mm. Coloration: dark reddish brown (almost black); antennae dark reddish brown (almost black); legs light reddish brown. Head (Figure 1 A) wider (HW: 0.64) than long (HL: 0.54); subrectangular; with rounded hind and anterior angles; large eyes slightly prominent and longer than temples; sides in front of eyes parallel up to base of antennae; at base of antennae with almost rectangular angle; anterior edge of clypeus as wide as half of width between base of antennae. Head with large, dense and deep setiferous punctures; with some coalescent punctures without evident ridges; each puncture with a yellow seta; interstices between punctures 1 / 2 to 1 / 5 as wide as diameter of punctures. Integument of epicranium, including surface of punctures, covered by dense microsculpturing formed by uniformly distributed micropunctures, appearing matt. Punctures on clypeus smaller and sparser than on vertex, surface matt. Neck appearing matt due to dense microsculpturing formed by uniformly distributed micropunctures; microsculpture on neck larger than microsculpture on head, pronotum, elytra, and abdomen. Antenna (Figure 1 B) slightly longer than head, pronotum and half of elytra together; antennomeres 2 – 10 gradually and smoothly widening anteriorly; antennomere 1 subrectangular, long and thick; antennomere 2 narrower (0.8) and shorter (0.45) than antennomere 1; antennomeres 3 – 4 wider than long, wider than antennomere 2 and narrower than antennomere 1; antennomeres 2 – 10 slightly wider than long; antennomere 11 narrower and longer than antennomere 10; all antennomeres with numerous long yellow setae; setae slightly shorter (0.9) as width of antennomeres. Pronotum (Figure 1 A) quadrate (PL: 0.60; PW: 0.60); widest at anterior third; anterior lateral margins convergent to widely rounded anterior angles in smooth convex curve; posterior lateral margins convergent to obtuse posterior angles in smooth concave curve; anterior margin straight. Pronotum with large, dense and deep setiferous punctures; some coalescent punctures without evident ridges; each puncture with a yellow seta; punctures as dense and deep as punctures on head; interstices between punctures 1 / 2 to 1 / 5 as wide as diameter of punctures. Median carina in wide depression; carina well developed for most of length, absent or weak anteriorly. Integument of pronotum, including surface of punctures, with similar dense microsculpturing formed by uniformly distributed micropunctures as head, appearing matt. Elytra (Figure 1 A) wider than pronotum, longer than wide (EL: 0.87; EW: 0.82); sides slightly convex; widest at middle; shoulders well developed, smoothly curved and shortly denticulate; posterior angles rounded; humeral angles of elytra strongly extended anteriorly. Elytra with large, dense and deep setiferous punctures; smaller punctures of elytra about equal to larger punctures on head and pronotum; punctures each with a yellow seta; setae in asymmetric position to center of punctures; interstices of punctures 1 / 2 to 1 / 4 as wide as diameter of punctures. Discal carina distinct; curved outward, following the lateral margin of elytra. Lateral carina indistinctly restricted shortly behind shoulders. Suture marked by line of small, shallow and coalescent punctures along entire length of elytra. Space between suture and discal carina with presence of three irregular and parallel rows of setiferous punctures along entire length of elytra; with some coalescent punctures without evident ridges. Laterally to discal carina another two irregular and parallel rows of setiferous punctures along entire length of elytra; submarginal surface with two rows of slightly smaller setiferous punctures. Integument of elytra, including carina and surface of punctures, with similar dense microsculpturing formed by uniformly distributed micropunctures as head and pronotum, appearing matt. Legs (Figure 1 A) almost completely covered with short yellow setae; surface appearing matt, with microridges organized with cells. Abdomen with dense and large setiferous punctures; each puncture carrying yellow seta; punctures becoming less deep from segments 3 to 8; punctures on segments 7 and 8 much less deep. Interstices between punctures in segment 3 similar to head, pronotum and elytra; interstices between punctures becoming larger from segments 3 to 8. Surface of abdomen with similar dense microsculpturing formed by uniformly distributed micropunctures as head, pronotum and elytra, appearing matt. Aedeagus as in Figure 1 C. In lateral view, paramere small and strongly curved, with its apex reaching half of median lobe; with short acute apex and without sensilla. Median lobe in lateral view curved, bulbous at base, apical region distinctly bigger than middle region; apex with small pointed to parameral side and with numerous sensilla in apical 2 / 3 region. Female similar to male. Variation. Total length 3.7 to 4 mm. Coloration, punctures pattern on the head, pronotum and elytra, and shapes of the antennomeres were consistent among the specimens examined.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123FFF815584FDCFFBBFFA3F.taxon	biology_ecology	Habitat. Captured manually under bark in decaying trunks and branches. The capture sites are destined for selective logging and the vegetation is second-growth Evergreen Tropical Forest “ Floresta Ombrófila Densa de Terras Baixas ” (Veloso et al. 1991).	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123FFF815584FDCFFBBFFA3F.taxon	distribution	Distribution. G. curupira sp. nov. is know from the municipalities of Itacoatiara and Itapiranga in Amazonas State, Brazil.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123FFF815584FDCFFBBFFA3F.taxon	etymology	Etymology. The specific epithet name is a noun in apposition and refers to a figure from Brazilian folklore (Curupira), of Amazonian origin, characterized as an entity that protects the forests. In a local legend (see Chaves 2021), at the exact point where the holotype of the species was collected, workers from a logging company reported having been terrorized by this folkloric figure after a day of work in the forest.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D1239FF835584FA6BFED7F9D7.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 43 CC 70 E 2 - 4 C 6 E- 46 A 1 - BE 67 - 41 F 6 B 1 CD 5 EAD (Figs 2 A – C)	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D1239FF835584FA6BFED7F9D7.taxon	materials_examined	Type material (15 ♂♂, 8 ♀♀). Holotype: BRAZIL: ♂, labeled: “ Brasil: AM, Carauari | Sítio da Tia Neuza, Manual | 01. i. 2020, Madeira podre | 4 ° 49 ’ 22.73 ’’ S / 66 ° 54 ’ 25.30 ’’ W | R. S. Moura | CESIT - UEA ”, “ HOLOTYPE | Glyptoma | luae sp. nov. | Desig. Klemann-Junior et al. 2024 ” (INPA). Paratypes (14 ♂♂, 8 ♀♀), labeled: “ Brasil: AM, Itapiranga | Faz. Caribi - Madeira podre | 16. xii. 2019 | 2 ° 32 ’ 55.74 ’’ S / 58 ° 48 ’ 24.21 ’’ W | L. Klemann-Junior; R. S. Moura | CESIT - UEA ” (5 ♂♂, 1 ♀ INPA); “ Brasil: AM, Itapiranga | Faz. Caribi - Madeira podre | 17. xii. 2019 | 2 ° 32 ’ 50.48 ’’ S / 58 ° 48 ’ 23.16 ’’ W | R. S. Moura; L. Klemann-Junior | CESIT - UEA ” (2 ♂♂ CEMT, 2 ♂♂ UFPR, 1 ♀ INPA; 2 ♀♀ UEA); “ Brasil: AM, Carauari | Col. Manual (Mad. Podre) | 05. i. 2020 | 4 ° 49 ’ 22,73 ’’ S / 66 ° 54 ’ 26,30 ’’ W | R. S. Moura; A. S. Santos | CESIT - UEA ” (1 ♀ CEMT); “ Brasil: AM [Amazonas state], Carauari | Sítio da Tia Neuza, Manual | 01. i. 2020, Madeira podre | 4 ° 49 ’ 22.73 ’’ S / 66 ° 54 ’ 25.30 ’’ W | R. S. Moura | CESIT - UEA ” (3 ♂♂ UEA, 1 ♂ CEMT, 1 ♂ UFPR, 1 ♀ CEMT, 2 ♀♀ UFPR). All paratypes with yellow label “ PARATYPE | Glyptoma | luae sp. nov. | Desig. Klemann-Junior et al. 2024 ”. Labels abbreviations, translations, and explanations: AM = Amazonas state; CESIT - UEA = Centro de Estudos Superiores de Itacoatiara - Universidade do Estado do Amazonas = written in a vertical line on the left side; Col. Manual (Mad. podre) = Coleta Manual (Madeira Podre) = captured manually under bark in decaying trunks and branches; Faz. = Fazenda = farm; Madeira podre = captured manually under bark in decaying trunks and branches; Manual = captured manually.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D1239FF835584FA6BFED7F9D7.taxon	diagnosis	Diagnosis. Among species of the G. cognatum group (with size between 1.8 and 3.2 mm, the setiferous punctures on the pronotum and elytra are at least partly shiny) G. luae sp. nov. is as long as G. schuhi and G. obscuricolle. However, G. luae can be differentiated from these two species by the presence of two almost regular and parallel rows of punctures along the entire length of the space between the suture and discal carina (at least partly three irregular rows of punctures in G. schuhi and two irregular rows of elongate punctures in G. obscuricolle); by the head, pronotum and elytra without microsculpture (isodiametric microsculpture in G. obscuricolle); by the larger punctures on the head (finely punctuate on G. obscuricolle); and by the distinctive form of the aedeagus. The median lobe of the aedeagus of G. luae is longer than the median lobe of G. obscuricolle. The basal bulb of G. luae, at the point of insertion of the parameres, has a straight upwards line before curved to the median lobe, whereas in G. obscuricolle has a downwards curve (in lateral view with apex pointed to right) before curved to the median lobe. Parameres of the aedeagus of G. luae is bigger (reaching almost the half of the median lobe), sinuate, and with a “ J ” shape, whereas in G. schuhi is short (reaching the basal fourth of the median lobe) and slightly sinuate.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D1239FF835584FA6BFED7F9D7.taxon	description	Description. Holotype male, BL: 3.13 mm, FL: 1.5 mm. Coloration: dark reddish brown; antennae and legs light reddish brown. Head (Figure 2 A) wider (HW: 0.51) than long (HL: 0.40); subrectangular; with rounded hind and anterior angles; large eyes slightly prominent and longer than temples; sides in front of eyes parallel up to base of antennae; at base of antennae with almost rectangular angle that narrowed to anterior edge of clypeus in a sinuate curve; anterior edge of clypeus less wide than half of width between base of antennae. Head with setiferous punctures moderately dense and deep; punctures not elongate; each puncture with a yellow seta; on average, interstices between punctures less wide than half of diameter of punctures. Integument and punctures of head without microsculpture, shiny. Neck appearing matt due to dense microsculpture formed by uniformly distributed micropunctures. Antenna (Figure 2 B) slightly longer than head and pronotum together; antennomeres 2 – 10 gradually and smoothly widening anteriorly; antennomere 2 slightly narrower than antennomere 1; antennomeres 2 – 10 slightly wider than long; antennomere 11 narrower and longer than antennomere 10; all antennomeres with numerous long yellow setae; setae approximately as long as width of antennomeres. Pronotum (Figure 2 A) wider than long (PL: 0.43; PW: 0.48); widest in anterior third; narrowed to anterior widely rounded angles in smooth convex curve; narrowed to posterior obtuse angles in smooth concave curve. Pronotum with setiferous punctures dense and deep; with some coalescent punctures without evident ridges; each puncture with a yellow seta; width of punctures irregular; interstitial ridges between punctures of variable size. Integument of pronotum without microsculpture; surface of punctures with few micropunctures; shiny. Elytra (Figure 2 A) wider than pronotum; longer than wide (EL: 0.67; EW: 0.62); wildest in middle; shoulders well developed and rounded; posterior angles rounded; anterior edge of elytra concave. Discal carina distinct; partly irregular or broken due to the punctures; slightly curved outward. Lateral carina indistinctly restricted shortly behind shoulders. Suture marked by line of 9 – 10 smaller, coalescent, setiferous punctures. Space between suture and discal carina with two almost regular and parallel rows of setiferous punctures along entire length of elytra, each row with 8 to 9 elongated punctures; some coalescent punctures without evident ridges. Laterally to discal carina another two almost regular and parallel rows of setiferous punctures along entire length of elytra; more lateral to these two lines, slightly smaller sparse setiferous punctures present. Each puncture with a yellow seta; setae in asymmetric position to centre of punctures; interstices of the punctures mostly restricted to small ridges. Integument of elytra without microsculpture; surface of punctures with few micropunctures; shiny. Legs (Figure 2 A) almost completely covered with yellow setae; surface appearing matt, with transverse striate microsculpture. Abdomen with dense and large setiferous punctation; each with one yellow seta; interstices between punctures restricted to small ridges; punctures on segments 3 - 6 deep; punctures on segments 7 and 8 less deep. Surface of abdomen with microsculpture formed by uniformly distributed micropunctures; micropunctures less deep on interstices between punctures and on segments 7 and 8; segments 3 – 6 appearing matt, except for shiny intersticies between punctures; segments 7 and 8 appearing shiny; anterior border of all segments appearing matt due to presence of micropunctures; posterior border of all segments appearing shiny due to absence of micropunctures. Aedeagus as in (Figure 2 C.) In lateral view, paramere small and pointed, slightly sinuate, with its apex reaching half of median lobe. Median lobe in lateral view curved, bulbous at base, with its apical region converging to narrow pointed apex; apical region with numerous sensilla. Female similar to male. Variation. Total length 3.0 to 3.2 mm. Three specimens are lighter reddish brown; punctures and interstices pattern on the head, pronotum and elytra, and shapes of the antennomeres were consistent among the specimens examined.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D1239FF835584FA6BFED7F9D7.taxon	biology_ecology	Habitat. Captured manually under bark in decaying trunks and branches. The capture sites are destined for selective logging or rural production and the vegetation is second-growth Evergreen Tropical Forest “ Floresta Ombrófila Densa de Terras Baixas ” (Veloso et al. 1991).	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D1239FF835584FA6BFED7F9D7.taxon	distribution	Distribution. Known from the municipalities of Itapiranga and Carauari in Amazonas State, Brazil.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D1239FF835584FA6BFED7F9D7.taxon	etymology	Etymology. The species is named after the daughter of the two first authors, Lua de Moura Klemann; it is a noun in apposition.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123BFF8D5584F933FC68FADB.taxon	description	https: // zoobank. org / urn: lsid: zoobank. org: act: 00 ECE 7 D 7 - 22 D 1 - 4 C 70 - BB 21 - 24 A 189 D 173 E 8 (Figs 3 A – C)	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123BFF8D5584F933FC68FADB.taxon	materials_examined	Type material (2 ♂♂, 1 unsexed due to lost abdomen) Holotype: BRAZIL: ♂, labeled: “ Brasil: Para]; Aldeia | Coraci, 12 km W. of Ca- | ninde. R. Gurupi. Leg: | B. Malkin. 24: IV: 1963. ”, “ under | bark ”, “ Holo- | type ”, “ Glyptoma | malkini | det. U. Irmler ” (FMNH). Paratypes (1 ♂, 1 unsexed), labeled: “ Brasil: Para; Aldeia | Coraci, 12 km W. of Ca- | ninde. R. Gurupi. Leg: | B. Malkin. 24: IV: 1963. ”, “ under | bark ”, “ Para- | type ”, “ Glyptoma | malkini | det. U. Irmler ” (1 ♂, 1 unsexed FMNH). Labels abbreviations, translations, and explanations: Para = Pará state; Aldeia | Coraci = Coraci indigenous village; Ca- | ninde. = Canindé Village; R. Gurupi. = Gurupi River.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123BFF8D5584F933FC68FADB.taxon	diagnosis	Diagnosis. Among species of the G. cognatum group (with body length between 1.8 and 3.2 mm, setiferous punctures on the pronotum and elytra at least partly shiny) G. malkini sp. nov. is similar to G. luae sp. nov. and G. nitens in the presence of two rows of punctures in the posterior half of the space between suture and discal carina. However, G. malkini can be differentiated from G. luae sp. nov. and G. nitens by the body length (body length of G. malkini is 2.1 mm; body length of G. luae sp. nov. is 3.0 - 3.2 mm; and body length of G. nitens is 2.6 – 2.9 mm); by the integument of the head appearing matt (shiny in G. luae sp. nov. and G. nitens); by the slightly elongate punctures on the head (punctures not elongate in G. luae sp. nov. and G. nitens); by the distinctly wider than long antennomeres 2 - 10 (slightly wider than long in G. luae sp. nov. and G. nitens); by the pronotum with dense microsculpturing formed by uniformly distributed micropunctures, appearing matt, and ridges in some parts without microsculpture, appearing shiny (the pronotum of G. luae sp. nov. the shiny integument lacks microsculpture and the surface of the punctures have few micropunctures; in G. nitens the microsculpture is restricted to the surface of punctures and the integument is shiny); by the presence of two parallel regular rows of punctures between suture and discal carina (there are two parallel, irregular rows of punctures in G. nitens); and by the distinctive shape of the aedeagus. The aedeagus has two sensilla at the base of the median lobe (the median lobe has numerous sensilla in G. luae sp. nov. and G. nitens); the parameres are narrow (broad in G. nitens); the parameres are strongly curved (it is slightly sinuate, with a “ J ” shape, in G. luae sp. nov.).	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123BFF8D5584F933FC68FADB.taxon	description	Description: Holotype male, BL: 2.1 mm, FL: 1.07 mm. Coloration: black; legs brown. Head (Figure 3 A) wider (HW: 0.38 mm) than long (HL: 0.26); subrectangular; eyes large, prominent; four times as long as temples; temples narrowed in smooth curve to neck; neck two thirds as wide as head; sides in front of eyes parallel; preocular part as long as length of eyes; anterior margin slightly convex. Head with large and dense setiferous punctation; vertex more densely punctate than clypeus; punctures slightly elongate; vertex with interstices between punctures approximately half as wide as diameter of punctures; surface of interstices and punctures with dense microsculpturing formed by uniformly distributed micropunctures; appearing matt. Neck appearing matt due to dense microsculpture formed by uniformly distributed micropunctures. Antennae (Figure 3 B) as long as head and pronotum combined; first antennomere approximately quadrate; antennomeres 2 – 10 distinctly wider than long; antennomere 2 one fourth wider than long; antennomere 10 twice as wide as long; antennomere 11 narrower and longer than antennomere 10; all antennomeres with long yellow apical setae; setae approximately as long as width of antennomeres. Pronotum (Figure 3 A) wider (PW: 0.35 mm) than long (PL: 0.32 mm); widest shortly behind anterior angles; anterior half slightly narrowed; posterior half narrowed in deep concave curve to posterior angles; posterior margin slightly concave; surface with deep, coarse, dense setiferous punctuation ;; interstices between punctures reduced to narrow ridges; interstices approximately one tenth as wide as diameter of punctures; yellow setae in asymmetric position to centre of punctures; seta in some cases inserted nearly on ridge; dense microsculpturing formed by uniformly distributed micropunctures, appearing matt; ridges in some parts without microsculpture, appearing shiny. Elytra (Figure 3 A) longer (EL: 0.49 mm) than wide (EW: 0.43 mm); sides slightly convex; widest close to middle; shoulders well developed and rounded; posterior angles rounded; shoulders not projected forward. Discal carina distinct; slightly curved outward. Lateral carina behind shoulder short and indistinct. Suture marked by a line of six smaller coalescent setiferous punctures. Space between suture and discal carina with two rows of setiferous punctures along entire length of elytra. Space laterad of discal carina with two additional rows of setiferous punctures along entire length of elytra. Each puncture carrying yellow seta; seta in asymmetric position to centre of punctures; interstices between punctures evident as small ridges. Integument of elytra without microsculpture; surface of punctures with dense microsculpturing formed by uniformly distributed micropunctures; appearing shiny. Legs almost completely covered with yellow setae; surface appearing matt, with microridges organized with cells. Abdomen with dense, large setiferous punctation; each puncture with yellow seta; insertion of setae slightly asymmetric; interstices between punctures restricted to small ridges. Surface of punctures with dense microsculpturing formed by uniformly distributed micropunctures; surface of ridges between punctures shiny. Aedeagus as in (Figure 3 C.) In lateral view, paramere curved, with its apex reaching base of 1 / 3 apical region of median lobe. Median lobe in lateral view curved, bulbous at base with two sensilla, its apical region converging to narrow pointed apex and with elongate tubular endophallus. Female unknown. Variation. No variations were observed among the three specimens examined.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123BFF8D5584F933FC68FADB.taxon	biology_ecology	Habitat. Collected under bark. The vegetation is Evergreen Tropical Forest.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123BFF8D5584F933FC68FADB.taxon	distribution	Distribution. Known from the municipality of Paragominas in Pará State, Brazil.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
F17D7D2D123BFF8D5584F933FC68FADB.taxon	etymology	Etymology. The species name honors the collector of the species, Borys Malkin, who collected Staphylinidae in many Central and South American countries; it is a noun in apposition.	en	Klemann-Junior, Louri, Moura, Roberta Souza De, Irmler, Ulrich, Asenjo, Angélico (2025): New species and new records of Glyptoma Erichson, 1839 (Coleoptera: Staphylinidae). Zootaxa 5584 (4): 539-550, DOI: 10.11646/zootaxa.5584.4.5, URL: https://doi.org/10.11646/zootaxa.5584.4.5
