taxonID	type	description	language	source
8C1F65DC43775B12B2AD9AA82298C51B.taxon	description	Description. Holotype, male. Head. Fastigium of vertex equal or slightly wider than half of scapus. Thorax. Pronotum short, slightly constricted in the middle, median sulcus located after the middle, cylindrical in prozona and somewhat raised in metazona, caudal margin of the disc concave, medial carina occurs as a yellowish line, disk bordered by large light lines slightly divergent in anterior and posterior margins constituting roughly as “) (“ shape; paranotal margin almost straight along prozona and oblique along metazona. Tegmina short, extending beyond the posterior margin of pronotum, stridulatory vein not totally covered by pronotum; stridulatory file with ca. 55 teeth. Male terminalia. Cercus cylindrical, gradually tapering toward apex, curvature is more prominent apically, incurved roughly as L-shaped, with a flattened apex and with 4 – 5 distinguishable denticles on external margin. Subgenital plate wider than long, with a wide roughly quadrangular median processes apically, distal margin is quadrangularly concave.	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
8C1F65DC43775B12B2AD9AA82298C51B.taxon	diagnosis	Diagnosis. The new species, P. anisozonatus sp. nov., shows sister group relationship with P. isozonatus, and each of them was consistently suggested as a distinct species by all species delimitation tests. The pair of P. isozonatus / P. anisozonatus sp. nov. can easily be distinguished by traditionally used phenotypic characters (Table 3 and Fig. 13). The typical cercus with flattened apex and prominent denticles located externally (no denticle on the internal side of apex), and male subgenital plate wider than long, are the most prominent characters distinguishing P. anisozonatus sp. nov. from P. isozonatus. Apart from the cercus and subgenital plate of the male, P. anisozonatus sp. nov. can be distinguished from P. isozonatus also by its male song – the syllable duration being 30.57 (27 – 34) ms in the first, while 13.6 (10 – 19) ms (Konya-Taşkent) or 6.46 (2 – 8) ms (Niğde-Çamardı) in the second (Table 3). Along with these phenotypic characters, there are six mutations, detected in the mitochondrial concatenated matrix by applying a PAUP analysis (File S 1), specific to the ancestral node of P. anisozonatus sp. nov., which we considered as a diagnostic character of this species; 1730 (C → T), 1799 (A → G), 1917 (A → C), 1928 (T → C), 2118 (C → T), 2223 (C → T).	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
8C1F65DC43775B12B2AD9AA82298C51B.taxon	etymology	Derivatio nominis. The name of the new species is constituted to express its close relation, but the clear distinction, from P. isozonatus.	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
0A243E972EA85A2DB1C8439A65CA5F10.taxon	distribution	Distribution. This species is known only from its type locality and close surroundings, along the Çoruh Valley in Erzurum and Artvin Provinces of Turkey (Fig. 1, Table 1, and Sevgili et al. (2018), Kaya (2018)).	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
D1368AA141DE578C9E4F72D5857CE681.taxon	description	Description. Holotype, male. Head. Fastigium of vertex equal or slightly narrower than half of scapus. Thorax. Pronotum short, slightly constricted in the middle, median sulcus located after the middle, cylindrical in prozona and distinctly raised in metazona, caudal margin of the disc concave, median carina occurs as a weak yellowish line; paranotal margin almost straight along prozona and oblique along metazona. Tegmina short, extending beyond the posterior margin of pronotum and reach to half of the second abdominal tergite; stridulatory vein almost totally covered by pronotum; stridulatory file with ca. 60 teeth. Male terminalia. Cercus cylindrical, gradually tapering toward apex, prominently incurved at apical half, incurved roughly as L-shaped, with a cylindrical distal branch almost as long as proximal branch and with 4 – 5 distinguishable denticles on external margin of distal branch and 3 – 4 denticles along the tip. Subgenital plate as wide as or slightly wider than long, with a quadrangularly concave caudal margin. Song. Male song consists of 11.3 (9 – 16) syllable duration (ms) and 14.65 (10 – 21) impulse number per syllable with occasional after-clicks following the main syllable at 20 – 30 ms. The peak frequency spectrum lies between 35 and 50 kHz. Thus, it is very similar to the song of P. denizliensis (see under the latter). Male song is exemplified in Figure 9 and song measurements are provided in Table 3. — Female. Similar to male in general. Pronotum slightly raised in metazona, tegmina well visible beyond hind margin of pronotum. Coloration. General coloration black with a creamish pattern; vertex black or with black dots on a creamish brown background, antennae black with regular white rings as in the group. Thorax. Disc of pronotum with black patterns or spots on a creamish brown background at the beginning of prozona, black in the middle and reddish brown in metazona; ventral half of the paranota creamish and dorsal half with black pattern; tegmina with typical black / light (marble or yellow) pattern; all legs are black dorsally. Abdomen. Abdominal terga black in front 3 / 4 and light in the remaining part, the black bands do not extend to subsequent tergum laterally. Female terminalia. Subgenital plate triangular, ovipositor typical of the group.	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
D1368AA141DE578C9E4F72D5857CE681.taxon	diagnosis	Diagnosis. The new species, P. boncukdagensis sp. nov., shows sister group relationships with P. ciplaki + P. datca. P. boncukdagensis sp. nov. was suggested as a separate identical species by bPTP and GMYC while was placed within P ciplaki by ASAP delimitation tests. However, the new species and P. ciplaki well differ from each other by male cercus. Cercus is weakly incurved, with rounded apex, denticles constitute a single row along the tip in P. ciplaki, while strongly incurved, L-shaped and with truncate apex, denticles constituting two lines, one along the tip and the other along external margin in P. boncukdagensis. Additionally, male subgenital plate is as wide as long or slightly wider than long in the new species, while it is longer than wide in P. ciplaki. The new species and P. datca are not monophyletic and no delimitation test suggested it belongs within P. datca, but the new species is rather similar to P. datca montana especially by the male cercus. The new species differs by the distal branch of cercus as long as proximal branch (longer than the half-length of the proximal branch), while it is at most as long as the half-length of the proximal branch in P. datca datca. Additionally, the distal branch of cercus is black in the new species while dark but not black in P. datca. Apart from the male cercus, P. datca and P. boncukdagensis sp. nov. can also be distinguished by the male calling song; a syllable consists of 1 – 4 and 10 – 21 impulses in the first and second species respectively (Table 3). Along with these phenotypic characters, there are 8 mutations, detected in the concatenated matrix of COI + ND 2 + VAL by applying a PAUP analysis (File S 1), specific to the ancestral node of P. boncukdagensis sp. nov., which we considered further diagnostic characters of the species, at the position 873 (T → C), 1322 (A → G), 1331 (C → T), 2039 (T → C), 2063 (A → G), 2498 (A → G), 2499 (G → T), 2500 (T → G) and 2897 (G → A).	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
D1368AA141DE578C9E4F72D5857CE681.taxon	etymology	Derivatio nominis. The name of the new species is established by the name of range area “ Boncuk Dağları ” Mts., located between Muğla and Denizli Provinces of Turkey.	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
3976BCF14E1F5B75B5A125C6C52C5E96.taxon	distribution	Distribution. Regarding the above-listed localities (Kaya 2018; Sevgili et al. 2018) and those given in Table 1, this species occurs in the lowlands of Fethiye, Dalaman, and Ortaca in the south of Muğla provinces of Turkey.	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
2D9A68A2CF585972ACA0EE5F0DB95C0D.taxon	distribution	Distribution. This species is reported from several localities along the Southern Taurus Mts., in the Konya, Niğde, and Karaman provinces of Turkey (Fig. 1, Table 1, and Kaya (2018)). The locality “ Konya-Hadim, road to Gündoğmuş, 36.88333 N, 32.11667 E ” reported by Kaya (2018) for P. isozonatus refers to P. anisozonatus sp. nov. (see below).	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
860F92C51CC85ACC8309370D23EA9A96.taxon	description	Description. Holotype, male. Head. Fastigium of vertex equal or slightly wider than half of the scapus. Thorax. The pronotum short, slightly constricted in the middle, median sulcus located after the middle, cylindrical in prozona and distinctly raised in metazona, caudal margin of the disc concave, medial carina occurs as a yellowish line or absent, disk bordered by large light lines divergent in anterior and posterior margins constituting roughly as “) (“ shape; paranotal margin almost straight along prozona and oblique along metazona. Tegmina short, extend beyond the posterior margin of pronotum, stridulatory vein hardly visible under the pronotum; the stridulatory file with 58 teeth. Male terminalia. Cercus cylindrical, gradually tapering toward apex, the curvature is more prominent apically, incurved roughly as L-shaped, with a robust, but slightly tapered apex and 2 – 3 hardly distinguishable denticles apically. The subgenital plate is as long as wide or slightly wider than long, distal margin is almost truncated.	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
860F92C51CC85ACC8309370D23EA9A96.taxon	diagnosis	Diagnosis. The three infraclades in Clade IIB, each of which was consistently suggested as distinct species by all species delimitation tests, show P. variicercis + (P. parazonatus sp. nov. + P. zonatus) relationships on the phylogenetic tree. However, they are very similar in the traditionally used structures / characters (Fig. 12). The distinct cercus, with almost blunt apex and indistinguishable denticles located at the tip, is the most prominent character distinguishing P. parazonatus sp. nov. from the other two related species. The black coloration of abdominal terga is more distinct in P. parazonatus sp. nov., but it is not reliable as it may differ in young and elder individuals, or may be locality-specific. Apart from phenotype, there are unambiguous mutations specific to the ancestral node of this species, which we considered as diagnostic characters of the species. These positions are (see File S 1); 1961 (A → G), 1259 (T → C), 1331 (A → T), and 1424 (T → C). The first is unique to P. parazonatus sp. nov., and the other three unambiguous mutations are not unique to this clade, but none of them is shared with the two closest relative species, P. variicercis and P. zonatus.	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
860F92C51CC85ACC8309370D23EA9A96.taxon	etymology	Derivatio nominis. The name of the new species is constituted by considering the phylogenetic position of P. parazonatus sp. nov. with P. zonatus on the phylogenetic tree as P. variicercis + (P. parazonatus sp. nov. + P. zonatus).	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
A300022E4564507EBBD5E22AD481A559.taxon	distribution	Distribution. This species is known only from its type locality and close surroundings, Nigde, Ulukışla (Fig. 1, Table 1, and Sevgili et al. (2018)).	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
794F46161DBF5A2A9E71F5CC77292B9F.taxon	distribution	Distribution. The type locality of the species was reported as ‘ Syria’ but see above.	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
58F00DFB51E95A7B9A85317317A20233.taxon	distribution	Distribution. This species is known only from the Erzurum province of Turkey (northeast part), plus some localities in the neighbouring provinces of Kars and Ardahan (Fig. 1, Table 1, and Sevgili et al. (2018)).	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
C4B44B0F53F457E0953BC75D9EED88A0.taxon	distribution	Distribution. This species is the only representative of the group in the Balkans, known only from the type locality, Vodno Mt., and a few closely situated locations in the Mariovo region of North Macedonia (Fig. 1, Table 1, and Sevgili et al. (2018)).	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
41AD0E4FCA255EFB81DC5FC67418AB13.taxon	distribution	Distribution. Possibly this is the most widespread species of the group. Its range covers associated altitudinal belts starting from the central part of the Anatolian Diagonal (or Maraş Triangle; see Çıplak et al. 1993) and extending to the Zagros Mountains (Van province of Turkey) in the east-west direction. Following the “ Previous records ” which review all previous published records and “ Material examined ” in Sevgili et al. (2018) and records in Kaya (2018), localities from the following provinces of Turkey were considered as P. zonatus; Bitlis, Kahramanmaraş, Malatya, Muş and Van. Apart from these provinces some localities in Erzincan (Kemaliye) and Erzurum (Kop Mt.) also refer to P. zonatus (see Fig. 1). The records by Sevgili et al. (2018) from Antalya, Konya, Niğde, and Muğla provinces of Turkey refer to other species (Fig. 1; see below). However, the material from Adana and Kayseri, the distributional border of P. zonatus and P. isozonatus, needs re-examination. The records of P. zonatus by Sevgili et al. (2018) and Kaya (2018) from the Munzur Mountains in Erzincan and Tunceli provinces of Turkey, the area situated between the two main branches of the Euphrates, the Murat and Karasu rivers (see Fig. 1), probably belong to P. parazonatus sp. nov. described here but require confirmation.	en	Uluar, Onur, Chobanov, Dragan P., Çıplak, Battal (2025): Merging taxonomy to systematics: A holistic approach to understanding the Poecilimon zonatus group (Orthoptera, Phaneropterinae). Arthropod Systematics & Phylogeny 83: 93-125, DOI: 10.3897/asp.83.e136516
