identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
E07687C30F74FFEDFF2C2F17FA42F0FC.text	E07687C30F74FFEDFF2C2F17FA42F0FC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Halone Walker 1854	<div><p>Key to the species of Halone in Cambodia based on the male genitalia</p><p>1 Uncus slender, with small apical spine; valva with apical process; saccular process not developed...................... 2</p><p>- Uncus stout, pointed; valva without apical process; saccular process well-developed................................ 4</p><p>2 Valva rounded, with two short apical processes............................................. H. specularis sp. nov.</p><p>- Valva truncated, with one apical and one basal area of costal margin............................................. 3</p><p>3 Basal process of costal margin longer than valva..................................................... H. ariadna</p><p>- Basal process of costal margin half size of valva...................................................... H. ivetae</p><p>4 Tegumen strongly shouldered, with a pair of dorsal lobe............................................. H. hollowayi</p><p>- Tegumen weakly shouldered, without dorsal lobe.......................................... H. angkorwat sp. nov.</p></div>	https://treatment.plazi.org/id/E07687C30F74FFEDFF2C2F17FA42F0FC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bayarsaikhan, Ulziijargal;Cha, Yeong-Bin;Kwon, Hyung Wook;Bae, Yang-Seop	Bayarsaikhan, Ulziijargal, Cha, Yeong-Bin, Kwon, Hyung Wook, Bae, Yang-Seop (2025): Genus Halone Walker (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with two new species and a newly recorded species. Zootaxa 5660 (1): 125-133, DOI: 10.11646/zootaxa.5660.1.9, URL: https://doi.org/10.11646/zootaxa.5660.1.9
E07687C30F74FFEDFF2C28F4FDF4F36D.text	E07687C30F74FFEDFF2C28F4FDF4F36D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Halone Walker 1854	<div><p>Genus Halone Walker, 1854</p><p>Halone Walker, 1854, List. Spec. Lepid. Insects Colln. Br. Mus. 2: 540. TS: Halone sobria Walker, 1854 . TL: Australia (Port Natal, Queensland).</p><p>Mosoda Walker, 1866, List. Spec. Lepid. Insects Colln. Br. Mus. 35: 1899. TS: Mosada anartoides Walker, 1866 . TL: Australia (Sydney).</p><p>Eurypepla Turner, 1922, Proc. R. Soc. Vict. 35: 30. TS: Eurypepla pteridaula Turner, 1922 . TL: Australia (Maclntyre Range, Queensland Nat. Park).</p><p>Eurypeplella Hedicke, 1923, Dt. Ent. Z. 1923 (2): 226. TS: Eurypepla pteridaula Turner, 1922 .</p><p>Mosada Turner, 1940, Proc. R. Soc. Qd. 51: 82. (missp.).</p><p>Species of the genus Halone can be recognized by the following characteristics: forewing ground color is creamy white or sordid white, with dark brown or pale brown markings or a large ochreous patch, and two dark discoidal spots or streaks; hindwing is light or pale gray (except for some Australian species with a distinct, dark terminal band); in the male genitalia, the valva has a very diverse form; the aedeagus is slender, strongly curved, with irregular cornuti; in the female genitalia, the corpus bursae is usually with a signum (Bucsek 2012, 2014).</p><p>Distribution. India (Singh et al. 2021), Laos (Bucsek 2020, Bayarsaikhan et al. 2021), Thailand (Černý &amp; Pinratana 2009), Cambodia (Bae et al. 2016, Bae &amp; Bayharsaikhan 2019), Vietnam (Bae &amp; Bayharsaikhan 2019; Bayarsaikhan et al. 2022), Peninsular Malaysia (Bucsek 2012; 2014), Papua New Guinea (Hampson 1914), Australia (Edwards 1996, Hebert et al. 2013).</p></div>	https://treatment.plazi.org/id/E07687C30F74FFEDFF2C28F4FDF4F36D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bayarsaikhan, Ulziijargal;Cha, Yeong-Bin;Kwon, Hyung Wook;Bae, Yang-Seop	Bayarsaikhan, Ulziijargal, Cha, Yeong-Bin, Kwon, Hyung Wook, Bae, Yang-Seop (2025): Genus Halone Walker (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with two new species and a newly recorded species. Zootaxa 5660 (1): 125-133, DOI: 10.11646/zootaxa.5660.1.9, URL: https://doi.org/10.11646/zootaxa.5660.1.9
E07687C30F74FFEBFF2C2C91FDF3F717.text	E07687C30F74FFEBFF2C2C91FDF3F717.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Halone angkorwat Bayarsaikhan & Cha & Kwon & Bae 2025	<div><p>Halone angkorwat Bayarsaikhan &amp; Bae, sp. nov.</p><p>LSID: urn:lsid:zoobank.org:act: F1EC0B11-F533-4827-AF88-B2A9AD943DAF</p><p>(Figures 2, 7, 12)</p><p>Type material Holotype. [CAMBODIA] ♂, Seima (N12˚14′54″, E107˚03′15″), 11.XI.2011 (Bae YS, Kim YK, Ju YD, Park BS, Lee HJ), adults and genitalia slides no. INU-1977C . Paratypes. (2 ♀) [CAMBODIA] ♀, same data as the holotype, adults and genitalia slides no. INU-10036C; ♀, Seima (N12˚16′44″, E107˚03′56″), 12.XI.2011 (Bae YS, Kim YK, Ju YD, Park BS, Lee HJ), adults and genitalia slides no. INU-10038C .</p><p>Diagnosis. By the wing pattern of this species, it is hardly distinguishable from many others of the Oriental Halone . The male genitalia structure of the new species is similar to H. arcuata Bayarsaikhan &amp; Bae, 2022 (in the present study: Figs. 16–18), but it can be distinguished from the latter by the almost straight, asymmetric saccular process that is longer than the tegumen, a triangular tegumen with a strongly sclerotized dorsal lobe, and the aedeagus vesica without a distinct cornutus. In H. arcuata, the strongly waved, sclerotized saccular process is shorter than the tegumen, the rectangular tegumen has a sclerotized, rounded dorsal lobe, and the aedeagus vesica contains several spine-shaped cornuti (Bayarsaikhan et al. 2022: 388, figs. 5a, 5b; in the present study: Fig. 17). In the female genitalia, the following features clearly separate the new species from H. arcuata, a weakly sclerotized postvaginal plate with weakly sclerotized and wrinkled lateral lobes; the lamella antevaginalis has a pair of large arch-shaped plates; and the ductus bursae membranous. In H. arcuata, the sclerotized postvaginal plate has strongly serrulate, triangular lateral lobes; the lamella antevaginalis has a large arch-shaped plate; and the ductus bursae is partly sclerotized (Bayarsaikhan et al. 2022: 389, fig. 9; in the present study: Fig. 18).</p><p>Description Adult (Figs. 2a, 2b). Length of the forewing 6.5–7 mm in both sexes. Head, patagium, tegula and thorax sordid white, mixed with brown scales. Forewing ground color sordid white, with pale brown sprinkles; antemedial, medial, and postmedial dark brown patches on the costal margin; medial area of forewing with two rounded, dark brown, distinct patches; cilia sordid white. Hindwing pale brown; cilia pale brown. Abdomen fuscous in both sexes. Male genitalia (Fig. 7). Uncus stout, pointed, curved. Tegumen weakly sclerotized, with a strongly sclerotized, rounded dorsal lobe. Valva membranous, narrow, weakly angled in the medial area of the sacculus, rounded apex with weakly covered with setae, almost the same length as the tegumen. The strongly sclerotized saccular process asymmetric, the more waved left process being longer than the almost straight right one, both processes dentate apical half, and longer than valva. Vinculum broadly “U”-shaped. Aedeagus stout, with a weakly sclerotized carina, and vesica with a weakly sclerotized, tubular elasma, and scobinate. Female genitalia (Fig. 12). Papillae anales weakly covered with setae. Weakly sclerotized postvaginal plate with weakly sclerotized and wrinkled laterial lobes. Broadly tubular antrum strongly sclerotized. Both apophysis slender, and the apophysis posterioris slightly longer than the apophysis anterioris. Strongly sclerotized lamella antevaginalis with a pair of large, arch-shaped plates. Membranous ductus bursae tubular. Corpus bursae globular, membranous, with a strongly dentate signum on the center of bursae.</p><p>Distribution. Cambodia (Mondulkiri Prov.).</p><p>Etymology. The specific epithet is a proper noun referring to Angkor Wat, the most famous and historically important cultural site in Cambodia.</p></div>	https://treatment.plazi.org/id/E07687C30F74FFEBFF2C2C91FDF3F717	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bayarsaikhan, Ulziijargal;Cha, Yeong-Bin;Kwon, Hyung Wook;Bae, Yang-Seop	Bayarsaikhan, Ulziijargal, Cha, Yeong-Bin, Kwon, Hyung Wook, Bae, Yang-Seop (2025): Genus Halone Walker (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with two new species and a newly recorded species. Zootaxa 5660 (1): 125-133, DOI: 10.11646/zootaxa.5660.1.9, URL: https://doi.org/10.11646/zootaxa.5660.1.9
E07687C30F72FFEBFF2C2F81FE67F18B.text	E07687C30F72FFEBFF2C2F81FE67F18B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Halone ivetae Bucsek 2020	<div><p>Halone ivetae Bucsek, 2020</p><p>(Figures 4, 10, 13)</p><p>Halone ivetae Bucsek, 2020, Entomofauna carpathica, 32 (2): 23. TL: Laos (Khammouane Prov.).</p><p>Type series. [ LAOS] ♂, holotype Khammouane prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=105.166664&amp;materialsCitation.latitude=17.566668" title="Search Plazi for locations around (long 105.166664/lat 17.566668)">Nakai env.</a>, 500 m, 17°34´N, 105°10´E, 25–29. IV. 2019, leg. K. Bucsek, SNM with a museum serial number SZ 10 951. Paratypes. 2 ♂ like holotype, CKB .</p><p>Distribution. Cambodia (present study), Laos (Bucsek 2020).</p><p>Material examined. [ CAMBODIA] ♀, Samkos (N12˚12′41″, E102˚54′38″), 19–21.VII.2012 (Bae YS, Ju YD, Le XV, Park BS, Lee HJ), adults and genitalia slides no. INU-10439C . LAOS: ♂, Laos, Bolikhamsai Prov., Tad Leuk Waterfall (N18˚23′42″, E103˚04′04.14″ Alt.: 216m), 01.VIII.2019 (Bae YS, Qi MJ, Lee DJ, Ko JH, Lee TG, Cha YB, Jang CM), adults and genitalia slides no. INU-10305L .</p><p>Remarks. Halone ivetae is newly recorded from Cambodia (Mondulkiri Province) herein. This species was previously reported from Laos (Khammouane Province) by Bucsek (2020). Here, we report it from the Bolikhamsai Province for the first time</p></div>	https://treatment.plazi.org/id/E07687C30F72FFEBFF2C2F81FE67F18B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bayarsaikhan, Ulziijargal;Cha, Yeong-Bin;Kwon, Hyung Wook;Bae, Yang-Seop	Bayarsaikhan, Ulziijargal, Cha, Yeong-Bin, Kwon, Hyung Wook, Bae, Yang-Seop (2025): Genus Halone Walker (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with two new species and a newly recorded species. Zootaxa 5660 (1): 125-133, DOI: 10.11646/zootaxa.5660.1.9, URL: https://doi.org/10.11646/zootaxa.5660.1.9
E07687C30F72FFEBFF2C2BDBFE7CF3D8.text	E07687C30F72FFEBFF2C2BDBFE7CF3D8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Halone specularis Bayarsaikhan & Cha & Kwon & Bae 2025	<div><p>Halone specularis Bayarsaikhan &amp; Bae, sp. nov.</p><p>LSID: urn:lsid:zoobank.org:act: 51E9327C-64CC-4633-AEA1-CE3DA9F0C6D8</p><p>(Figures 3, 8)</p><p>Type material. Holotype. [CAMBODIA] ♂, Koh Kong (N11˚41′44.9″, E103˚06′51.8″, Alt.: 278m), 25.XII.2014 (Bae YS, Ju YD, Qi MJ, Bayarsaikhan U, Park BS, Na SM, Kim JW, Lee DJ), adults and genitalia slides no. INU-1978C.</p><p>Diagnosis. By the wing pattern of this species, it is hardly distinguishable from many others of the Oriental Halone . The male genitalia structure of the new species is similar to H. ariadna Bucsek, 2012, but it can be distinguished from the latter by its more rounded valva, which has two short, angled processes at the upper and lower angles of the valva, and the aedeagus vesical having a large, spine-shaped cornutus. In H. ariadna, the irregularly rectangular valva has a short, truncated process at the lower angle of the valva and a long, spine-shaped basal process on the costal margin, and the aedeagus vesica has two stouts, spine˗shaped cornuti (Bae &amp; Bayarsaikhan 2019: 396, fig. 3; in the present study: fig. 9).</p><p>Description. Adult (Fig. 3). Length of the forewing 7 mm in male. Head, patagium, tegula, and thorax pale gray, with pale brown sprinkles and the thorax darker than the other parts. Forewing ground color pale gray, slightly tinged with yellowish, with pale brown sprinkles; distinct basal, antemedial, medial, postmedial, and subterminal dark brown patches on the costal margin; rounded small terminal dots dark brown; medial area of the forewing with two rounded patches (the discal area’s one being larger than the other), dark brown and distinct; cilia pale gray, slightly tinged with yellowish. Hindwing white-gray, with an indistinct brown discal streak; cilia white-gray. Abdomen pale gray. Male genitalia (Fig. 8). Uncus tapering, weakly curved apically, with a small apical spine. Tegumen roundly rectangular, sclerotized, and not shouldered distinctly. Short, stout valva rounded, with strongly sclerotized edge; apex of valva weakly covered with setae and two short, stout processes of the same length as each other, with the process of the upper angle with a slender apical spine. Vinculum broadly “V”˗shaped, almost the same length as the uncus and tegumen. Aedeagus slightly waved, with a weakly sclerotized carina, and vesica with a large spine-shaped cornutus, and scobinate. Female genitalia. Not collected.</p><p>Distribution. Cambodia (Koh Kong Prov.).</p><p>Etymology. The specific epithet is derived from the Latin “ speculum ” meaning “ glasses ”, referring to the valva shape of male genitalia.</p></div>	https://treatment.plazi.org/id/E07687C30F72FFEBFF2C2BDBFE7CF3D8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bayarsaikhan, Ulziijargal;Cha, Yeong-Bin;Kwon, Hyung Wook;Bae, Yang-Seop	Bayarsaikhan, Ulziijargal, Cha, Yeong-Bin, Kwon, Hyung Wook, Bae, Yang-Seop (2025): Genus Halone Walker (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with two new species and a newly recorded species. Zootaxa 5660 (1): 125-133, DOI: 10.11646/zootaxa.5660.1.9, URL: https://doi.org/10.11646/zootaxa.5660.1.9
E07687C30F73FFE9FF2C2C9EFE25F7DF.text	E07687C30F73FFE9FF2C2C9EFE25F7DF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Halone hollowayi Bae & Bayarsaikhan, HT 2019	<div><p>Halone hollowayi Bae &amp; Bayarsaikhan, 2019</p><p>(Figures 5, 11, 14)</p><p>Halone hollowayi Bayarsaikhan &amp; Bae, 2019, Zootaxa, 4586 (2): 396, figs. 2a, 2b, 5, 6. TL: Cambodia (Kampot Prov., Bokor).</p><p>Type series. Holotype. [Cambodia] 1 ♂, Kampot Prov., Bokor (N10˚37′42.1″, E104˚03′53.9″), 15.I.2013 (YS Bae, YD Ju, XV Le, BS Park, HJ Lee), adult and genitalia slide no. INU-10033C . Paratypes (2 ♂, 3 ♀) Cambodia: 2 ♀, Pursat Prov., Pursat (N12˚03′46.7″, E103˚14′36.1″), 2–3.V.2010 (YS Bae, XV Le, DY Cha), adults and genitalia slides no. INU-1386C, 1932C; 1 ♂, Mondulkiri Prov., Seima (N12˚18′29″, E107˚04′55″), 11.I.2013 (YS Bae, YD Ju, XV Le, BS Park, HJ Lee), adult and genitalia slide no. INU-10034C ; 1 ♂, 1 ♀, Kampot Prov., Bokor (N10˚39′11″, E104˚03′11″, Alt.: 950m), 26–27.XII.2014 (YS Bae, YD Ju, MJ Qi, U Bayarsaikhan, BS Park, SM NA, JW Kim, DJ Lee), adults and genitalia slides no. INU-1568C, 1569C .</p><p>Distribution. Cambodia (Bae &amp; Bayarsaikhan 2019), Laos (Bayarsaikhan et al. 2021).</p><p>Remarks. This species was previously reported from Cambodia (Kampot, Mondulkiri, and Pursat provinces) by Bae &amp; Bayarsaikhan (2019).</p></div>	https://treatment.plazi.org/id/E07687C30F73FFE9FF2C2C9EFE25F7DF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bayarsaikhan, Ulziijargal;Cha, Yeong-Bin;Kwon, Hyung Wook;Bae, Yang-Seop	Bayarsaikhan, Ulziijargal, Cha, Yeong-Bin, Kwon, Hyung Wook, Bae, Yang-Seop (2025): Genus Halone Walker (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with two new species and a newly recorded species. Zootaxa 5660 (1): 125-133, DOI: 10.11646/zootaxa.5660.1.9, URL: https://doi.org/10.11646/zootaxa.5660.1.9
E07687C30F70FFE9FF2C2E99FC79F19F.text	E07687C30F70FFE9FF2C2E99FC79F19F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Halone ariadna Bucsek 2012	<div><p>Halone ariadna Bucsek, 2012</p><p>(Figures 6, 9, 15)</p><p>Halone ariadna Bucsek, 2012, Erebidae, Arctiinae of Malay Peninsula-Malaysia: 85, pl. 15: 212. TL: peninsular Malaysia (Cameron Highlands, Pahang Dist.).</p><p>Type series. [CAMBODIA] 1 ♂, 20km SE Koh Kong, Tatai river (N11˚34′, E103˚07′, Alt.: 50˗300m), 3–19.V.2005 (leg. E Jendek &amp; O. Šauša); 1 ♂, Kirirom National Park (N11˚21′, E104˚04′, Alt.: 780m), 17.I.200? (leg. M. &amp; S. Murzin, CKB) .</p><p>Distribution. Cambodia (Bucsek 2012; Bae et al. 2016), Laos, Vietnam (Bae &amp; Bayarsaikhan 2019), Peninsular Malaysia (Bucsek 2012).</p><p>Material examined. [ CAMBODIA] 1 ♂ ♂, Pursat, Pursat Prov. (N12˚05′34.2″, E103˚11′10.1″), 4. V.2010 (YS Bae, XV Le, DY Chae); 2 ♀, Mondulkiri, Mondulkiri Prov. (N12˚57′47.8″, E107˚10′52.8″), 6.X.2010 (YS Bae, XV Le, DY Chae), adults and genitalia slides no. INU-10041C, 10042C; 1 ♂, Mondulkiri, Mondulkiri Prov. (N12˚10′58.1″, E106˚58′15″), 9.X.2010 (YS Bae, XV Le, DY Chae), adults and genitalia slides no. INU-10043C; 1 ♂, Central Cardamom, Koh Kong Prov. (N12˚20′09″, E107˚18′54″), 28. VI.2011 (YS Bae, XV Le, U Bayarsaikhan, BS Park, MS Kim, KW Kim), adults and genitalia slides no. INU-10044C ; 1 ♂, Central Cardamom, Koh Kong Prov. (N11˚58′47″, E103˚22′22″), 29. VI.2011 (YS Bae, XV Le, U Bayarsaikhan, BS Park, MS Kim, KW Kim), adults and genitalia slides no. INU-1009 ; 1 ♂, 1 ♀, Samkos, Koh Kong Prov. (N12˚12′48.1″, E102˚54′12.7″), 24.XII.2014 (YS Bae, YD Ju, MJ Qi, U Bayarsaikhan, BS Park, SM Na, JW Kim, DJ Lee) .</p><p>Remarks. This species was previously reported from Cambodia (Koh Kong, Mondulkiri, and Pursat provinces) by Bucsek (2012), Bae et al. (2016), and Bae &amp; Bayarsaikhan (2019).</p></div>	https://treatment.plazi.org/id/E07687C30F70FFE9FF2C2E99FC79F19F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Bayarsaikhan, Ulziijargal;Cha, Yeong-Bin;Kwon, Hyung Wook;Bae, Yang-Seop	Bayarsaikhan, Ulziijargal, Cha, Yeong-Bin, Kwon, Hyung Wook, Bae, Yang-Seop (2025): Genus Halone Walker (Lepidoptera, Erebidae, Arctiinae) from Cambodia, with two new species and a newly recorded species. Zootaxa 5660 (1): 125-133, DOI: 10.11646/zootaxa.5660.1.9, URL: https://doi.org/10.11646/zootaxa.5660.1.9
