taxonID	type	description	language	source
514AFCE05B2A5ABE8B0BD2F27531210A.taxon	description	Figs 8, 9, 10	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
514AFCE05B2A5ABE8B0BD2F27531210A.taxon	etymology	Etymology. furtiva (L., hidden, concealed), from the rarity, and microhabitat preference, of this species.	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
514AFCE05B2A5ABE8B0BD2F27531210A.taxon	diagnosis	Diagnosis. The epigynum of C. furtiva sp. nov. differs from that of C. heterops Chamberlin & Ivie (1937) in possessing a slightly more sclerotized and rugous epigynal plate anterior to the epigastric furrow, with white (glandular?) material lying anterior to the spermathecae, and longer spermathecae that nearly meet at the midline.	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
514AFCE05B2A5ABE8B0BD2F27531210A.taxon	description	Description of ♀ holotype. (SDSU _ TAC 000889; Figs 8 B – H, 9 A, J – M). Color in alcohol greenish brown with dark gray dorsal pattern. Carapace with dark markings extending from eyes to cephalic area, and sides of cephalic area; abdomen grayish with dark pattern dorsally, laterally and around spinnerets. Sternum unmarked. Legs darker at distal femora, basal tibiae, and basal metatarsi. Cheliceral promargin with three teeth (two basal together, one distal parted), retromargin with seven very small teeth. Total length 3.53, carapace length 1.40, carapace width 1.05, cephalic region width 0.60, posterior eye row width 0.45. Anterior median eyes missing, represented by small dark blotches of black pigment. Eye diameters ALE: PLE: PME = 0.09: 0.08: 0.08. Sternum length 0.70, sternum width 0.67. Leg formula 1423. Leg I article lengths (1.27, 0.49, 1.08, 1.05, 0.75 = 4.64) leg IV article lengths (1.37, 0.45, 1.25, 2.67, 0.90 = 6.64). Abdomen 2.17 long. Leg macrosetae (Fig. 9 C – I, paratype MACN - Ar 46970) absent on femora, 1 - 1 on patellae (dorsal), thick on legs I – III, weaker on IV. Tibia I ventral 2 - 2 - 0, prolateral 1 - 1 - 0, dorsal 1 - 0 weaker; metatarsus I prolateral 1 - 1 - 1 - 1, ventral 2 - 2 - 2 - 0 not well paired. Tibia IV prolateral 1 - 1, ventral 1 - 1, retrolateral 0 - 1, dorsal 1 - 0 - 1; metatarsus IV prolateral 1 - 1, ventral 0 - 1 - 1, retrolateral 0 - 0 - 2. Patellae with basal fracture region seen as darker indentations at the sides (Fig. 9 A, B). Spinnerets short, colulus a hairy patch (Fig. 9 J). Epigynal plate oval with sinuous transversal ridge and rugous median area (Fig. 9 K, L). Wide proximal copulatory ducts filled with whitish material (Fig. 9 K), spermathecae posterior, transversal. Copulatory openings not seen, probably in the anterior ridges and leading to the wide, soft copulatory ducts and from there to posterior spermathecae. Bennett’s gland large, on ectal side of spermatheca (Fig. 9 M), fertilization ducts posteriorly placed.	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
514AFCE05B2A5ABE8B0BD2F27531210A.taxon	distribution	Distribution and natural history. Confirmed specimens from Del Norte and Humboldt counties, California (Fig. 10), at lower elevations (120 – 150 m). Topotypic specimens were abundantly collected under rock piles or logs, and in leaf litter, in wet mixed conifer forest. Males were only represented as penultimates. Robb Bennett (pers. comm.) has examined female specimens from Siskiyou County (southeast of Seiad Valley, Fig. 10), currently housed in the American Museum of Natural History, that appear to match this species.	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
F2A4040E5FF95633B2C103F53A888CFE.taxon	description	Figs 1, 6, 7, 10	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
F2A4040E5FF95633B2C103F53A888CFE.taxon	etymology	Etymology. armilla (L., a ring, bracelet), from the conspicuous ring-like patellar fracture lines (Fig. 1 C).	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
F2A4040E5FF95633B2C103F53A888CFE.taxon	description	Description of ♂ holotype. (SDSU _ G 1090; Fig. 6 A, B) Color in alcohol pale cream to white, fangs slightly darker. Carapace essentially bare, unmarked; abdomen dusky dorsally and ventrally, slightly darker dorsal longitudinal bars posteriorly. Total length 2.3, carapace length 1.15, carapace width 0.85, cephalic region width 0.5, posterior eye row width 0.325. Eye diameters AME: ALE: PME: PLE = 0.025: 0.075: 0.05: 0.075. Leg I article lengths (1.65, 0.5, 1.9, 1.6, 0.9 = 6.4), leg IV article lengths (1.45, 0.4, 1.5, 1.5, 0.8 = 5.6). Leg macrosetae sparse, one or two on tibia III and V (prolateral and ventral), and metatarsus III and IV (prolateral and ventral, distal cluster). Male palp with shelf-like distal patellar apophysis. Basal retrolateral tibial apophysis (RTA) a minute spike, with sclerotized parallel weak ridges bordering a slight excavation, extending to a short and blade-like distal RTA. Bulb with a short dorsal tegular process at base of whip-like embolus which extends to ~ 3 o’clock then projects ventrally, protected by conductor. Triangular conductor with spatulate distal, and sharply pointed basal lobes. Beak-like ventral lobe of conductor enclosing embolus. Median apophysis absent. Cymbium elongate, distal projection approximately as long as bulb.	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
F2A4040E5FF95633B2C103F53A888CFE.taxon	description	Description of ♀ paratype. (SDSU _ G 1086 A; Fig. 7 A – E) Color in alcohol pale cream to orange, fangs slightly darker. Carapace essentially bare, unmarked; abdomen without markings. Total length 2.9, carapace length 1.3, carapace width 0.925, cephalic region width 0.55, posterior eye row width 0.325. Eye diameters AME: ALE: PME: PLE = 0.025: 1: 0.05: 1. Leg I article lengths (1.57, 0.5, 1.6, 1.4, 0.9 = 5.9), leg IV article lengths (1.55, 0.4, 1.5, 1.5, 0.7 = 5.7). Leg macrosetae sparse, 1 on dorsal patella III and IV, 1 – 2 on tibia III and V (prolateral and ventral), and metatarsus III and IV (prolateral and ventral, distal cluster). Epigynal plate with a semicircular ridge along the anterior edge (Fig. 7 D, E). Paired lateral ear-shaped copulatory openings, sclerotized laterally and ventrally, lie ventral to this ridge. Thin copulatory ducts arise from copulatory openings and loop dorsally to meet in the midline, then extend ventrally in parallel, diverge at the bottom, then extend dorsally to connect with spermathecae. Paired spermathecae bilobed, medial lobe slightly smaller and more oval-shaped than larger and more spherical lateral lobes, connected by a short, looped duct. Fertilization ducts short and indistinct, extending ventrally from medial lobes.	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
F2A4040E5FF95633B2C103F53A888CFE.taxon	distribution	Distribution and natural history. Known from five separate locations (three in close vicinity near Beaver Creek), spanning from northern records near Cave Junction, Oregon to southern records along the Salmon River in Siskiyou County, California (Fig. 10). All collecting events are from mixed coniferous forests (including Pseudotsuga menziesii, Pinus sp., Arbutus menziesii) at mid-elevations (550 – 800 m) in the Klamath-Siskiyou Mountains. We anticipate additional populations in intervening and neighboring locations. Spiders found abundantly beneath rocks in shaded rock piles, often under rocks lying directly on the surface, versus deeper in rock piles. Spiders were generally found without associated webs, although some subadult specimens found along Beaver Creek at Fishtrap Creek (MCH 24 _ 057) were found near sparse webbing (Fig. 1 D). Adults have been collected in April, July, and August; only subadults were found during the July collecting event at Fishtrap Creek (MCH 24 _ 057).	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
D143EFD9B16158E1A385DBA7ACE1A587.taxon	description	Figs 1, 6, 7	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
D143EFD9B16158E1A385DBA7ACE1A587.taxon	etymology	Etymology. A modification of siskiyou, from the Klamath-Siskiyou Mountains that encompass much of the known distribution of this genus. The etymology of siskiyou, possibly indigenous, remains uncertain. Grammatical gender treated as feminine.	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
D143EFD9B16158E1A385DBA7ACE1A587.taxon	diagnosis	Diagnosis. With a conspicuous fracture line near the base of patellae I – IV, like close relatives Calymmaria and Willisus (Roth 1981: fig. 1; Heiss and Draney 2004: fig. 3). The male palp of Siskiyu gen. nov. is diagnosed from Calymmaria in possessing a dorsal tegular process, a thinner embolus with the distal tip projecting ventrally, associated with a trilobed conductor (Fig. 6 A, B). Calymmaria palps lack a tegular process, have generally thicker (or sometimes forked) emboli, with conductors that lack dorsal processes (Heiss and Draney 2004: figs 6, 7). Siskiyu gen. nov. male palps are most like those of Willisus but differ in the condition of the dorsal tegular process (more robust and toothed in Willisus), shape of the embolus (sinuate in Willisus), and shapes of both the conductor and RTA lobes (Roth 1981: figs 3 – 5). Siskiyu gen. nov. female genitalia differ from Calymmaria in possessing subequal, bilobed spherical spermathecae (Fig. 7 D, E), in contrast to the simple paired spherical spermathecae of most Calymmaria (Heiss and Draney 2004: figs 4, 5), and different from the distinctive C. alleni epigynum; Heiss and Draney 2004: figs 8, 9). Siskiyu gen. nov. female genitalia are distinguished from those of Willisus which include short copulatory ducts and oblique kidney-bean shaped spermathecae (Roth 1981: figs 6, 7).	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
D143EFD9B16158E1A385DBA7ACE1A587.taxon	description	Description. Adults 2 – 3 mm in length. Dorsal carapace mostly bare and lacking pigmentation, longitudinal fovea ~ 2 × length of PME diameter. Eight eyes, anterior eye row slightly recurved, posterior eye row slightly procurved. AMEs smallest, ~ ¼ diameter of ALEs, separated by diameter. Lateral eyes approximately contiguous, approximately equal in size; PMEs ½ diameter of PLEs, slightly closer to PLEs. Clypeus as high as ALEs. Chelicerae straight, longer than width of cephalic region, lateral boss present; anterior margin with three teeth, posterior margin with five or six teeth. Labium nearly square, narrowing slightly anteriorly. Endites slightly convergent, ~ 2 × as long as wide, with serrula. Sternum approximately heart-shaped but with truncated anterior edge, narrowed posteriorly, ending at posterior edge of hind coxae; sparsely covered with fine hairs. Trochanters without notches. Legs with fine hairs, mostly unmarked. Leg formula 1, 4, 2, 3. Patella-tibia I 1.6 – 2 × longer than carapace. Femurs lacking macrosetae; patellar, tibial, and metatarsal macrosetae restricted to posterior legs. Leg patellae with conspicuous cleavage plane basally at ⅕ of patella length (Fig. 7 C). Tarsi and metatarsi with dorsal rows of 3 – 6 long trichobothria. Paired tarsal claws with ten teeth, median claw very small, semicircular hairs between claws not obvious. Abdomen with scattered simple hairs, more ventrally than dorsally. Colulus inconspicuous, with six setae. Anterior spinnerets are stout, conical, separated by basal width, distal segment hemispherical. Posterior spinnerets longer than anterior with wedge-shaped distal segments ~ 1.3 × longer than length of basal segment, pointed medially. Median spinnerets lying directly posterior and approximately equal in height to anterior spinnerets, closely adjacent. Female palp with tibial and tarsal macrosetae, clothed with fine hairs. Female genitalia with paired lateral openings, elongate copulatory ducts, bilobed spermathecae. Male palp with distal patellar apophysis; tibia with basal and distal processes. Bulb with small tegular process, slender embolus protected by beak-like ventral process of triangular conductor.	en	Hedin, Marshal, Ramírez, Martín J., Monjaraz-Ruedas, Rodrigo (2025): Phylogenomics of North American cybaeid spiders (Araneae, Cybaeidae), including the description of new taxa from the Klamath Mountains Geomorphic Province. ZooKeys 1226: 47-75, DOI: 10.3897/zookeys.1226.140204
