identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
D42CA7775415CB5BFF6DFDD8783EFA9E.text	D42CA7775415CB5BFF6DFDD8783EFA9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Adelops kiesenwetteri Dieck 1869	<div><p>Adelops kiesenwetteri Dieck, 1869 (Figs. 1, 26)</p><p>Adelops Kiesenwetteri Dieck, 1869: 350 .</p><p>Current accepted name</p><p>Troglocharinus kiesenwetteri (Dieck, 1869): Fresneda et al., 2024: 775.</p><p>Type locality</p><p>Montserrat (caves of) [original citation: “Grotten des Montserrat in Catalonien”].</p><p>Type material</p><p>Syntypes: MZB 78-2710, MZB 78-2711 (2 exs.), type loc. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Dieck (1869) neither establishes a holotype, nor states the exact number of specimens in the type series nor where it is deposited. Thanks to intercession by C. Bolívar, Zariquiey was able to study various cotypes sent by Dieck to Pérez Arcas and Escalera, that formed part of the MNCNM collection (Zariquiey, 1924: 4). Zariquiey subsequently describes the material that he studied and indicates textually “Cuevas y Avencs de Montserrat. Cova del Salitre (Tipos), siete ejemplares Co-tipos! del Museo de Madrid; un ejemplar cotipo? ex coll. Bedel...” (Zariquiey, 1924: 13). Based on this information, and in accordance with the inscription on its labels, specimens MZB 78-2710 and MZB 78-2711 from the Zariquiey’s collection, would seem to belong to this series of seven cotypes mentioned above. Both specimens have red labels with the inscriptions “Grotte pres du MONTSERRAT ” and “COTIPUS”. Specimen MZB 78-2710 also has a label with the inscription “CO-TIPOS de DIECK ex Museo de MADRID”. A third specimen (MZB 78-2712) has labels with the inscripions “ Montserrat ” and “Ex. coll. BEDEL” (the latter a red label) and seems to correspond to the specimen cited by Zariquiey as a doubtful cotype.</p><p>In the MNCNM collection (box 27) we have located a specimen from the Pérez Arcas collection with the hand-written inscription “Dieck” on the back of the label, as mentioned by Zariquiey (1924: 4). We have also found four damaged specimens that would seem to correspond to the material sent by Dieck to Escalera (none of these specimens are identified as belonging to the type series). The characteristics of the cards on which the specimens in the MCNB are mounted, as well as the transcription of their locality of origin, coincide with the those of the four above-mentioned specimens, which indicates that they all form part of the same group of specimens (with which they complete the series of seven cotypes studied by Zariquiey) .</p></div>	https://treatment.plazi.org/id/D42CA7775415CB5BFF6DFDD8783EFA9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775415CB5AFF6DF8A678CCFEAB.text	D42CA7775415CB5AFF6DF8A678CCFEAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillochlamys aurouxi Espanol 1965	<div><p>Anillochlamys aurouxi Español, 1965 (Fig. 27)</p><p>Anillochlamys aurouxi Español, 1965b: 98; figs. 3–6.</p><p>Current accepted name</p><p>Anillochlamys aurouxi Español, 1965</p><p>Type locality</p><p>La Covatilla; Cova Bartolo [original citation: “Castellón: Cueva Covatilla, Ahín, en la Sierra de Espadán ...; cueva Bartolo, Ahín, inmediata a la anterior”].</p><p>Type material</p><p>Syntypes: MZB 78-1728 / MZB 78-1742 (26 ♂♂, 33 ♀♀), La Covatilla, 18-IV-1965, I. González, L. Auroux &amp; F. Español leg. │ MZB 78-1743 / MZB 78-1762 (28 ♂♂, 36 ♀♀), Cova Bartolo, 18-IV-1965, I. González, L. Auroux &amp; F. Español leg.</p><p>Remarks</p><p>Español (1965b) refers to “numerosos ejemplares” [numerous specimens] for both localities without specifying how many or establishing a holotype. The subsequent designation of a holotype by Escolà (2000) is not valid (Article 73.1.3. ICZN 1999). This specimen (MZB 78-1728) has a red label with the inscription “Typus” hand-written by F. Español; the remaining specimens have red printed labels with the inscription “ Paratypus ”. Escolà (2000) mentions a total of 58 paratypes from La Covatilla and 63 from Cova Bartolo.</p></div>	https://treatment.plazi.org/id/D42CA7775415CB5AFF6DF8A678CCFEAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775414CB5AFF6DFCB17F93FB55.text	D42CA7775414CB5AFF6DFCB17F93FB55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillochlamys avariae Comas 1977	<div><p>Anillochlamys avariae Comas, 1977 (Fig. 28)</p><p>Anillochlamys avariae Comas, 1977a: 21; fig. 1.</p><p>Current accepted name</p><p>Anillochlamys bueni Jeannel, 1909: Salgado &amp; Fresneda, 2004a: 364.</p><p>Type locality</p><p>Cova del Forat [original citation: “Cova del Forat, Barig, Valencia ”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1709, type loc., 27-II-1977, G. Avaria &amp; A. Sendra leg.</p><p>Paratypes: MZB 78-1710 / MZB 78-1712, MZB 78-1715 (3 ♂♂, 2 ♀♀), same data as holotype. │ MZB 78-1713, MZB 78-1714 (3 ♀♀), type loc., 19-XII-1976, G. Avaria &amp; A. Sendra leg. │ MZB 78-1716 / MZB 78-1718 (3 ♂♂, 2 ♀♀), Cueva del Perro (as “ Cova del Perro ”), 27-II-1977, G. Avaria &amp; A. Sendra leg. │ MZB 78-1719 (remains 1 ♂), Avenc de la Vinya Vella (quoted as “ Avenc Vinabella ”), 14-XII-1975, J. V. González leg.</p><p>Remarks</p><p>Type series consisting of the ♂ holotype and 16 paratypes (6 ♂♂ and 10 ♀♀) (Comas, 1977a). Although the author only quotes two males from Cueva del Perro, in the MCNB collection there are in fact three males from such locality, identically labelled. The remaining paratypes are deposited in the JCC (Comas, 1977a).</p></div>	https://treatment.plazi.org/id/D42CA7775414CB5AFF6DFCB17F93FB55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775414CB5AFF6DF9EA7A55F8A8.text	D42CA7775414CB5AFF6DF9EA7A55F8A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillochlamys baguenai Jeannel 1930	<div><p>Anillochlamys baguenai Jeannel, 1930</p><p>Anillochlamys Baguenai Jeannel, 1930: 91 .</p><p>Current accepted name</p><p>Anillochlamys subtruncata Jeannel, 1930: Salgado et al., 2008: 759.</p><p>Type locality</p><p>Cova de l’Alt del Pi [original citation: “Province de Valencia: cueva del Altopino, à Barraix, prês de Estivella ”] .</p><p>Type material</p><p>Holotype: MZB 78-2156 (1 ♂), type loc., 11-VIII-1928, M. Báguena leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Male holotype, originally designated as “Type”, fixed by monotypy (Article 73.1.2.). Consists of the remains of a specimen in Zariquiey’s collection, with no appendices except the femur and tibia of the hind right leg, as described by Jeannel (1930). Español (1965c) indicated that this specimen would probably be deposited in Jeannel’s collection.</p></div>	https://treatment.plazi.org/id/D42CA7775414CB5AFF6DF9EA7A55F8A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540BCB44FF6DFB257FD1FE8F.text	D42CA777540BCB44FF6DFB257FD1FE8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillochlamys moroderi subsp. subtruncatus Jeannel 1930	<div><p>Anillochlamys moroderi subtruncatus Jeannel, 1930 (Fig. 29)</p><p>Anillochlamys Moroderi spp. subtruncatus Jeannel, 1930: 90.</p><p>Current accepted name</p><p>Anillochlamys subtruncata Jeannel, 1930: Salgado et al., 2008: 759.</p><p>Type locality</p><p>Cueva Negra [original citation: “Province de Valencia: cueva Negra, prês de Montanejos ”] .</p><p>Type material</p><p>Syntypes: MZB 78-2157 (1 ♀), type loc., 10-I-1930, R. Zariquiey leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Type series consisting of four females according to Jeannel (1930), who does not state where they are deposited. Two specimens of the type series have been located in the MNHNP’s general collection (box 52). Both are identified with the printed inscription “ TYPE ”, one in black letters on a red label and the other in red letters on a white label, as in the case of MZB 78-2157. These three specimens all have identical collection labels.</p></div>	https://treatment.plazi.org/id/D42CA777540BCB44FF6DFB257FD1FE8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540ACB44FF6DFC957A9AFB9D.text	D42CA777540ACB44FF6DFC957A9AFB9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillochlamys negrei Comas 1990	<div><p>Anillochlamys negrei Comas, 1990</p><p>Anillochlamys negrei Comas, 1990: 69; figs. 1, 4–10.</p><p>Current accepted name</p><p>Anillochlamys moroderi Bolívar, 1923: Salgado &amp; Fresneda, 2004a: 367.</p><p>Type locality</p><p>Túnel dels Sumidors [original citation: “cueva «Túnel dels Sumidors», situada en el término municipal de Vallada en la comarca de La Costera del País Valencià ”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1720, type loc., 27-IV-1969, J. Bartolomé leg.</p><p>Paratypes: MZB 78-1721 / MZB 78-1724 (2 ♂♂, 4 ♀♀), same data as holotype. │ MZB 78-1725 (1 ♀), type loc., 8-XII-1976, G. Avaria &amp; A. Sendra leg. │ MZB 78-1726 (1 ♀), type loc., 6-III-1983, R. Lluc leg. │ MZB 78-1727 (1 ♀), type loc., S.E.V.E. leg .</p><p>Remarks</p><p>Type series consisting of a ♂ holotype and 14 paratypes (4 ♂♂ and 10 ♀♀); the rest of the paratypes are deposited in the JCC (Comas, 1990) .</p></div>	https://treatment.plazi.org/id/D42CA777540ACB44FF6DFC957A9AFB9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540ACB47FF6DF9A27F20FE63.text	D42CA777540ACB47FF6DF9A27F20FE63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillochlamys raholai Zariquiey 1922	<div><p>Anillochlamys raholai Zariquiey, 1922 (Figs. 2, 30)</p><p>Anillochlamys Raholai Zariquiey, 1922b: 159; plate XI.</p><p>Current accepted name</p><p>Pseudospeonomus raholai (Zariquiey, 1922): Comas et al., 2007: 124.</p><p>Type locality</p><p>Cova Tassana [original citation: “Cova Tasana, junto a la cala Reparada y dentro de la bahía de Joncus, situada en terrenos de los Barones de la Roda y término municipal de Cadaqués (Prov. Gerona)”].</p><p>Type material</p><p>Holotype ♂: MZB 78-3018, type loc., VIII-1919, R. Zariquiey leg.</p><p>Paratypes: MZB 78-2225, MZB 78-3017 (2 ♀♀), same data as holotype .</p><p>Remarks</p><p>Male holotype originally designated as a “Tipo” (= holotype, Article 73.1.1.). Subsequently Zariquiey (1922b) cites the capture of various specimens from the type locality at the end of August 1919 (with no further details). In the MCNB collection we have found three specimens with identical collection labels and with red labels with the written inscription “COTIPO” (MZB 78-2225) or the printed inscription “Typus” (MZB 78-3017 and MZB 78-3018). Oleguer Escolà, the former curator of the MCNB arthropod collection, identified by handwritten labels the specimen MZB 78-3018 (lacking the posterior part of its abdomen) as the holotype designated by Zariquiey (1922b), indicating that its copulatory organ has been extracted and subsequently lost. The author of the description gives the length of the holotype (2.5 mm) and also indicates its relative dimensions (length and width) of the antennal segments, the relative lengths of the segments of the rear tarsus, and describes and illustrates the aedeagus of the supposed (there is no way of knowing for sure) male holotype. Neither can we be sure how Zariquiey made these measurements. We have no reason to doubt the identification of the holotype made by Escolà, who we assume must have had additional information at his disposal that enabled him to reach this conclusion.</p><p>In the MNCNM (box 2) we have located a specimen labelled as a cotype by C. Bolívar, which also has the red printed label of a syntype (nº 11491) added at a later date. Jeannel (1924a) states for this species “…type: cova Tasana (coll. Zariquiey et coll. Biospeologica)”. In the general collection of the MHNNP (box 52), we have located specimens with identical collection dates but not labelled as belonging to the type series .</p></div>	https://treatment.plazi.org/id/D42CA777540ACB47FF6DF9A27F20FE63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775409CB47FF6DFCF97F9EFAE5.text	D42CA7775409CB47FF6DFCF97F9EFAE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillochlamys raholai subsp. luisbofilli Zariquiey 1940	<div><p>Anillochlamys raholai luisbofilli Zariquiey, 1940 (Fig. 3)</p><p>Anillochlamys raholai ssp. luis-bofilli Zariquiey, 1940: 529.</p><p>Current accepted name</p><p>Pseudospeonomus raholai (Zariquiey, 1922): Comas et al., 2007: 124.</p><p>Type locality</p><p>Cova de l’Aigua de Malaterra [original citation: “Cueva de Mala-terra, junto al Puig del Áliga, termino municipal de Rosas, provincia de Gerona”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2913, type loc. (ex coll. R. Zariquiey).</p><p>Paratypes: MZB 78-1763, MZB 78-2914 (2 ♀♀), same data as holotype (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Male holotype designated originally by Zariquiey (1940) as “Tipo” (Article 73.1.1.), deposited in his collection. Subsequently, he reports the collection in a trap of eight specimens without specifying their sex. We have only been able to find three of these specimens (1 ♂ and 2 ♀♀), with a collection date of November 1934. Each has a red label with the hand-written inscription “TIPO”. Oleguer Escolà has identified the male as the holotype designated by Zariquiey, and has added the respective red holotype and paratype labels to these specimens. Unless evidence to contrary comes to light, we accept as a holotype the specimen identified as such by Escolà. In “box 52” of the MNHNP collection, we have found a male labelled “COTYPE” with identical collection labels .</p></div>	https://treatment.plazi.org/id/D42CA7775409CB47FF6DFCF97F9EFAE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775409CB46FF6DF87A7F8AFED3.text	D42CA7775409CB46FF6DF87A7F8AFED3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anillochlamys urgellesi Espanol 1965	<div><p>Anillochlamys urgellesi Español, 1965 (Fig. 31)</p><p>Anillochlamys urgellesi Español, 1965c: 452; figs. 2, 3.</p><p>Current accepted name</p><p>Paranillochlamys urgellesi (Español, 1965): Salgado &amp; Fresneda, 2004a: 373.</p><p>Type locality</p><p>Cova Bonica [original citation: “Tarragona: Cova Bonica, sobre la Font dels Fumats, cerca de la cumbre del Montsiá”].</p><p>Type material</p><p>Syntypes: MZB 78-1764, MZB 78-1765 (1 ♂, 1 ♀), type loc., 16-VIII-1964, I. González &amp; A. Avalo leg. │ MZB 78-1766 / MZB 78-1772 (4 ♂♂, 3 ♀♀), type loc., 20-IX-1964, F. Español leg.</p><p>Remarks</p><p>Type series consisting of nine specimens (Español, 1965c), all deposited in the MCNB. On the collection labels of specimens MZB 78-1764 and MZB 78-1765 the date is given as August 1964; the former has a red label with the inscription “ Holotypus ” hand-written by F. Español. The remaining specimens have red labels with the inscription “ Paratypus ” hand-written by the author, who, nevertheless, does not establish a holotype in the original description. The subsequent designation of a holotype by Escolà (2000) is not valid (Article 73.1.3.). Escolà (2000) excludes from the type series the female specimen dated from August 1964 referred to by Español (1965) .</p></div>	https://treatment.plazi.org/id/D42CA7775409CB46FF6DF87A7F8AFED3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775408CB46FF6DFC697A7CFB9D.text	D42CA7775408CB46FF6DFC697A7CFB9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Antrocharidius orcinus subsp. acevedoi Espanol 1953	<div><p>Antrocharidius orcinus acevedoi Español, 1953 (Fig. 32)</p><p>Antrocharidius orcinus ssp. acevedoi Español, 1953a: 94 .</p><p>Current accepted name</p><p>Troglocharinus orcinus acevedoi (Español, 1953): Rizzo &amp; Comas, 2015: 109.</p><p>Type locality</p><p>Forat del Castell de la Formiga [original citation: “Forat del Castell de la Formiga, sobre Vilaverd”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2133, type loc., F. Español leg.</p><p>Paratypes: MZB 78-2134 / MZB 78-2139 (5 ♂♂, 2 ♀♀), type loc., F. Español, M. Acevedo, I. Fernández &amp; E. Schütte leg. │ MZB 78-2140 / MZB 78-2144 (4 ♂♂, 1 ♀), Avenc del Roc de les Abelles, F. Español, M. Acevedo, I. Fernández &amp; E. Schütte leg.</p><p>Remarks</p><p>Male holotype originally designated as a “Tipo” (Article 73.1.1.). Español (1953a) does not give the number of specimens in the type series, merely mentioning two lots from the two named caves. There is no record of any material deposited in any other collection (Español, 1953a only explicitly refers to the deposit of the holotype in the MCNB).</p></div>	https://treatment.plazi.org/id/D42CA7775408CB46FF6DFC697A7CFB9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775408CB46FF6DF9A27AB9F844.text	D42CA7775408CB46FF6DF9A27AB9F844.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Antrocharidius orcinus subsp. lagari Espanol 1953	<div><p>Antrocharidius orcinus lagari Español, 1953 (Fig. 33)</p><p>Antrocharidius orcinus ssp. lagari Español, 1953a: 93; figs. 2–4.</p><p>Current accepted name</p><p>Troglocharinus orcinus lagari (Español, 1953): Rizzo &amp; Comas, 2015: 109.</p><p>Type locality</p><p>Avenc Nou de la Font Freda [original citation: “avenc Nou de la Font Freda, junto al caserio del Bonretorn, en las estribaciones sud-orientales de la sierra de la Musara, entre l’Albiol y Alcover”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2019, type loc., 1-X-1951, F. Español leg.</p><p>Paratypes: MZB 78-1970 / MZB 78-2018, MZB 78-2020, MZB 78-2021 (108 ♂♂, 144 ♀♀), type loc., 1-X-1951, F. Español, A. Lagar, E. Schütte &amp; I. Fernández leg.</p><p>Remarks</p><p>Male holotype originally designated as a “Tipo” (Article 73.1.1.). Español (1953a) does not detail the number of specimens in the type series nor indicates where it is deposited; he only indicates that the holotype is in the MCNB. Twenty-six paratypes (12 ♂♂ and 14 ♀♀) are deposited in the FLC (Fresneda &amp; Salgado, 2016); a male paratype is in the MNCNM collection.</p></div>	https://treatment.plazi.org/id/D42CA7775408CB46FF6DF9A27AB9F844	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540FCB41FF6DFD907E12FD22.text	D42CA777540FCB41FF6DFD907E12FD22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aranzadiella leizaolai Espanol 1972	<div><p>Aranzadiella leizaolai Español, 1972</p><p>Aranzadiella leizaolai Español, 1972a: 61; figs. 1–3.</p><p>Current accepted name</p><p>Aranzadiella leizaolai Español, 1972</p><p>Type locality</p><p>Sima Kobeta [original citation: “sima de Kobeta, Olatz, en el extremo NW. de Guipúzcoa ”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1923, type loc., 18-VII-1966, F. Leizaola leg.</p><p>Paratypes: MZB 78-1924 (2 ♀♀), same data as holotype .</p><p>Remarks</p><p>Español (1972a) does not specify the number of paratypes (all females), indicating that these specimens were deposited in the collections of the MCNB and the Aranzadi Museum in San Sebastián (currently, the SCA).</p></div>	https://treatment.plazi.org/id/D42CA777540FCB41FF6DFD907E12FD22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540FCB41FF6DFE3A7ECCF97C.text	D42CA777540FCB41FF6DFE3A7ECCF97C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathysciola (Bathysciola) breuili Bolivar 1921	<div><p>Bathysciola (Bathysciola) breuili Bolívar, 1921 (Figs. 4, 34)</p><p>Bathysciola (s. str.) breuili Bolívar, 1921: 525; fig. 18.</p><p>Current accepted name</p><p>Zariquieyella breuili (Bolívar, 1921): Fresneda et al., 2024: 691.</p><p>Type locality</p><p>Cueva de Landarbaso [original citation: “Provincia de Guipúzcoa: cueva de Landarbaso (término municipal de Rentería, partido de San Sebastián)”].</p><p>Type material</p><p>Paratypes: MZB 78-2239 (1 ♂), type loc., 18-VIII-1919, H. Breuil &amp; R. Jeannel leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Bolívar (1921) cites various specimens collected by H. Breuil and R. Jeannel on 18 August 1919 deposited in the Bioespeologica collection and in the Madrid Museum. The author originally designated a male holotype using the term “Tipo” (Article 73.1.1.), indicating that it was deposited in the Biospeologica collection. Fresneda &amp; Salgado (2006) studied three and two specimens from the type series, deposited, respectively, in the MNHNP general collection (box 60) and the MNCNM (box 16). The inscription on their labels is transcribed by Fresneda &amp; Salgado (2006, 2016). The specimens in the MNCNM collection (male and female) were not originally identified as belonging to the type series. The male MZB 78-2239 has collection labels that are identical to those of the other collections mentioned here, with the inscription “TYPE” in red letters on a white label. Although the holotype was deposited in the Biospeologica collection, we are not aware that it has been located or identified from amongst the specimens from the MNHNP (Fresneda &amp; Salgado, 2006 refer to the group of studied specimens as a type series).</p></div>	https://treatment.plazi.org/id/D42CA777540FCB41FF6DFE3A7ECCF97C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540FCB40FF6DFA4B7A3DFDF3.text	D42CA777540FCB40FF6DFA4B7A3DFDF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathysciola diegoi Salgado & Fresneda 2001	<div><p>Bathysciola diegoi Salgado &amp; Fresneda, 2001: 184; figs. 1–4.</p><p>Current accepted name</p><p>Zariquieyella diegoi (Salgado &amp; Fresneda, 2001): Fresneda et al., 2024: 691.</p><p>Type locality</p><p>Cueva de Mendia Landa [original citation: “cueva de Mendia Landa ... en las proximidades de Arive, cerca de la carretera que une esta población con el pueblo de Aria, valle de Aézkoa (Navarra), cuenca hidrogràfica del río Irati”].</p><p>Type material</p><p>Paratypes: MZB 2004-0949 (1 ♀), type loc., 19-VII-2000, D. Rodríguez, J. M. Salgado &amp; J. Fresneda leg.</p><p>Remarks</p><p>Type series consisting of 5 ♂♂ and 8 ♀♀ (Salgado &amp; Fresneda, 2001). Specimens from the type series deposited in other collections: ♂ holotype in the FLC; paratypes in CZULE, FLC and IBE (Salgado &amp; Fresneda, 2001; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA777540FCB40FF6DFA4B7A3DFDF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540ECB40FF6DFF897F7AFAE5.text	D42CA777540ECB40FF6DFF897F7AFAE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathysciola fadriquei Fresneda & Comas 2007	<div><p>Bathysciola fadriquei Fresneda &amp; Comas, 2007 (Fig. 35)</p><p>Bathysciola fadriquei Fresneda &amp; Comas, 2007: 326; figs. 1–3, 4, 9, 10.</p><p>Current accepted name</p><p>Zariquieyella fadriquei (Fresneda &amp; Comas, 2007): Fresneda et al., 2024: 691.</p><p>Type locality</p><p>Cueva de Lizara [original citation: “Huesca, Jaca,Aragüés del Puerto, Cueva de Lizara, UTM: X 694400-Y 4737375- Z 1.470 m (datum europeo de 1950)”].</p><p>Type material</p><p>Paratypes: MZB 2008-0056 / MZB 2008-0059, MZB 2008-0638, MZB 2008-0639 (3 ♂♂, 3 ♀♀), type loc., 12-IV-2006 / 10-IX-2006 (trap), F. Fadrique leg.</p><p>Remarks</p><p>Type series consisting of 14 ♂♂ and 15 ♀♀ from two cavities in the Huescan Pyrenees (Fresneda &amp; Comas, 2007). Specimens of the type series deposited in other collections: male holotype in the JCC; paratypes in the CZULE, FLC, JCC and MNCNM (Fresneda &amp; Comas, 2007; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA777540ECB40FF6DFF897F7AFAE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540ECB43FF6DF87A7953FE47.text	D42CA777540ECB43FF6DF87A7953FE47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathysciola (Bathysciola) mystica Fresneda & Fery 2009	<div><p>Bathysciola (Bathysciola) mystica Fresneda &amp; Fery, 2009</p><p>Bathysciola (Bathysciola) mystica Fresneda &amp; Fery, 2009: 4; figs. 5, 7, 9, 11, 13.</p><p>Current accepted name</p><p>Zariquieyella mystica (Fresneda &amp; Fery, 2009): Fresneda et al., 2024: 691.</p><p>Type locality</p><p>Plan Batalhèr [original citation “Plan Batalher, Vielha-Mijaran, Lleida, España, 1.230 m. UTM: 31T 317700 473277”].</p><p>Type material</p><p>Paratypes: MZB 87-6299 (2 ♂♂, 1 ♀), Les, 26-III-1916, F. Léon-Hilaire leg. (coll. R. Zariquiey)│ MZB 87-6298 (1 ♂, 2 ♀♀), Les, 30-XI-1916, F. Léon-Hilaire leg. (coll. R. Zariquiey)│ MZB 87-6308 (1 ♂, 1 ♀), Les, 16-I-1916, F. Léon-Hilaire leg. (coll. R. Zariquiey) │ MZB 87-6307 (2 ♀♀), Bossòst, 14-VII-1916, F. Léon-Hilaire leg. (coll. R. Zariquiey)│ MZB 87-6306 (4 ♂♂, 4 ♀♀), Val d’Aran, VIII-1916, R. Zariquiey leg. (coll. R. Zariquiey)│ MZB 87-6302 (7 ♂♂, 1 ♀), Grotte de Gourgue, 20-XII-1919, Ch. Fagniez leg. │ MZB 87-6310 (9 ♂♂, 2 ♀♀), Bagnères-de-Luchon (“feuilles seches sur noisetière”), H. du Buysson leg.</p><p>Remarks</p><p>Type series consisting of 81 specimens from various localities in the French departments of Ariège and Haute-Garonne, and from the Val d’Aran (Lleida province, Spain) (Fresneda &amp; Fery, 2009). Most specimens from the Val d’Aran are in the MCNB collection and were previously referred to (see Fresneda &amp; Fery, 2009) and identified on their labels as Bathysciola schiodtei (Kiesenwetter, 1850) . Specimens from the type series deposited in other collections (Fresneda &amp; Fery, 2009; Fresneda &amp; Salgado, 2016): holotype in the FLC; paratypes in AFC, CVC, FLC, HFC, IBE, MPC and ZSM.</p></div>	https://treatment.plazi.org/id/D42CA777540ECB43FF6DF87A7953FE47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540DCB43FF6DFCDD7982FA81.text	D42CA777540DCB43FF6DFCDD7982FA81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathysciola (Bathysciola) obermaieri Bolivar 1918	<div><p>Bathysciola (Bathysciola) obermaieri Bolívar, 1918 (Figs. 5, 36)</p><p>Bathysciola (s. str.) obermaieri Bolívar, 1918: 447; figs. 1–4.</p><p>Current accepted name</p><p>Zariquieyella obermaieri (Bolívar, 1918): Fresneda et al., 2024: 691.</p><p>Type locality</p><p>Cueva de la Ermita de Santa Elena [original citation: “Provincia de Huesca: cueva de la Ermita de Santa Elena, cerca de Biescas”].</p><p>Type material</p><p>Paratypes: MZB 78-2490 (1 ♂), type loc., 24-VII-1918 (ex coll. R. Zariquiey).</p><p>Remarks</p><p>The type series consists of 20 specimens (Bolívar, 1918), of which only the holotype ♂ is indicated as having been deposited in the MNCNM collection (originally designated as “Tipo”, Article 73.1.1.). Specimen MZB 78- 2490 deposited in the MCNB collection has printed capture labels (only the collection date is hand-written) and a determination label hand-written by C. Bolivar with the inscription “co-TIPO”, as well as a red label printed with “Cotypus”. The inscription on the labels is practically identical to that transcribed by Fresneda &amp; Salgado (2006, 2016) for the above-mentioned specimens from the MNCNM (2 ♂♂ and 2 ♀♀) and MNHNP (two specimens); these six specimens bear determination labels hand-written by Bolívar with the inscription “TIPO”. In our visit to the MNCNM we found the holotype and four paratypes (box 16), duly identified by red printed labels added at a later date. As well, we found five other specimens with identical capture labels that, although not labelled as part of the type series, probably do form part of that series.</p></div>	https://treatment.plazi.org/id/D42CA777540DCB43FF6DFCDD7982FA81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540DCB42FF6DF8A67FD0FCDB.text	D42CA777540DCB42FF6DF8A67FD0FCDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bathysciola (Bathysciola) zariquieyi Bolivar 1919	<div><p>Bathysciola (Bathysciola) zariquieyi Bolívar, 1919 (Figs. 6A–C)</p><p>Bathysciola (s. str.) zariquieyi Bolívar, 1919: 18; figs. 1–4.</p><p>Current accepted name</p><p>Catalaniana zariquieyi zariquieyi (Bolívar, 1919): Fresneda et al., 2024: 413.</p><p>Type locality</p><p>Vallvidrera [original citation: “Vallvidrera (Barcelona)”].</p><p>Type material</p><p>Paratypes: MZB 78-2546 (1 ♂), type loc., Sres. Zariquiey leg. (ex coll. R. Zariquiey) │ MZB 78-2547 (1 ♂), Montseny, Sres. Zariquiey leg. (ex coll. R. Zariquiey) .│ MZB 78-1447 (1 ♀), Vilafranca del Penedès [quoted as “Villafranca del Panadés”], Sres. Zariquiey leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Bolívar (1919) originally designated as the holotype a male from Vallvidrera (using the term “Tipo”, Article 73.1.1.). Subsequently, he mentions two other localities, Montseny and Vilafranca del Penedès, also in the province of Barcelona. In light of the range in body length given in the description, the studied material originally consisted of a number of specimens (Bolívar, 1919, states neither how many specimens nor the place of deposit). We thus deduce that the author based his description of this new taxon on specimens from these three localities.</p><p>Box 16 in the MNCNM collection contains the specimen from the type series whose labels are transcribed by Fresneda &amp; Salgado (2006, 2016); in our visit to this collection we observed that a red label with the printed inscription “ Holotipo ” has been added. We also found at least one specimen from Montseny (not identified as belonging to the type series) whose label of origin is almost identical to that of specimen MZB 78-2547 and to the type specimen deposited in the MNHNP (box 59) mentioned by Fresneda &amp; Salgado (2006, 2016). The collection dates on these labels are, respectively, June 1916 (MNCNM), 9 June 1916 (MNHNP) and 8 June 1916 (MCNB).</p><p>Specimens MZB 78-2546 and MZB 78-2547 have labels of origin hand-written by Zariquiey and red labels with the printed inscription “ Typus ”. Specimen MZB 78-2546 also has a determination label hand-written by Bolívar with the inscription “TIPOS”; its collection label contains the date “XI-15” [November 1915], possibly due to an error (the original description only states that the material was collected in April and June). The specimen MZB 78-1447 bears a label with the hand-written inscription “TIPO” and other red label with the inscription “CoTypus” (the letters “Co” handwritten, and the word “ Typus ” printed). The non-printed part of the collection label was hand-written by Zariquiey (collection date 24 june 1917). This specimen has been cited by Lagar (1954), Fresneda &amp; Salgado (2006) and Fresneda &amp; Salgado (2016) who also transcribed its associated labels.</p></div>	https://treatment.plazi.org/id/D42CA777540DCB42FF6DF8A67FD0FCDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540CCB42FF6DFE617FA9F8CE.text	D42CA777540CCB42FF6DFE617FA9F8CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Breuilia cendreroi Bolivar 1923	<div><p>Breuilia cendreroi Bolívar, 1923</p><p>Breuilia cendreroi Bolívar, 1923: 424 .</p><p>Current accepted name</p><p>Espanoliella tibialis (Jeannel, 1909): Guéorguiev, 1976: 126.</p><p>Type locality</p><p>Cueva del Castillo [original citation: “Cueva del Castillo, cerca de Noja ... término municipal de Castillo de las Siete Villas ... partido judicial de Santoña, provincia de Santander”].</p><p>Type material</p><p>Paratypes: MZB 78-2484, MZB 78-2485 (2 ♂♂), type loc., O. Cendrero leg. (ex coll. R. Zariquiey). │ MZB 78- 2226, MZB 78-2227 (1 ♂, 1 ♀), Cueva de Paulino CoterÓn, O. Cendrero leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Bolívar (1923) does not specify the number of specimens in the type series, that originate from five localities in Cantabria (he reports “numerosos ejemplares de esta especie recogidos por el Sr. Cendrero” [numerous specimens of this species collected by Sr. Cendrero]). The specimens deposited in the MCNB collection are identified as cotypes on their determination labels, hand-written by Bolívar; specimens MZB 78-2226 and MZB 78-2227 also have red cotype labels. A male specimen with accession number MZB 78-2228 exists from the cave Paulino Coterón, with an identical collection label and a label with the inscription “prép nº 59”, but is not identified as belonging to the type series. The ♂ holotype, originally designated as “Tipo” (Article 73.1.1.) is deposited in MNCNM (Bolívar, 1923). Along with this specimen (box 14), we have also found a number of specimens from the localities recorded by Bolívar (1923), probably belonging to the type series, although they are not labelled as such. In the MNHNP general collection (box 55) we have found at least one specimen from the type locality labelled as a cotype.</p></div>	https://treatment.plazi.org/id/D42CA777540CCB42FF6DFE617FA9F8CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777540CCB4CFF6DFA567F1AFE63.text	D42CA777540CCB4CFF6DFA567F1AFE63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Breuilia tibialis Jeannel 1909	<div><p>Breuilia tibialis Jeannel, 1909 (Fig. 37)</p><p>Breuilia tibialis Jeannel, 1909: 469 .</p><p>Current accepted name</p><p>Espanoliella tibialis (Jeannel, 1909): Guéorguiev, 1976: 126.</p><p>Type locality</p><p>Cueva de San Roque [original citation: “... cueva de San Roque, à Valle (provincia de Santander)”].</p><p>Type material</p><p>Syntypes: MZB 78-2229 (1 ♂, 1 ♀), type loc., VIII-1909, H. Breuil leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>The type series consists of a large number of specimens (Jeannel, 1909) originally deposited in the Biospeologica collection (Jeannel, 1924a). The specimens in the MCNB (mounted on the same pin) have a label with the determination followed by the term “ TYPE ” hand-written in red letters, and hand-written collection labels. In box 55 of the general collection of the MNHNP we have found a pin with a number of labels including the three original labels associated with the specimens MZB 78-2229, all identical and written by the same hand. Of note too, is a red label with the inscription “ TYPE ” printed in black letters. In the same collection we have found a second pin bearing a white label printed with “ TYPE ” in red letters. Otherwise, in the MNCNM collection (box 14) we have found a specimen labelled as a cotype (a white label with the hand-written inscription “Co TYPE ” in red, followed by another a red label with the printed inscription “Cotipo”). These specimens’ collection labels are identical to those of the specimens MZB 78-2229. It is worth noting that the collector of the specimens does not figure on the capture labels of the studied material (instead, the inscription “R. Jeannel” appears printed).</p></div>	https://treatment.plazi.org/id/D42CA777540CCB4CFF6DFA567F1AFE63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775402CB4CFF6DFCF97AF2FA81.text	D42CA7775402CB4CFF6DFCF97AF2FA81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Breuilia urdialensis Bolivar 1917	<div><p>Breuilia urdialensis Bolívar, 1917</p><p>Breuilia urdialensis Bolívar, 1917: 210; fig. 6.</p><p>Current accepted name</p><p>Espanoliella urdialensis (Bolívar, 1917): Guéorguiev, 1976: 127.</p><p>Type locality</p><p>Cueva de Mingobalsa [original citation: “Provincia de Santander: Cueva de Mingobalsa, en Los Corrales, cerca de Castro-Urdiales”].</p><p>Type material</p><p>Paratypes: MZB 71-9606 (1 ♀), MZB 78-2483 (1 ♂), type loc., IX-1916, C. Bolívar leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Bolívar (1917) does not state the number of specimens in the type series, and only indicates that the male “Tipo” (= holotype, Article. 73.1.1.) is deposited in his collection (currently part of the MNCNM collection). Subsequently, he mentions material he collected from the type locality and five other caves in September 1916. The specimens in the MCNB collection have printed collection labels (the collection date, though, is hand-written), as well as a red label with the printed inscription “Cotypus”. The male MZB 78-2483 has a determination label hand-written by Bolívar with the inscription “co-TIPOS”. Along with the holotype we have found in the MNCNM (box 14) a series of specimens from the caves mentioned by Bolívar but not identified as part of the type series. In the MNHNP general collection (box 55), we have found at least one specimen from the type locality labelled “cotype” with similar collection labels .</p></div>	https://treatment.plazi.org/id/D42CA7775402CB4CFF6DFCF97AF2FA81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775402CB4FFF6DF8A6798DFF1B.text	D42CA7775402CB4FFF6DF8A6798DFF1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Breuilites eloyi Salgado 1980	<div><p>Breuilites eloyi Salgado, 1980</p><p>Breuilites eloyi Salgado, 1980a: 160, figs. 1–5.</p><p>Current accepted name</p><p>Breuilites eloyi Salgado, 1980</p><p>Type locality</p><p>Cueva del Sidrón [original citation: “Cueva del Sidrón ... en Borines (Oviedo)”].</p><p>Type material</p><p>Paratypes: MZB 78-1921, MZB 78-1922 (2 ♂♂, 3 ♀♀), type loc., 13-V-1979, E. Samartino leg.</p><p>Remarks</p><p>Type series consisting of 6 ♂♂ and 10 ♀♀, with four paratypes (2 ♂♂ and 2 ♀♀) deposited in the MCNB collection according to Salgado (1980a). In the MCNB collection there are three females mounted on the same pin, together with one of the males (MZB 78-1922). The other male (MZB 78-1921) is mounted on another pin with its genitalia preserved in Canada balsam. Specimens belonging to the type series deposited in other collections: holotype, allotype and paratypes (CZULE) (Salgado, 1980a).</p></div>	https://treatment.plazi.org/id/D42CA7775402CB4FFF6DF8A6798DFF1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775401CB4FFF6DFC217F5FFC29.text	D42CA7775401CB4FFF6DFC217F5FFC29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cantabrogeus antimachus Salgado & Luque 2012	<div><p>Cantabrogeus antimachus Salgado &amp; Luque, 2012</p><p>Cantabrogeus antimachus Salgado &amp; Luque, 2012 in Salgado et al.: 33; figs. 1, 19, 20, 26.</p><p>Current accepted name</p><p>Cantabrogeus antimachus Salgado &amp; Luque, 2012</p><p>Type locality</p><p>Cueva de Covallarco (main access) [original citation: “ España, Cantabria, Carcabal-Merilla, T. M. de San Roque de Riomiera, cueva de Covallarco, (acceso principal), ED50-UTM: 30TVN 4052289764, 400 m ”].</p><p>Type material</p><p>Paratypes: MZB 2013-1841, MZB 2017-0319, MZB 2017-0320 (3 ♀♀), Cueva de Covallarco, third entry (“3ª entrada (ED50 -UTM: 30TVN 4058789790, 420 m)”), 26-IX-2009, C. G. Luque &amp; E. Muñoz leg.</p><p>Remarks</p><p>Type series consisting of 183 specimens, all females (parthenogenetic species) (Salgado et al., 2012). Specimens from the type series deposited in other collections: holotype in the CZULE; paratypes in the entomological collections of the authors of the description and in the collections of the BMNHL, CZULE, FLC, IBE, MCNA, MNCNM and PMGC (Salgado et al., 2012; Cieslak et al., 2014; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775401CB4FFF6DFC217F5FFC29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775401CB4FFF6DF93E7A89F934.text	D42CA7775401CB4FFF6DF93E7A89F934.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cantabrogeus cultellus Salgado & Luque 2012	<div><p>Cantabrogeus cultellus Salgado &amp; Luque, 2012</p><p>Cantabrogeus cultellus Salgado &amp; Luque, 2012 in Salgado et al.: 36; figs. 2, 8, 9, 13, 17, 21, 22, 27.</p><p>Current accepted name</p><p>Cantabrogeus cultellus Salgado &amp; Luque, 2012</p><p>Type locality</p><p>Cueva de la Matorra IV [original citation: “ España, Cantabria, Solares, T. M. de Medio Cudeyo, cueva de la Matorra IV, ED50-UTM: 30TVP 4016503440, 60 m ”] .</p><p>Type material</p><p>Paratypes: MZB 2013-1837, MZB 2017-0314 / MZB 2017-0318 (3 ♂♂, 3 ♀♀), type loc., 15-XI-2008, C. G. Luque &amp; J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of 148 ♂♂ and 182 ♀♀ from five cavities in Cantabria (Salgado et al., 2012). Specimens of the type series deposited in other collections: ♂ holotype in the CZULE; paratypes in the entomological collection of the authors of the description and in the CZULE, FLC, IBE, MCNA, MNCNM and PMGC (Salgado et al., 2012; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775401CB4FFF6DF93E7A89F934	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775401CB49FF6DFA4C7A3DFE8F.text	D42CA7775401CB49FF6DFA4C7A3DFE8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cantabrogeus fresnedai Salgado & Luque 2012	<div><p>Cantabrogeus fresnedai Salgado &amp; Luque, 2012</p><p>Cantabrogeus fresnedai Salgado &amp; Luque, 2012 in Salgado et al.: 38; figs. 3, 10, 14, 23, 28.</p><p>Current accepted name</p><p>Cantabrogeus fresnedai Salgado &amp; Luque, 2012</p><p>Type locality</p><p>Cueva de La Peña Caborco [original citation: “ España, Cantabria, Esles, T. M. de Sta. Mª de CayÓn, cueva de La Peña Caborco, ED 50-UTM: 30TVN 3431591964, 260 m ”] .</p><p>Type material</p><p>Paratypes: MZB 2013-1839, MZB 2017-0321, MZB 2017-0322 (1 ♂, 2 ♀♀), type loc., 30-VI-2010, L. Labrada &amp; C. G. Luque leg.</p><p>Remarks</p><p>Type series consisting of 12 ♂♂ and 26 ♀♀ (Salgado et al., 2012). Specimens of the type series deposited in other collections: ♂ holotype in the CZULE; paratypes in the entomological collections of the authors of the description and in the CZULE, FLC, IBE, MCNA, MNCNM and PMGC (Salgado et al., 2012; Cieslak et al., 2014; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775401CB49FF6DFA4C7A3DFE8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775407CB49FF6DFC957A10FB9D.text	D42CA7775407CB49FF6DFC957A10FB9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Espanoliella luquei Salgado & Fresneda 2005	<div><p>Espanoliella luquei Salgado &amp; Fresneda, 2005</p><p>Espanoliella luquei Salgado &amp; Fresneda, 2005: 88; figs. 1–9.</p><p>Current accepted name</p><p>Amphogeus luquei (Salgado &amp; Fresneda, 2005): Fresneda et al., 2024: 508.</p><p>Type locality</p><p>Cueva del Polvorín [original citation: “Polvorín or Merino Cave, Barrio of Dueso, Santoña — Cantabria (España)”].</p><p>Type material</p><p>Paratypes: MZB 2010-0173, MZB 2017-1265 / MZB 2017-1267 (2 ♂♂, 2 ♀♀), type loc., 31-V-2003, D. Rodríguez &amp; J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of 66 ♂♂ and 99 ♀♀ from two cavities in Cantabria (Salgado &amp; Fresneda, 2005). The ♂ holotype is deposited in the CZULE (Fresneda &amp; Salgado, 2016); other paratypes in the following collections: CZULE, FLC, FMNHC, MCNA, MNCNM, MNHNP, PMGC and RMC, (Salgado &amp; Fresneda, 2005; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775407CB49FF6DFC957A10FB9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775407CB49FF6DF9A278F7F8A8.text	D42CA7775407CB49FF6DF9A278F7F8A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Fresnedaella lucius Salgado, Labrada & Luque 2011	<div><p>Fresnedaella lucius Salgado, Labrada &amp; Luque, 2011</p><p>Fresnedaella lucius Salgado, Labrada &amp; Luque, 2011: 7; figs. 1, 2.</p><p>Current accepted name</p><p>Fresnedaella lucius Salgado, Labrada &amp; Luque, 2011</p><p>Type locality</p><p>Cueva de La Canal de la Cubía [original citation: “ España, Cantabria, barrio de la Cubía, término municipal de Selaya, Cueva de La Canal de la Cubía, coordenadas ED50-UTM 30TVP3792286405, altitud 450 m s.n.m.”].</p><p>Type material</p><p>Paratypes: MZB 2013-1838, MZB 2017-0311 / MZB 2017-0313 (2 ♂♂, 2 ♀♀), type loc., 15-VI-2010, Labrada &amp; Glez. Luque leg.</p><p>Remarks</p><p>Type series consisting of 31 ♂♂ and 49 ♀♀ (Salgado et al., 2011). Specimens of the type series deposited in other collections: ♂ holotype deposited in the CZULE; paratypes in the entomological collections of the authors of the description and in the AFC, CZULE, FLC, IBE (for molecular study), MCNA, MNCNM, MNHNP and PMGC, (Salgado et al., 2011; Cieslak et al., 2014; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775407CB49FF6DF9A278F7F8A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775406CB48FF6DFD907A1DFCDE.text	D42CA7775406CB48FF6DFD907A1DFCDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Notidocharis espanyoli Comas 1987	<div><p>Notidocharis espanyoli Comas, 1987 (Fig. 38)</p><p>Notidocharis espanyoli Comas, 1987: 135; figs. 1–7.</p><p>Current accepted name</p><p>Uhagoniella franzi (Jeannel, 1956): Fresneda et al., 2024: 166.</p><p>Type locality</p><p>Cueva de la Pruneda [original citation: “ Cueva de la Pruneda, Bobia de Abajo, Asturias ”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2217, type loc., 2-VIII-1976, J. Comas leg.</p><p>Paratypes: MZB 78-2218 (1 ♀), Cueva de Merín, 7-VIII-1976, J. Comas leg.</p><p>Remarks</p><p>On the label of the holotype, the date of collection is given as 2 August 1986; type series consisting of 3 ♂♂ and 4 ♀♀ from three localities in Asturias (Comas, 1987). The remaining paratypes are deposited in the EVC and JCC (Comas, 1987).</p></div>	https://treatment.plazi.org/id/D42CA7775406CB48FF6DFD907A1DFCDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775406CB48FF6DFE667862F861.text	D42CA7775406CB48FF6DFE667862F861.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paranillochlamys velox Zariquiey 1940	<div><p>Paranillochlamys velox Zariquiey, 1940 (Figs. 7, 39)</p><p>Paranillochlamys velox Zariquiey, 1940: 532; figs. 1–6.</p><p>Current accepted name</p><p>Paranillochlamys velox Zariquiey, 1940</p><p>Type locality</p><p>Avenc de la Fotx [original citation: “Avenc del Fotx, junto a Tivissa, provincia de Tarragona”].</p><p>Type material</p><p>Syntypes: MZB 78-2158 (1 ♂), 78-3016 (1 ♀), type loc., F. Español leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>The type series consist of two males and a female, deposited in the collection of the Barcelona Museum and in the collection of the author of the description (Zariquiey, 1940). We have located a male and a female with capture labels hand-written by Zariquiey and red labels with the hand-written inscription (probably by Zariquiey himself) “TIPO”. In the MNHNP general collection (box 52) there is a specimen labelled as follows:</p><p>[ Paratypus] (hand-written by F. Español on a red label) [CATALONIA / Tivissa /Avenc Fotx] (printed) [Español leg.] [ Paranillochlamys / velox nov. gen. n. sp. / Zariquiey] (the latter two labels hand-written by Español) .</p><p>It is likely, although we cannot be absolutely sure, that the labels with the inscription “TIPO” of the specimens deposited in the MCNB correspond to an attempt to designate a male holotype and a female allotype (this would explain the designation of “ paratype ” for the specimen deposited in the MNHNP). The capture labels of the two specimens in the MCNB collection give the date 3 February 1935 . In the general collection other specimens, captured by F. Español at the same site, have been located, most with a capture date of December 1935 according to the labels (not labelled as types). This date is posterior to the VI International Entomological Congress held in Madrid in September 1935, whose proceedings, published in 1940, include the description of this new taxon. Each male specimen in the series is labelled with the symbol ♂ followed by a correlative number from 1 to 12 (the male MZB 78-2158 is number 8 of the series). Español (1950b) cites material captured in December 1934 without specifying how many specimens. This material has not been located in the MCNB collection —unless it corresponds to the above-mentioned specimens, most labelled December 1935 .</p></div>	https://treatment.plazi.org/id/D42CA7775406CB48FF6DFE667862F861	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775405CB4BFF6DFD907ED9FC0A.text	D42CA7775405CB4BFF6DFD907ED9FC0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parvospeonomus cruillensis Comas 2021	<div><p>Parvospeonomus cruillensis Comas, 2021</p><p>Parvospeonomus cruillensis Comas, 2021: 30; figs. 1b, 2b, 3b, 4b.</p><p>Current accepted name</p><p>Parvospeonomus cruillensis Comas, 2021</p><p>Type locality</p><p>Cova de Cruïlles [original citation: “Cv. Cruïlles, Aiguafreda (Vallès Oriental) … ETRS89 Fus: 31N / 438339, 4625636 / 678 m ”].</p><p>Type material</p><p>Paratypes: MZB 2021-0789 / MZB 2021-0791 (3 ♀♀), type loc., 27-VIII-2020, A. Meseguer, L. Auroux, J. Comas &amp; J. Pallisé leg. │ MZB 2021-0853 / MZB 2021-0855 (1♂, 2 ♀♀), type loc., 27-VIII-2020, A. Meseguer, L. Auroux, J. Comas &amp; J. Pallisé leg. (ex coll. X. Bellés) │ MZB 2021-0792, MZB 2021-0793 (1 ♂, 1 ♀), Cova de l’Infern, 27-VIII-2020, J. Pallisé leg. │ MZB 2021-0856, MZB 2021-0857 (1 ♂, 1 ♀), Cova de l’Infern, 27-VIII-2020, J. Pallisé leg. (ex coll. X. Bellés) │ MZB 2021-0794 (1 ♀), Avenc Enfonsat A, 10-IV-1971, J. Comas leg. │ MZB 2021-0858 (1 ♀), Avenc Enfonsat A, 10-IV-1971, J. Comas leg. (ex coll. X. Bellés) .</p><p>Remarks</p><p>Type series consisting of 19 ♂♂ and 37 ♀♀ from four cavities in the province of Barcelona (Comas, 2021). Specimens of the type series deposited in other collections (Comas, 2021): ♀ holotype (JCC); paratypes in the FLC, JCC. The paratypes originally deposited in the XBC (Comas, 2021) have been donated to the MCNB collection.</p></div>	https://treatment.plazi.org/id/D42CA7775405CB4BFF6DFD907ED9FC0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775405CB4BFF6DF9117ED9F97C.text	D42CA7775405CB4BFF6DF9117ED9F97C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parvospeonomus garrigai Comas 2021	<div><p>Parvospeonomus garrigai Comas, 2021</p><p>Parvospeonomus garrigai Comas, 2021: 32; figs. 1c, 2c, 3c, 4c.</p><p>Current accepted name</p><p>Parvospeonomus garrigai Comas, 2021</p><p>Type locality</p><p>Cova dels Ermitans [original citation: “Cv. dels Ermitans, Caldes de Montbui (Vallès Oriental) … ETRS89 Fus: 31N / 428588, 4612604 / 520 m ”].</p><p>Type material</p><p>Paratypes: MZB 2021-0795 / MZB 2021-0798 (1 ♂, 3 ♀♀), type loc., 23-VII-2020, J. Pallisé leg. │ MZB 2021- 0849 / MZB 2021-0852 (1 ♂, 3 ♀♀), type loc., 23-VII-2020, J. Pallisé leg. (ex coll. X. Bellés).</p><p>Remarks</p><p>Type series consisting of 14 ♂♂ and 51 ♀♀ from four cavities in the province of Barcelona (Comas, 2021). Specimens of the type series deposited in other collections (Comas, 2021): ♀ holotype (JCC); paratypes in the FLC, JCC. The paratypes originally deposited in the XBC (Comas, 2021) have been donated to the MCNB collection.</p></div>	https://treatment.plazi.org/id/D42CA7775405CB4BFF6DF9117ED9F97C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775405CB4AFF6DFA03780FFCDB.text	D42CA7775405CB4AFF6DFA03780FFCDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Perriniella bofilli Zariquiey 1924	<div><p>Perriniella bofilli Zariquiey, 1924 (Fig. 40)</p><p>Perriniella Bofilli Zariquiey, 1924: 14; figs. 7, 8.</p><p>Current accepted name</p><p>Perriniella bofilli Zariquiey, 1924</p><p>Type locality</p><p>Cova de les Encantades [original citation: “«Cova de les Encantades», a dos horas de Queralps (Caralps), al N. de Ribas, valle del río Fresser, Prov. Gerona”].</p><p>Type material</p><p>Syntypes: MZB 78-2057 / MZB 78-2066, MZB 78-2688 / MZB 78-2694 (3 ♂♂, 17 ♀♀), type loc., X-1923 (trap), R. Zariquiey leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Zariquiey (1924) mentions as types a number of specimens collected in traps without giving the exact number or where they are deposited. In the MCNB collection we have found 20 specimens, all with the red labels “ Typus ” except for MZB 78-2066 that has a red label with the inscription “Co-Typus”. Most collection labels give the date October 1923 (in the case of three specimens, the exact date of 28 October 1923 is given). A part of the type series is deposited in the MNHNP collection (Jeannel, 1924a). We found four specimens in the MNHNP general collection (box 73) with similar labels to those of the material deposited in the MCNB bearing the printed inscription “ TYPUS ”. (Fresneda &amp; Salgado, 2016, transcribe the labels associated with these specimens). In the MNCNM collection (box 26), we have found a specimen labelled as a cotype (on two labels, one of which is red), and with a collection label that resembles those of the specimens deposited in the MCNB.</p></div>	https://treatment.plazi.org/id/D42CA7775405CB4AFF6DFA03780FFCDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775404CB4AFF6DFE6079DDF9E9.text	D42CA7775404CB4AFF6DFE6079DDF9E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Quaestus) acuminatus Salgado 1999	<div><p>Quaestus (Quaestus) acuminatus Salgado, 1999 (Fig. 41)</p><p>Quaestus (Quaestus) acuminatus Salgado, 1999a: 211; figs. 2–5.</p><p>Current accepted name</p><p>Quaestus acuminatus Salgado, 1999: Fresneda et al., 2024: 487.</p><p>Type locality</p><p>Cueva la Xiana [original citation: “ Cueva la Xiana ... in Tospe (Municipality of Arriondas, Asturias, Spain)”].</p><p>Type material</p><p>Paratypes: MZB 98-0901 / MZB 98-0907, MZB 2010-0178, MZB 2017-1385 / MZB 2017-1387 (4 ♂♂, 7 ♀♀), Cueva de Pico de Biesques, 9-IX-1995, J. M. Salgado &amp; D. Rodríguez leg.</p><p>Remarks</p><p>Type series consisting of 177 ♂♂ and 303 ♀♀ from eight caves in Asturias (Salgado, 1999a). Male holotype and female allotype deposited in the CZULE; paratypes deposited in other collections: ACC, CZULE, MNHNP and PMGC (Salgado, 1999a). Fresneda &amp; Salgado (2016) indicated that paratypes from the Cueva la Cueva and Cueva de los Padrones are deposited in the MCNB; nevertheless, we have only been able to find the specimens from the aforementioned Cueva de Pico de Biesques.</p></div>	https://treatment.plazi.org/id/D42CA7775404CB4AFF6DFE6079DDF9E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775404CB75FF6DFB7E7F4AF868.text	D42CA7775404CB75FF6DFB7E7F4AF868.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Quaestus) asturicus Fresneda & Salgado 2001	<div><p>Quaestus (Quaestus) asturicus Fresneda &amp; Salgado, 2001</p><p>Quaestus (Quaestus) asturicus Fresneda &amp; Salgado, 2001: 104; figs. 2–7.</p><p>Current accepted name</p><p>Quaestus asturicus Fresneda &amp; Salgado, 2001: Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva de Cuetu Lluengu [original citation: “cueva de Cuetu Lluengu ... Riensena, (Asturias)”].</p><p>Type material</p><p>Paratypes: MZB 2010-0172, MZB 2017-1262 / MZB 2017-1264 (2 ♂♂, 2 ♀♀), type loc., 26-IX-1998, J. M. Salgado &amp; D. Rodríguez leg.</p><p>Remarks</p><p>Type series consisting of 52 ♂♂ and 36 ♀♀ from two cavities in Asturias (Fresneda &amp; Salgado, 2001). Specimens from the type series deposited in other collections (Fresneda &amp; Salgado, 2001, 2016): ♂ holotype in CZULE; paratypes in the CZULE, FLC, MCNA, MNCNM, MNHNP, PMGC and RMC.</p></div>	https://treatment.plazi.org/id/D42CA7775404CB75FF6DFB7E7F4AF868	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543ACB74FF6DF9597FE2F976.text	D42CA777543ACB74FF6DF9597FE2F976.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Asturianella) incognitus Salgado & Fresneda 2004	<div><p>Quaestus (Asturianella) incognitus Salgado &amp; Fresneda, 2004b: 39; figs. 1–9.</p><p>Current accepted name</p><p>Quaestus incognitus Salgado &amp; Fresneda, 2004: Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva de Julió [original citation: “Cave of Julió, Caleao, T. M. de Caso (Asturias)”].</p><p>Type material</p><p>Paratypes: MZB 2010-0170, MZB 2017-1255 / MZB 2017-1258 (3 ♂♂, 2 ♀♀), Cueva de Rearco, 15-IV-2003, J. M. Salgado &amp; D. Rodríguez leg. │ MZB 2021-0815, MZB 2021-0816 (1 ♂, 1 ♀), Cueva de Rearco, 22-IV-1989, J. M. Salgado &amp; D. Rodríguez leg. (ex coll. X. Bellés).</p><p>Remarks</p><p>Type series consisting of 260 ♂♂ and 347 ♀♀ from four cavities in Asturias (Salgado &amp; Fresneda, 2004b). Specimens of the type series deposited in other collections: ♂ holotype in the MHNG; paratypes in the CZULE, FLC, FMNHC, MHNG, MNHNP, PMGC and XBC (Salgado &amp; Fresneda, 2004b; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA777543ACB74FF6DF9597FE2F976	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543ACB74FF6DFD907BA6FC0A.text	D42CA777543ACB74FF6DFD907BA6FC0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Quaesticulus) bustilloi Salgado & Fresneda 2009	<div><p>Quaestus (Quaesticulus) bustilloi Salgado &amp; Fresneda, 2009</p><p>Quaestus (Quaesticulus) bustilloi Salgado &amp; Fresneda, 2009: 3; figs. 1, 2a–d, 3a,c,d.</p><p>Current accepted name</p><p>Schaufussiella bustilloi (Salgado &amp; Fresneda, 2009): Fresneda et al., 2025: 199.</p><p>Type locality</p><p>Cueva de la Canal [original citation: “ España, Asturias, Villaorille-Quirós, Cueva de la Canal, UTM: 29T QH 421869, 740 m.”].</p><p>Type material</p><p>Paratypes: MZB 2010-0177, MZB 2017-1271 / MZB 2017-1274 (4 ♂♂, 1 ♀), Cueva de la Peña de Cotrutes, 8-V-2004, J. M. Salgado &amp; D. Rodríguez leg.</p><p>Remarks</p><p>Male holotype deposited in the CZULE (Biodiversity and Environmnetal Management Department, University of León) (Sagado &amp; Fresneda, 2009). A total of 132 male paratypes and 118 female paratypes (designated as allotypes) deposited in the MCNB and the following collections: AFC, CBC, CZULE, FLC, MCNA, MNCNM, MNHNP, MRSNT and PMGC (Sagado &amp; Fresneda, 2009; Fresneda &amp; Salgado, 2016). The specimens from the type series are from two caves in Asturias. Fresneda &amp; Salgado (2016) indicate that paratypes from the type locality are deposited in the MCNB but we have only been able to find the specimens from the aforementioned Cueva de la Peña de Cotrutes.</p></div>	https://treatment.plazi.org/id/D42CA777543ACB74FF6DFD907BA6FC0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543ACB77FF6DFA0D7831FDF3.text	D42CA777543ACB77FF6DFA0D7831FDF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Speogeus) jubilationis Salgado & Fresneda 2010	<div><p>Quaestus (Speogeus) jubilationis Salgado &amp; Fresneda, 2010</p><p>Quaestus (Speogeus) jubilationis Salgado &amp; Fresneda, 2010: 100; figs. 1, 2a,c,e,g,i.</p><p>Current accepted name</p><p>Quaestus jubilationis Salgado &amp; Fresneda, 2010: Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva de Covallos [original citation:“ España, Asturias,Pando-Villaviciosa, Cueva de Covallos, UTM 30TUP072138, 235 m s.n.m.”].</p><p>Type material</p><p>Paratypes: MZB 2013-1840, MZB 2017-0308 / MZB 2017-0310 (2 ♂♂, 2 ♀♀), type loc., 10/ 17-X-2009 (trap), J. M. Salgado &amp; D. Rodríguez leg.</p><p>Remarks</p><p>Type series consisting of 25 ♂♂ and 41 ♀♀ (Salgado &amp; Fresneda, 2010). Specimens of the type series deposited in other collections: ♂ holotype in the CZULE; paratypes in the CZULE, FLC, MCNA, MNCNM, MNHNP and PMGC (Salgado &amp; Fresneda, 2010; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA777543ACB77FF6DFA0D7831FDF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775439CB77FF6DFF887BB8FAC9.text	D42CA7775439CB77FF6DFF887BB8FAC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Speogeus) littoralis Salgado 1999	<div><p>Quaestus (Speogeus) littoralis Salgado, 1999</p><p>Quaestus (Speogeus) littoralis Salgado, 1999b: 82; figs. 1–6.</p><p>Current accepted name</p><p>Quaestus littoralis Salgado, 1999: Fresneda et al., 2024: 489.</p><p>Type locality</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-5.1308336&amp;materialsCitation.latitude=43.466667" title="Search Plazi for locations around (long -5.1308336/lat 43.466667)">Cueva del Barredo</a> [original citation: “ Grotte du Barredo (43º28'N – 5º7'51"W) ... La Torre (Asturies, Espagne)”] .</p><p>Type material</p><p>Paratypes: MZB 2010-0171, MZB 2017-1259 / MZB 2017-1261 (2 ♂♂, 2 ♀♀), type loc., 4-VI-1994, J. M. Salgado leg.</p><p>Remarks</p><p>Male holotype and female allotype are deposited in the MHNG. Paratypes (43 ♂♂ and 45 ♀♀) are deposited in the MCNB and in the CZULE, FLC, MCNA, MHNG, MNHNP and PMGC (Salgado, 1999b; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775439CB77FF6DFF887BB8FAC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775439CB77FF6DF85E7FE2F838.text	D42CA7775439CB77FF6DF85E7FE2F838.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Speogeus) mermejaensis Salgado & Fresneda 2004	<div><p>Quaestus (Speogeus) mermejaensis Salgado &amp; Fresneda, 2004</p><p>Quaestus (Speogeus) mermejaensis Salgado &amp; Fresneda, 2004b: 41; figs. 18–23.</p><p>Current accepted name</p><p>Quaestus mermejaensis Salgado &amp; Fresneda, 2004: Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva del Prau [original citation: “Cave of the Prau, Puente de Piedra-Buspriz, T. M. de Caso (Asturies, Spain)”].</p><p>Type material</p><p>Paratypes: MZB 2010-0174, MZB 2017-1268 / MZB 2017-1270 (2 ♂♂, 2 ♀♀), type loc., 31-VII-2002, J. M. Salgado &amp; D. Rodríguez leg. │ MZB 2021-0813, MZB 2021-0814 (1 ♂, 1 ♀), type loc., 17-VI-1989, J. M. Salgado &amp; D. Rodríguez leg. (ex coll. X. Bellés).</p><p>Remarks</p><p>Type series consisting of 54 ♂♂ and 75 ♀♀ from three cavities in Asturias (Salgado &amp; Fresneda, 2004b). Specimens of the type series deposited in other collections: ♂ holotype in the MHNG; paratypes in CZULE, FLC, FMNHC, MHNG, MNHNP, PMGC and XBC (Salgado &amp; Fresneda, 2004b; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775439CB77FF6DF85E7FE2F838	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775438CB71FF6DFB417AC9FDF3.text	D42CA7775438CB71FF6DFB417AC9FDF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Quaesticulus) pasensis Salgado, Labrada & Luque 2010	<div><p>Quaestus (Quaesticulus) pasensis Salgado, Labrada &amp; Luque, 2010</p><p>Quaestus (Quaesticulus) pasensis Salgado, Labrada &amp; Luque, 2010: 28; figs. 1–7.</p><p>Current accepted name</p><p>Luquesia pasensis (Salgado, Labrada &amp; Luque, 2010): Fresneda et al., 2024: 518.</p><p>Type locality</p><p>Cueva de El Churrón [original citation: “ SPAIN: Cantabria: Luena: El Churrón Cave ( The Churrón-Millajo Cave karst system, 30TVN 2988980475, 255 m)”] .</p><p>Type material</p><p>Paratypes: MZB 2010-0176, MZB 2017-1275 / MZB 2017-1277 (2 ♂♂, 2 ♀♀), Cueva del Rellano, 23-VIII-2008, C. G. Luque &amp; L. Labrada leg.</p><p>Remarks</p><p>Salgado et al. (2010) refer to six paratypes (3 ♂♂ and 3 ♀♀) deposited in the MCNB collection with the same collection dates as the holotype. Nevertheless, in the MCNB collection there are only four paratypes (2 ♂♂ and 2 ♀♀) from Cueva del Rellano, with the date 23 August 2008 on the original label (on which Salgado and González are referred to mistakenly as the collectors). Male holotype and female allotype deposited in the CZULE (Salgado et al., 2010). The authors of the description refer to numerous paratypes (79 ♂♂ and 87 ♀♀) from five caves in Cantabria deposited in the MCNB and other collections: CZULE, FLC, MCNA, MNCNM, MNHNP and PMGC (Salgado et al., 2010) .</p></div>	https://treatment.plazi.org/id/D42CA7775438CB71FF6DFB417AC9FDF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543FCB71FF6DFF887F1FFB2D.text	D42CA777543FCB71FF6DFF887F1FFB2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Quaestus) sajaensis Salgado 1998	<div><p>Quaestus (Quaestus) sajaensis Salgado, 1998 (Fig. 43)</p><p>Quaestus (Quaestus) sajaensis Salgado, 1998: 258; figs. 2–5.</p><p>Current accepted name</p><p>Quaestus sajaensis Salgado, 1998: Fresneda et al., 2024: 491.</p><p>Type locality</p><p>Cueva El Poyo [original citation: “ Cueva El Poyo ... Saja (Cantabria)”].</p><p>Type material</p><p>Paratypes: MZB 98-0896 / MZB 98-0900 (2 ♂♂, 3 ♀♀), type loc., 26-V-1995, J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of 21 ♂♂ and 50 ♀♀ from two localities in Cantabria (Salgado 1998). Specimens of the type series deposited in other collections: ♂ holotype and ♀ allotype in the CZULE; paratypes in CZULE, FLC, FMNHC, MCNA, MNHNP and PMGC (Salgado 1998; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA777543FCB71FF6DFF887F1FFB2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543FCB71FF6DF83279A7F860.text	D42CA777543FCB71FF6DF83279A7F860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quaestus (Quaestus) spinulosus Salgado 1998	<div><p>Quaestus (Quaestus) spinulosus Salgado, 1998</p><p>Quaestus (Quaestus) spinulosus Salgado, 1998: 261; figs. 6–10.</p><p>Current accepted name</p><p>Quaestus spinulosus Salgado, 1998: Fresneda et al., 2024: 491.</p><p>Type locality</p><p>Cueva les Xianes [original citation: “Cueva les Xianes, La Piñera-Sevares, Asturias (España)”].</p><p>Type material</p><p>Paratypes: MZB 98-0892 / MZB 98-0895, MZB 2010-0175, MZB 2017-1388 / MZB 2017-1390 (4 ♂♂, 4 ♀♀), type loc., 12-X-1996, J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of 18 ♂♂ and 30 ♀♀ (Salgado 1998). Specimens of the type series deposited in other collections: ♂ holotype and ♀ allotype in the CZULE, paratypes in the CZULE, FLC, FMNHC, MCNA, MNHNP and PMGC (Salgado 1998; Fresneda &amp; Salgado, 2016.</p></div>	https://treatment.plazi.org/id/D42CA777543FCB71FF6DF83279A7F860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543ECB73FF6DFB6D7A0BFE1F.text	D42CA777543ECB73FF6DFB6D7A0BFE1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spelaeochlamys verai Comas 1977	<div><p>Spelaeochlamys verai Comas, 1977 (Fig. 44)</p><p>Spelaeochlamys verai Comas, 1977b: 217; figs. 1–5.</p><p>Current accepted name</p><p>Spelaeochlamys ehlersi verai Comas, 1977: Salgado &amp; Fresneda, 2004a: 381.</p><p>Type locality</p><p>Cova del Tío Melchor [original citation: “Cova del Tío Melchor, Castalla, Alicante”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1909, type loc., 15-IV-1976, J. M. Vera leg.</p><p>Paratypes: MZB 78-1910 (1 ♀), same data as holotype. │ MZB 78-1911 / MZB 78-1913 (6 ♂♂, 15 ♀♀, 1 indet.), type loc., 30-V-1976, O. Escolà &amp; J. Comas leg.</p><p>Remarks</p><p>The type series consists of 15 ♂♂ and 33 ♀♀ (Comas, 1977b). On the labels of the paratypes with register numbers MZB 78-1911 / MZB 78-1913 the collection date appears as 31 May 1976. Paratypes are also deposited in the JCC (Comas, 1977b).</p></div>	https://treatment.plazi.org/id/D42CA777543ECB73FF6DFB6D7A0BFE1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543DCB73FF6DFF25788CFB2D.text	D42CA777543DCB73FF6DFF25788CFB2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharidius galani Espanol 1970	<div><p>Speocharidius galani Español, 1970 (Fig. 45)</p><p>Speocharidius galani Español, 1970: 59; figs. 1, 2.</p><p>Current accepted name</p><p>Speocharidius (Kobiella) galani Español, 1970: Fresneda &amp; Salgado, 2000: 49.</p><p>Type locality</p><p>Guardetxe Aurreko Leizea 1 [original citation: “Guardetxe-Aurre’ko Leizea 1, entre Usurbil y Orio, en la orilla izquierda del Oria, Guipúzcoa ”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2151, type loc., 9-VII-1968, C. Galán leg.</p><p>Paratypes: MZB 78-2152 / MZB 78-2155 (11 ♀♀), MZB 71-2603 (remains 1 ♂), same data as holotype .</p><p>Remarks</p><p>Type series consisting of the ♂ holotype and 18 paratypes (4 ♂♂ and 14 ♀♀) deposited in the MCNB and SCA collections (Español, 1970). Genitalia preserved in Canada balsam and mounted on the same pin as the holotype. Remains of a male specimen preserved in the same way with its genitalia. This material (MZB 71-2603) has a label with the following inscription: [ Speocharidius / sp. n. ♂ / Guardetxe leg.] .</p></div>	https://treatment.plazi.org/id/D42CA777543DCB73FF6DFF25788CFB2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543DCB73FF6DF83278F3F860.text	D42CA777543DCB73FF6DF83278F3F860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharidius vivesi Espanol & Belles 1980	<div><p>Speocharidius vivesi Español &amp; Bellés, 1980 (Fig. 46)</p><p>Speocharidius vivesi Español &amp; Bellés, 1980: 33; figs. 2, 3, 6.</p><p>Current accepted name</p><p>Speocharidius (Speocharidius) vivesi Español &amp; Bellés, 1980: Fresneda &amp; Salgado, 2000: 49.</p><p>Type locality</p><p>Ekainko Leizea [original citation: “Ekain’ko Leizea, Cestona ( Guipúzcoa)”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2145, type loc., 26-III-1978, E. Vives leg.</p><p>Paratypes: MZB 78-2146 / MZB 78-2150 (2 ♂♂, 3 ♀♀), same data as holotype .</p><p>Remarks</p><p>On the labels of specimens with the accession numbers MZB 78-2148, MZB 78-2149 and MZB 78-2150 the collection date is given as 23 March 1978. Although Español &amp; Bellés (1980) refer to the deposit of paratypes in the EVC, the six specimens of the type series are deposited in the MCNB .</p></div>	https://treatment.plazi.org/id/D42CA777543DCB73FF6DF83278F3F860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543CCB72FF6DFE3A78E7FA15.text	D42CA777543CCB72FF6DFE3A78E7FA15.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis (Speogeus) avicularis Salgado 1985	<div><p>Speocharis (Speogeus) avicularis Salgado, 1985 (Fig. 47)</p><p>Speocharis (Speogeus) avicularis Salgado, 1985: 263; figs. 1–6.</p><p>Current accepted name</p><p>Quaestus avicularis (Salgado, 1985): Fresneda et al., 2024: 489.</p><p>Type material</p><p>Cueva de los Pandanes [original citation: “Cueva de los Pandanes ... en Soto de Agues (Asturias)”].</p><p>Types</p><p>Paratypes: MZB 78-2159, MZB 78-2160 (3 ♂♂, 3 ♀♀), Cueva del Troncu, 27-IX-1984, J. M. Salgado leg.</p><p>Remarks</p><p>On the labels (hand-written by O. Escolà) the collection date is given as 24 September 1984. Type series consists of 69 ♂♂ and 46 ♀♀ from three caves in Asturias (Salgado, 1985). Fresneda &amp; Salgado (2016) refer to the deposit in the MCNB collection of paratypes from these three localities, although in the Museum there are only the six aforementioned specimens from Cueva del Troncu. Types deposited in other collections (Salgado, 1985; Fresneda &amp; Salgado, 2016): ♂ holotype, ♀ allotype and paratypes in the CZULE.</p></div>	https://treatment.plazi.org/id/D42CA777543CCB72FF6DFE3A78E7FA15	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543CCB72FF6DFD9079ABFD22.text	D42CA777543CCB72FF6DFD9079ABFD22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis amicalis Salgado 1984	<div><p>Speocharis amicalis Salgado, 1984</p><p>Speocharis amicalis Salgado, 1984: 262; figs. 10, 16, 18.</p><p>Current accepted name</p><p>Quaestus amicalis (Salgado, 1984): Fresneda et al., 2024: 487.</p><p>Type locality</p><p>Cueva Subterráneo [original citation: “ Cueva subterráneo, Sellaño, provincia de Oviedo”].</p><p>Type material</p><p>Paratypes: MZB 78-2174, MZB 78-2175 (4 ♂♂, 4 ♀♀), type loc., 16-IX-1983, J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of holotype ♂ and 344 paratypes (156 ♂♂ and 188 ♀♀) from two cavities in Asturias (Salgado, 1984). Specimens of the type series in other collections: holotype in the CZULE; paratypes in the CZULE and FLC (Salgado, 1984; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA777543CCB72FF6DFD9079ABFD22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777543CCB7DFF6DFB2B79E3FE3B.text	D42CA777543CCB7DFF6DFB2B79E3FE3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis begoniae Negre 1965	<div><p>Speocharis begoniae Nègre, 1965 (Figs. 8, 48)</p><p>Speocharis begoniae Nègre, 1965: 93; figs. 1, 4.</p><p>Current accepted name</p><p>Amphogeus angustitarsis (Español, 1950): Fresneda et al., 2024: 507.</p><p>Type locality</p><p>Cueva de Urrekoba [original citation: “ Cueva de Urrekoba situada en el desfiladero de Atxarte, en la falda del Monte Unzillaitz, Ayuntamiento de Abadiano, cerca de Durango”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2219, type loc., 31-III-1948, J. Nègre, L. &amp; M. B. Hérault leg.</p><p>Paratypes: MZB 78-1920, MZB 78-2220, MZB 78-2221 (3 ♂♂, 1 ♀), type loc., VIII-1963, Grupo EspeleolÓgico Vizcaíno leg.</p><p>Remarks</p><p>Nègre (1965) does not state exactly the number of paratypes (“una pequeña serie de ejemplares recogidos en la misma cueva, en agosto 1963 ” [a short series of specimens collected in one cave in August 1963]) nor their deposit details. We have located the holotype in the MCNB collection (first deposited in the author’s collection) as well as four paratypes. It is likely that this material came to the MCNB collection via the donation of the author’s private collection. Hand-written on the capture labels the author is given as the collector, who refers to the traps placed on his behalf by the Vizcaíno Speleological Society .</p></div>	https://treatment.plazi.org/id/D42CA777543CCB7DFF6DFB2B79E3FE3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775433CB7DFF6DFF417AE4FB55.text	D42CA7775433CB7DFF6DFF417AE4FB55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis (Speocharis) bergidi Salgado 1983	<div><p>Speocharis (Speocharis) bergidi Salgado, 1983</p><p>Speocharis (Speocharis) bergidi Salgado, 1983: 277; figs. 1–5.</p><p>Current accepted name</p><p>Leonesiella bergidi (Salgado, 1983): Salgado, 1996: 153.</p><p>Type locality</p><p>Cueva de la Gruta [original citation: “ Cueva de La Gruta, La Barosa, provincia de León”].</p><p>Type material</p><p>Paratypes: MZB 78-2208 / MZB 78-2210 (2 ♂♂, 2 ♀♀), type loc., 28-III-1982, J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of 9 ♂♂ and 10 ♀♀ (Salgado 1983). Types deposited in other collections (Salgado, 1983; Fresneda &amp; Salgado, 2016): ♂ holotype in CZULE; paratypes in the CZULE and MNCNM (ex coll. Instituto Español de Entomología).</p></div>	https://treatment.plazi.org/id/D42CA7775433CB7DFF6DFF417AE4FB55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775433CB7DFF6DF9EA7E44F838.text	D42CA7775433CB7DFF6DF9EA7E44F838.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis cantabricus subsp. angustitarsis Espanol 1950	<div><p>Speocharis cantabricus angustitarsis Español, 1950</p><p>Speocharis cantabricus angustitarsis Español, 1950a: 107 .</p><p>Current accepted name</p><p>Amphogeus angustitarsis (Español, 1950): Fresneda et al., 2024: 507.</p><p>Type locality</p><p>Cueva de San Lorenzo n. 1; Cueva de Azko [original citation: “ Vizcaya: cueva de San Lorenzo en Mañaria ... cueva de Azco, Mañaria ...”].</p><p>Type material</p><p>Syntypes: MZB 78-1916, MZB 78-1917 (2 ♂♂, 3 ♀♀), Cueva de San Lorenzo, IX-1935, F. Español &amp; R. Zariquiey leg. │ MZB 78-1918, MZB 78-1919 (1 ♂, 2 ♀♀), Cueva de Azko, IX-1935, F. Español &amp; R. Zariquiey leg.</p><p>Remarks</p><p>Español (1950a) neither establishes a holotype nor states how many examples belong to the type series, and merely indicates two collection sites and the collection date. Eight specimens have been deposited in the MCNB collection (five from Cueva de San Lorenzo and the other three from Cueva de Azko). Each specimen has red labels hand-written by F. Español with the inscription “Tipo” for MZB 78-1916 ( San Lorenzo) and “ Paratipo ” for the others. The first of these specimens was designated incorrectly by Escolà (2000) as the holotype (Article 73.1.3.) on a hand-written red label. We have been unable to find any material from the type series in any other collection.</p></div>	https://treatment.plazi.org/id/D42CA7775433CB7DFF6DF9EA7E44F838	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775432CB7FFF6DFBC37833FE47.text	D42CA7775432CB7FFF6DFBC37833FE47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis espanoli Salgado 1978	<div><p>Speocharis espanoli Salgado, 1978</p><p>Speocharis espanoli Salgado, 1978: 15; fig. 2.</p><p>Current accepted name</p><p>Quaestus espanoli (Salgado, 1978): Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva La Cuevona [original citation: “Prov. de Oviedo: ... Cueva La Cuevona, cerca de Ribadesella].” Type material</p><p>Holotype ♂: MZB 78-2161, type loc., H. Obermaier leg.</p><p>Paratypes: MZB 78-2162 / MZB 78-2164 (1 ♂, 2 ♀♀), same data as holotype .</p><p>Remarks</p><p>Type series consisting of ♂ holotype and four paratypes (1 ♂ and 3 ♀♀) deposited in the MCNB collection (Salgado, 1978). Three of the four mentioned paratypes have been located .</p></div>	https://treatment.plazi.org/id/D42CA7775432CB7FFF6DFBC37833FE47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775431CB7FFF6DF95A7E5CF8D0.text	D42CA7775431CB7FFF6DF95A7E5CF8D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis (Speocharinus) llosesi Espanol & Escola 1977	<div><p>Speocharis (Speocharinus) llosesi Español &amp; Escolà, 1977</p><p>Speocharis (Speocharinus) llosesi Español &amp; Escolà, 1977b: 26; figs. 1–4.</p><p>Current accepted name</p><p>Speocharinus llosesi (Español &amp; Escolà, 1977b): Bellés et al., 1978: 61</p><p>Type locality</p><p>Morterón n. 1 del Hoyo Salzoso [original citation: “Morterón del Hoyo Salzoso ... a 870 m de altitud junto a los Trillos”].</p><p>Type material</p><p>Syntypes: MZB 78-0968 / MZB 78-0972 (4 ♂♂, 1 ♀), type loc., 9-IV-1977, R. Lloses leg.</p><p>Remarks</p><p>Español &amp; Escolà (1977b) do not establish a holotype. The ♂ specimen MZB 78-0968 has a red label with the printed inscription “Typus” and corresponds to the designation of the holotype made incorrectly by Escolà (2000) (Article 73.1.3.). Label 3 transcripted by Escolà for this specimen is missing. (in place of this label there is the above-mentioned red label). The remaining specimens have red printed labels with the inscription ‘Paratypus’, as mentioned by Escolà (2000). Type series wholly deposited in the MCNB (Español &amp; Escolà, 1977b).</p></div>	https://treatment.plazi.org/id/D42CA7775431CB7FFF6DF95A7E5CF8D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775431CB7FFF6DFCDD7F04FBC2.text	D42CA7775431CB7FFF6DFCDD7F04FBC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis jeannei Coiffait 1965	<div><p>Speocharis jeannei Coiffait, 1965 (Fig. 49)</p><p>Speocharis jeannei Coiffait, 1965: 292; figs. 18–21.</p><p>Current accepted name</p><p>Quaestus jeannei (Coiffait, 1965): Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva de la Vega de Teón [original citation: “ Cueva de la Vega de Teón, 960 m, Covadonga (Oviedo)”].</p><p>Type material</p><p>Paratypes: MZB 78-2186, MZB 78-2187 (3 ♂♂), type loc., Jeanne leg.</p><p>Remarks</p><p>Type series consisting of holotype and 11 paratypes (6 ♂♂ and 5 ♀♀) (Coiffait, 1965). The holotype is deposited in the HCC; the remaining paratypes are deposited in the CJC and HCC (Coiffait, 1965).</p></div>	https://treatment.plazi.org/id/D42CA7775431CB7FFF6DFCDD7F04FBC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775431CB7EFF6DFA68793BFDAF.text	D42CA7775431CB7EFF6DFA68793BFDAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis longicornis Salgado 1989	<div><p>Speocharis longicornis Salgado, 1989</p><p>Speocharis longicornis Salgado, 1989: 127; figs. 4–6.</p><p>Current accepted name</p><p>Quaestus longicornis (Salgado, 1989): Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva de los Pandanes [original citation: “Cueva de los Pandanes ... en Soto de Agues (Asturias)”].</p><p>Type material</p><p>Paratypes: MZB 78-2211, MZB 78-2212 (5 ♂♂, 5 ♀♀), type loc., 27-IX-1984, J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of 292 ♂♂ and 327 ♀♀ from three cavities in Asturias (Salgado, 1989). Types deposited in other collections (Salgado, 1989; Fresneda &amp; Salgado, 2016): ♂ holotype and ♀ allotype in the CZULE; paratypes in BMNHL, CZULE, FLC and MNHNP.</p></div>	https://treatment.plazi.org/id/D42CA7775431CB7EFF6DFA68793BFDAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775430CB7EFF6DFFB57FEFFB2D.text	D42CA7775430CB7EFF6DFFB57FEFFB2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis luctuosus Salgado 1984	<div><p>Speocharis luctuosus Salgado, 1984</p><p>Speocharis luctuosus Salgado, 1984: 260; figs. 9, 14, 15, 17.</p><p>Current accepted name</p><p>Quaestus luctuosus (Salgado, 1984): Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva del Agua [original citation: “Cueva del Agua o del Venero, a 1,5 km de las Cuevas, provincia de Oviedo”].</p><p>Type material</p><p>Paratypes: MZB 78-2172, MZB 78-2173 (2 ♂♂, 2 ♀♀), type loc., 23-IX-1981, J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of ♂ holotype and 23 paratypes (11 ♂♂ and 12 ♀♀) (Salgado, 1984). The holotype and the remaining paratypes are deposited in the CZULE (Salgado, 1984; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775430CB7EFF6DFFB57FEFFB2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775430CB7EFF6DF8327E5DF844.text	D42CA7775430CB7EFF6DF8327E5DF844.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis mariscali Salgado 1979	<div><p>Speocharis mariscali Salgado, 1979</p><p>Speocharis mariscali Salgado, 1979: 387; fig. 1.</p><p>Current accepted name</p><p>Schaufussiella mariscali (Salgado, 1979): Fresneda et al., 2025: 200.</p><p>Type locality</p><p>Cueva de los Tocinos [original citation: “Cueva de los Tocinos, al sur de la vertiente Cantábrica, a unos 800 m. de Caldas de Luna, provincia de León”].</p><p>Type material</p><p>Paratypes: MZB 78-2215, MZB 78-2216 (2 ♂♂, 2 ♀♀), type loc., 28-V-1977, J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of 8 ♂♂ and 6 ♀♀ (Salgado, 1979). Types deposited in other collections (Salgado, 1979): ♂ holotype in CZULE; paratypes in CZULE and MNCNM (ex coll. Instituto Español de Entomología).</p></div>	https://treatment.plazi.org/id/D42CA7775430CB7EFF6DF8327E5DF844	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775437CB78FF6DFBC97E3CFDF3.text	D42CA7775437CB78FF6DFBC97E3CFDF3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis mierensis Bolivar 1911	<div><p>Speocharis mierensis Bolívar, 1911 (Fig. 50)</p><p>Speocharis Mierensis Bolívar, 1911: 570; figs. p. 571.</p><p>Current accepted name</p><p>Hypogesia minos (Jeannel, 1909): Fresneda et al., 2024: 517.</p><p>Type locality</p><p>Cueva de Cuerdavilloso [original citation: “ Cueva de Cuerdavilloso, Rucandio (Santander) ”].</p><p>Type material</p><p>Syntypes: MZB 78-2481 (1 ♀), type loc., C. Bolívar leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Bolívar (1911) neither establishes the holotype, the number of specimens in the type series, nor where it is deposited. The specimen in the MCNB collection is identified as a cotype on the determination label hand-written by C. Bolívar and on another red label. Deposited in the MNCNM (box 4) there is a series of specimens headed by one identified by Bolívar as the “TIPO”. All of these specimens have subsequently been labelled as syntypes on red labels. In box 54 of the MNHNP there are at least four specimens labelled as cotypes (material examined by J. M. Salgado) .</p></div>	https://treatment.plazi.org/id/D42CA7775437CB78FF6DFBC97E3CFDF3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775436CB78FF6DFF897868F95D.text	D42CA7775436CB78FF6DFF897868F95D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis nadali Salgado 1978	<div><p>Speocharis nadali Salgado, 1978 (Figs. 9A–B, 51)</p><p>Speocharis nadali Salgado, 1978: 16; fig. 3.</p><p>Current accepted name</p><p>Cantabrogeus nadali (Salgado, 1978): Salgado et al., 2008: 222.</p><p>Type locality</p><p>Cueva de la Cañuela [original citation: “Prov. de Santander: ... Cueva de la Cañuela, cerca de Arredondo, al descender el Puerto de Alisas, en la carretera de la Santander a Ramales”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2192, type loc., 3/ 5-VIII-1954, Nègre, Dresco &amp; Derouet leg.</p><p>Paratypes and possible paratypes: MZB 78-2193 / MZB 78-2198 (5 ♂♂, 7 ♀♀), same data as holotype .</p><p>Remarks</p><p>According to Salgado (1978) the type series consists of a holotype ♂ and eight paratypes (2 ♂♂ and 6 ♀♀) collected on 3 and 5 August 1954 (Nègre, Dresco &amp; Derouet leg.) and deposited in the MCNB. The material we have found with labels stating that it belongs to the type series corresponds to 13 specimens (6 ♂♂ and 7 ♀♀). Specimen MZB 78-2192 has a red label with the printed inscription “Tipus” and its scientific name hand-written by O. Escolà; the remaining specimens have different kinds of red labels with the type-written inscription “ Paratipo ”.All originate from the type locality according to the capture labels hand-written by the different authors (among them J. M. Salgado). The labels of 3 ♂♂ (including the specimen labelled as “Typus”) and 3 ♀♀ give a collection date of 4 August 1954 (Nègre leg.). All the males except one have labels that indicate that their genitalia were studied or that they were prepared for microscopic study, although none of the genitalia are mounted with their corresponding specimens and none have been found. The characteristics and aspect of most of the the cards on which the specimens are mounted are similar and it is possible that they have been removed from their original pins, re-labelled and mounted on other pins to facilitate their study. Despite the lack of any pertinent documentation or indication, it is possibly that all these specimens were used in the description of this new species .</p></div>	https://treatment.plazi.org/id/D42CA7775436CB78FF6DFF897868F95D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775436CB7BFF6DFBE27AB9FC07.text	D42CA7775436CB7BFF6DFBE27AB9FC07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis (Speocharis) nietoi Salgado 1988	<div><p>Speocharis (Speocharis) nietoi Salgado, 1988 (Fig. 10)</p><p>Speocharis (Speocharis) nietoi Salgado, 1988: 61; figs. 2–7.</p><p>Current accepted name</p><p>Quaestus nietoi (Salgado, 1988): Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva del Sierru [original citation: “Cueva del Sierru, en San Pedro de Tolivia (Asturias)”].</p><p>Type material</p><p>Possible paratypes: MZB 78-2213, MZB 78-2214 (5 ♂♂, 5 ♀♀), type loc., 13-IV-1985?, J. M. Salgado leg.</p><p>Remarks</p><p>When citing the information regarding the type series, Salgado (1988: 61) does not give a data for the paratypes. However, subsequently (p. 65) he does assign a date of 13 April 1985, given as the capture date of the holotype and allotype, to a series of 166 ♂♂ and 141 ♀♀ specimens (which match the number of specimens in the type series). The labels of the material deposited in the MCNB (5 ♂♂ and 5 ♀♀) have the date 9 March 1986, which corresponds to a large group of specimens from the same locality, also mentioned by the author (p. 65) but excluded from the type series (Article 72.4.6.). These specimens have red labels bearing the inscription “Paratipus / Speocharis nietoi n. sp. / J. Mª Salgado det.”. The labels (including the capture labels) were hand-written by O. Escolà. Salgado (1988) indicates that the holotype ♂ and allotype ♀ are deposited in his own collection, and that the paratypes are in his own collection and in the collections of the CZULE and MNHNP. We believe that once the description had been published, the author’s idea was to donate paratypes to the MCNB, However, either the MCNB specimens are true paratypes and the date on the capture label is wrong (i.e. it should be 13 April 1985), or they correspond, as the date indicates, to the large series captured on 9 March 1986 that was erroneously labelled as a paratype. The precision of the published information and the information on the labels admits no other explanation bearing in mind, as well, that Salgado (1988) cites over 3000 specimens from the type locality. Although it is likely that these specimens are really paratypes (indeed, Fresneda &amp; Salgado, 2016, mention the deposit of paratypes in the MCNB), we cannot be absolutely sure that this is so.</p></div>	https://treatment.plazi.org/id/D42CA7775436CB7BFF6DFBE27AB9FC07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775435CB7BFF6DF91D7A77F95D.text	D42CA7775435CB7BFF6DF91D7A77F95D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis nuptialis Espanol 1973	<div><p>Speocharis nuptialis Español, 1973</p><p>Speocharis nuptialis Español, 1973: 88; figs. 1–4.</p><p>Current accepted name</p><p>Quaestus nuptialis (Español, 1973): Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva del Triángulo [original citation: “Cueva del Triángulo o de Calderón, Velilla de Carrión, prov. de Palencia”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2190, type loc., 15-VIII-1972, L. Auroux leg.</p><p>Paratypes: MZB 78-2191 (1 ♂ “adelfotípico”), same data as holotype .</p><p>Remarks</p><p>Type series consisting of the two cited specimens (Español, 1973); on their labels the collection date is given as 12 August 1972.</p></div>	https://treatment.plazi.org/id/D42CA7775435CB7BFF6DF91D7A77F95D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775435CB7AFF6DFBE27A5AFC4F.text	D42CA7775435CB7AFF6DFBE27A5AFC4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis obermaieri Bolivar 1923	<div><p>Speocharis obermaieri Bolívar, 1923 (Figs. 11A–B)</p><p>Speocharis obermaieri Bolívar, 1923: 427 .</p><p>Current accepted name</p><p>Quaestus occidentalis (Jeannel, 1911): Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva de la Verde [original citation: “ Cueva de la Verde, en Garandiella, Porrúa, término municipal y partido de Llanes, provincia de Oviedo”].</p><p>Type material</p><p>Paratypes: MZB 71-8181 / MZB 71-8184 (1 ♂, 3 ♀♀), MZB 78-2475, MZB 78-2476 (2 ♀♀), type loc., H. Obermaier leg. (ex coll. R. Zariquiey). │ MZB 71-3746 / MZB 71-3750 (3 ♂♂, 2 ♀♀), type loc., H. Obermaier leg. (ex coll. J. Nègre) .</p><p>Remarks</p><p>Bolívar (1923) does not specify the number of specimens in the type series. The identification labels of specimens MZB 71-3746 and MZB 78-2475 (hand-written by Bolívar) state “Cotipos”. Along with these specimens we have located in their respective original places of deposit the remaining specimens with the same or very similar printed labels, although most of them are not labelled as types. In Nègre’s collection we have also found a male from the same locality (MZB 71-3751) whose labels, unlike the above-mentioned specimens, are hand-written and provide no information regarding the collector. One of these labels includes the inscription “Mat. Bolivar / Mus. Madrid ”. The ♂ holotype, originally designated as “Tipo” (Article 73.1.1.), is deposited in the MNCNM (box 9), along with numerous specimens of the same origin that probably belong to the type series, although they are not identified as such. Jeannel (1924a) reports the deposit of type specimens in the Biospeologica collection. In the MNHNP collection (box 53) at least two specimens identified as cotypes have been found (material examined by J. M. Salgado).</p></div>	https://treatment.plazi.org/id/D42CA7775435CB7AFF6DFBE27A5AFC4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775434CB7AFF6DFED57F1AF8A5.text	D42CA7775434CB7AFF6DFED57F1AF8A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis olajensis Salgado 1978	<div><p>Speocharis olajensis Salgado, 1978 (Figs. 12, 52)</p><p>Speocharis olajensis Salgado, 1978: 12; fig. 1.</p><p>Current accepted name</p><p>Quaestus olajensis (Salgado, 1978): Fresneda et al., 2024: 489.</p><p>Type locality</p><p>Cueva del Carrascal [original citation: “Provincia de León: ... Cueva del Carrascal, a unos 90 minutos del pueblo de Sta. Olaja de la Varga”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2199, type loc., 22-IX-1974, J. M. Salgado leg.</p><p>Paratypes: MZB 78-2201, MZB 78-2202 (3 ♂♂, 3 ♀♀), same data as holotype. │ MZB 78-2203, MZB 78-2204 (1 ♂, 3 ♀♀), Cueva de Covarrubias, 22-IX-1974, J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of 27 ♂♂ and 34 ♀♀ from caves in the province of LeÓn (Salgado, 1978). The collection labels of the specimens in the MCNB from Cueva del Carrascal and Cueva de Covarrubias (hand-written by Salgado) give, respectively, the dates 20 September 1974 and 23 September 1974. Along with this material there is also a pin, with accession number MZB 78-2200, with two male genitalia preserved in Canadian balsam and mounted on acetate labels. Each one has a hand-written label stating “C. del Carrascal / Sta. Olaja de la Varga / León — nº 2” and “C. del Carrascal / Sta. Olaja de la Varga / León — nº 3”. Given how they have been prepared (very similar to the preparation of the holotype), it seems likely that these genitalia were taken from two paratypes or, at least, were used as study material. Nevertheless, we cannot be sure which specimens they correspond to. The remaining paratypes are deposited in the CZULE collection (Salgado, 1978; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775434CB7AFF6DFED57F1AF8A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777542ACB64FF6DFD90791DFC6E.text	D42CA777542ACB64FF6DFD90791DFC6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis oxypterus Bolivar 1917	<div><p>Speocharis oxypterus Bolívar, 1917</p><p>Speocharis oxypterus Bolívar, 1917: 207; figs. 1, 2.</p><p>Current accepted name</p><p>Amphogeus oxypterus (Bolívar, 1917): Fresneda et al., 2024: 508.</p><p>Type locality</p><p>Cueva de Juan Gómez [original citation: “Provincia de Santander: cueva de Juan Gómez, en Sámano, a unos 5 km. de Castro-Urdiales”].</p><p>Type material</p><p>Paratypes: MZB 78-2482 (1 ♀), type loc., IX-1916, C. Bolívar leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Bolívar (1917) does not specify how many specimens there are in the type series. The specimen deposited in the MNCB, damaged, is identified as a cotype, both on the identification label hand-written by Bolívar and on another red label. The ♂ holotype, originally designated as “Tipo” (Article 73.1.1.), is deposited in the MNCNM (box 4). Together with this specimen, we have also found at least eight specimens that, given the associated information and the format of their capture labels, probably belong to the type series, although they do not bear labels that state this. We have located in the MNHNP at least six specimens (box 54) labelled as cotypes with similar capture labels (material examined by J. M. Salgado) .</p></div>	https://treatment.plazi.org/id/D42CA777542ACB64FF6DFD90791DFC6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777542ACB64FF6DFEF6791DF860.text	D42CA777542ACB64FF6DFEF6791DF860.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis pachecoi Bolivar 1915	<div><p>Speocharis pachecoi Bolívar, 1915 (Figs. 13A–B)</p><p>Speocharis Pachecoi Bolívar, 1915: 401 .</p><p>Current accepted name</p><p>Schaufussiella pachecoi (Bolívar, 1915): Fresneda et al., 2025: 200.</p><p>Type locality</p><p>Cueva de la Peña [original citation: “Provincia de Oviedo: cueva de la Peña ... en San Román de Candamo, Ayuntamiento de Pravia, valle del río Nalón”].</p><p>Type material</p><p>Syntypes: MZB 71-8204 (1 ♂), MZB 78-2476 / MZB 78-2480 (2 ♂♂, 5 ♀♀), type loc., 20-VIII-1915, H. Pacheco &amp; J. Cabré leg. (ex coll. R. Zariquiey). │ MZB 71-3756 (1 ♂), type loc., 20-VIII-1915, H. Pacheco &amp; J. Cabré leg. (ex coll. J. Nègre).</p><p>Remarks</p><p>Bolívar (1915) neither establishes a holotype nor indicates where the studied material is deposited (“numerosos ejemplares” [many specimens]). The specimens MZB 78-2476, MZB 78-2477 and 78-2478 have red printed cotype labels; furthermore, specimen MZB 78-2477 has a determination label hand-written by Bolívar with the inscription “co-TIPOS”. Specimens MZB 71-3756 and MZB 71-8204 are also identified as cotypes (on white labels). Four other specimens, with accession numbers MZB 78-2479 and MZB 78-2480, have labels of origin (H. Pacheco leg.) and, although not labelled as cotypes, they must in fact belong to the type series. In the MNCNM collection (box 12) we have found many specimens that probably represent most of the type series, given the information on the printed collection labels and their characteristics (none of them have labels that state that they are type specimens). In the MNHNP general collection (box 53) we have found at least two specimens with the printed inscription “COTYPE” (material examined by J. M. Salgado) .</p></div>	https://treatment.plazi.org/id/D42CA777542ACB64FF6DFEF6791DF860	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775429CB67FF6DFD90781FFD22.text	D42CA7775429CB67FF6DFD90781FFD22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis pseudoccidentalis Salgado 1980	<div><p>Speocharis pseudoccidentalis Salgado, 1980</p><p>Speocharis pseudoccidentalis Salgado, 1980b: 269; figs. 1–8.</p><p>Current accepted name</p><p>Quaestus pseudoccidentalis (Salgado, 1980): Fresneda et al., 2024: 491.</p><p>Type locality</p><p>Cueva del Sidrón [original citation: “Cueva del Sidrón, Barines-Villamayor, provincia de Oviedo”].</p><p>Type material</p><p>Paratypes: MZB 78-2169 / MZB 78-2171 (10 ♂♂, 5 ♀♀), type loc., 31-VIII-1979, E. Samartino leg.</p><p>Remarks</p><p>Type series consisting of 43 ♂♂ and 35 ♀♀ (Salgado, 1980). Types deposited in other collections (Salgado, 1980; Fresneda &amp; Salgado, 2016): ♂ holotype in the CZULE; paratypes in the CZULE, FLC, MNCNM (ex coll. Instituto Español de Entomología), and in the E. Samartino collection .</p></div>	https://treatment.plazi.org/id/D42CA7775429CB67FF6DFD90781FFD22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775429CB67FF6DFE3A7E4BFA5C.text	D42CA7775429CB67FF6DFE3A7E4BFA5C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis recordationis Salgado 1982	<div><p>Speocharis recordationis Salgado, 1982</p><p>Speocharis recordationis Salgado, 1982: 54; figs. 3d, 7, 8.</p><p>Current accepted name</p><p>Quaestus recordationis (Salgado, 1982): Fresneda et al., 2024: 491.</p><p>Type locality</p><p>Cueva del Agua [original citation: “Cueva del Agua o del Venero, a 1 Km. de Las Cuevas, provincia de Oviedo”].</p><p>Type material</p><p>Paratypes: MZB 78-2188, MZB 78-2189 (3 ♂♂, 3 ♀♀), type loc., 23-IX-1981, J. M. Salgado leg.</p><p>Remarks</p><p>Type series consisting of 27 ♂♂ and 32 ♀♀ (Salgado, 1982). The author cites 4 ♂♂ and 4 ♀♀ deposited in the MCNB collection. Types deposited in other collections (Salgado, 1982; Fresneda &amp; Salgado, 2016): ♂ holotype in the CZULE; paratypes in the CZULE, FLC and MNCNM (ex coll. Instituto Español de Entomología).</p></div>	https://treatment.plazi.org/id/D42CA7775429CB67FF6DFE3A7E4BFA5C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775429CB66FF6DF8E37870FDD7.text	D42CA7775429CB66FF6DF8E37870FDD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis sellai Bolivar 1923	<div><p>Speocharis sellai Bolívar, 1923</p><p>Speocharis sellai Bolívar, 1923: 426 .</p><p>Current accepted name</p><p>Quaestus sellai (Bolívar, 1923): Fresneda et al., 2024: 491.</p><p>Type locality</p><p>Cueva del Penicial [original citation: “cueva del Penicial, en Nueva ... término municipal y partido de Llanes, provincia de Oviedo”].</p><p>Type material</p><p>Paratypes: MZB 78-2165 / MZB 78-2168, MZB 78-2473, MZB 78-2474 (2 ♂♂, 4 ♀♀), type loc., Conde de la Vega de Sella &amp; Obermaier leg. (ex coll. R. Zariquiey) . │ MZB 71-3763 / MZB 71-3771 (2 ♂♂, 7 ♀♀), type loc., Conde de la Vega de Sella &amp; Obermaier leg. (ex coll. J. Nègre) .</p><p>Remarks</p><p>Male holotype originally designated as “Tipo” (Article 73.1.1.). Bolívar (1923) does not specify how many specimens there are in the type series, which originates from four caves in Asturias. The specimens originally deposited in Zariquiey’s collection have red labels with the printed inscription “Cotypus”. Specimen MZB 78-2473 also has a determination label hand-written by Bolívar with the inscription “cotipos”. In Nègre’s collection we have found nine specimens with identical printed collection labels, of which the first (MZB 71-3763) also has the same label as mentioned above, hand-written by Bolívar. The holotype is deposited in the MNCNM (Bolívar, 1923). Together with this specimen, in the same collection (box 9) we have also found a number of specimens that seem to belong to the type series given the characteristics of their labels and the information they contain; however, they bear no labels that identify them as types. In the MNHNPs general collection (box 53) with have located at least one specimen labelled as a cotype (material examined by J. M. Salgado).</p></div>	https://treatment.plazi.org/id/D42CA7775429CB66FF6DF8E37870FDD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775428CB66FF6DFF6D7E11FAC6.text	D42CA7775428CB66FF6DFF6D7E11FAC6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis suevensis Salgado 1991	<div><p>Speocharis suevensis Salgado, 1991</p><p>Speocharis suevensis Salgado, 1991: 122; figs. 1–5, 15.</p><p>Current accepted name</p><p>Quaestus suevensis (Salgado, 1991): Fresneda et al., 2024: 491.</p><p>Type locality</p><p>Cueva de la Carretera [original citation: “ Cueva de la Carretera o Pitufos, altitud 110 m ... Bodes (Asturias)”].</p><p>Type material</p><p>Paratypes: MZB 78-2177 / MZB 78-2185 (4 ♂♂, 5 ♀♀), type loc., 13-V-1989, J. M. Salgado leg.</p><p>Remarks</p><p>On the labels of the paratypes in the MCNB the collection date is given as 13 May 1979, probably due to an error when copying the year. The type series consists of the ♂ holotype and the ♀ allotype, as well as 220 paratypes (106 ♂♂ and 114 ♀♀) (Salgado, 1991). The holotype and allotype are deposited in the author’s collection (CZULE); the remaining paratypes are deposited in the author’s collection (CZULE) and in the FLC and MNHNP (Salgado, 1991; Fresneda &amp; Salgado, 2016). Of the material deposited in the MCNB, Salgado (1991) cites 3 ♂♂ and 3 ♀♀.</p></div>	https://treatment.plazi.org/id/D42CA7775428CB66FF6DFF6D7E11FAC6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775428CB66FF6DF84C7FEAF837.text	D42CA7775428CB66FF6DF84C7FEAF837.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speocharis zariquieyi Jeannel 1924	<div><p>Speocharis zariquieyi Jeannel, 1924</p><p>Speocharis Zariquieyi Jeannel, 1924a: 59 .</p><p>Current accepted name</p><p>Uhagoniella zariquieyi (Jeannel, 1924): Fresneda et al., 2024: 166.</p><p>Type locality</p><p>Puerto de Leitariegos (south slope) [original citation: “León: Caboalles, au pied du versant sud du pic de Leitariegos”].</p><p>Type material</p><p>Syntypes: MZB 78-2472 (1 ♂, 3 ♀♀), type loc., Paganetti &amp; Hümmler leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Type series consisting of a large number of specimens (with no more information) deposited originally in the Biospeologica collection (Jeannel, 1924a). The specimens in the MCNB, mounted on the same pin, have a label with the inscription “ TYPE ” printed in red letters. In the MNHNP general collection (box 52) we have located part of the type series, examined by J. Mª Salgado: two specimens and a genitalia with red labels with the inscription “ TYPE ” printed in black, and eight specimens with the same label “ TYPE ” as MZB 78-2472 .</p></div>	https://treatment.plazi.org/id/D42CA7775428CB66FF6DF84C7FEAF837	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777542ECB60FF6DFD907F55FC96.text	D42CA777542ECB60FF6DFD907F55FC96.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomites velox Jeannel 1910	<div><p>Speonomites velox Jeannel, 1910 (Fig. 53)</p><p>Speonomites velox Jeannel, 1910a: CLI; figs. 1, 2.</p><p>Current accepted name</p><p>Speonomites velox Jeannel, 1910</p><p>Type locality</p><p>Cova Fosca (de Vilanova) [original citation: “... dans une petite grotte, située sur le revers méridional de la sierra de Montsech, dans le bassin du rio Segre, nommée «Cova Fosca», près de Villanova di Meya (partido de Balaguer, provincia de Lerida)”].</p><p>Type material</p><p>Syntypes: MZB 78-2646 (1 ♂), type loc., 8-IX-1910, R. Jeannel &amp; E. G. Racovitza leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Type series consisting of 13 specimens originally deposited in the Biospeologica collection (Jeannel, 1910a, 1924a). The specimen in the MCNB has a label with the inscription “ TYPE ” printed in red letters and hand-written collection labels. In the MNHNP’s general collection (box 72) we have found at least one specimen with identical collection labels and with a red label with the inscription “ TYPE ” printed in black letters.</p></div>	https://treatment.plazi.org/id/D42CA777542ECB60FF6DFD907F55FC96	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777542ECB63FF6DFEAD7FDAFE1F.text	D42CA777542ECB63FF6DFEAD7FDAFE1F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus akarsticus Escola 1980	<div><p>Speonomus akarsticus Escolà, 1980</p><p>Speonomus akarsticus Escolà, 1980a: 27; figs. 1–3.</p><p>Current accepted name</p><p>Stygiophyes akarsticus (Escolà, 1980): Fresneda, 1998: 61.</p><p>Type locality</p><p>Mines dels Cubilars [original citation: “Mines dels Cubilars (Torre de Cabdella, prov. Lleida. Margen izquierda del Flamicell)”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1111, type loc., 12-IV-1970, Auroux, Ubach &amp; Escolà leg.</p><p>Paratypes: MZB 78-1112 / MZB 78-1118 (4 ♂♂, 4 ♀♀), MZB 78-1119 (remains 1 ex.), same data as holotype. │MZB 78-1121 (1 ♀), Avenc del Solanet, 27-XII-1975, Ubach, Nubiola &amp; Escolà leg. │ MZB 69-0161 / MZB 69-0166 (281 ♂♂, 502 ♀♀), MZB 72-4075 (2 ♀♀), MZB 75-9329 (1 ♂), MZB 78-1122 / MZB 78-1169 (87 ♂♂, 23 ♀♀), 78-3076 (2 ♂♂, 1 ♀) Avenc del Solanet, 27-XII-1975 / 24-X-1976 (trap), Rosaura, Clariana, Centelles &amp; Escolà leg. │ MZB 78-1170 / MZB 78-1172 (3 ♂♂, 5 remains), Avenc del Tinc-ca-tinc, 12-X-1978, Escolà leg. │ MZB 78-1173 / MZB 78-1178 (5 ♂♂, 16 ♀♀), Avenc del Tinc-ca-tinc, 17-XII-1978, Bellés, Cuñé &amp; Escolà leg.</p><p>Remarks</p><p>The type series consists of 992 specimens from three localities in the Lleida Pyrenees, most (887) of which were trapped in Avenc del Solanet (Escolà, 1980a, 1981). We have located 939 specimens in the MCNB collection (783 of which preserved in alcohol). Some of these trapped specimens have deteriorated (many are disarticulated): the remains of a specimen with accession number MZB 78-1119 from Mines dels Cubilars, also labelled as a paratype, should be added to the nine specimens cited from this locality by Escolà (1980a) (this specimen is included on the record card of the author’s data file). Specimen MZB 78-1120 (not labelled as a paratype) corresponds to the genitalia, supposedly extracted from one of the males from Mines dels Cubilars. The collection label of male MZB 78-1122 gives a date of 27 December 1975, whilst for specimens MZB 78-1123 and MZB 78-1124 (1 ♂ and 1 ♀) the date cited is 24 October 1976 (these two dates correspond, respectively, to the date the traps were placed in Avenc del Solanet and when they were removed). Escolà (1980a) only mentions a female collected on 27 December 1975 (which must correspond to MZB 78-1121) and so, given that these specimens are labelled as paratypes, we must suppose that they were either trapped or were not mentioned by the author in his original description. It is reasonable to think that these discrepancies are due to the difficulty in keeping a track of and labelling so many specimens, and so we believe that the cited specimens should be included in the type series. We have located another 127 specimens trapped in Avenc del Solanet (register number MZB 69-0167), labelled by Escolà as “en mal estado” [‘in poor condition’]. This material, preserved in alcohol, were probably excluded from the type series due to their bad state of preservation.</p><p>Some paratypes have been deposited in the XBC, according to Fresneda &amp; Salgado (2016). Likewise, according with our data, two paratypes were sent to Michael Perreau and other two to Dante Vailati .</p></div>	https://treatment.plazi.org/id/D42CA777542ECB63FF6DFEAD7FDAFE1F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777542DCB63FF6DFF2478C8FA81.text	D42CA777542DCB63FF6DFF2478C8FA81.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus aldomai Fresneda & Hernando 1988	<div><p>Speonomus aldomai Fresneda &amp; Hernando, 1988 (Fig. 54)</p><p>Speonomus aldomai Fresneda &amp; Hernando, 1988: 164; figs. 3–5, 7, 9–10, 12, 15, 17–20.</p><p>Current accepted name</p><p>Stygiophyes aldomai aldomai (Fresneda &amp; Hernando, 1988): Fresneda, 1998: 61.</p><p>Type locality</p><p>Graller de les Planelles [original citation: “Graller de les Planelles (Corroncui, El Pont de Suert, Lleida)”].</p><p>Type material</p><p>Holotype ♂: MZB 99-1341, type loc., 31-V-1985, J. Fresneda &amp; J. Aldomà leg.</p><p>Paratypes: MZB 99-0557 / MZB 99-0560 (35 ♂♂, 31 ♀♀), type loc., 27-VII-1986 / 27-V-1989 (trap), O. Escolà leg.</p><p>Remarks</p><p>Type series consisting of 64 ♂♂ and 75 ♀♀ (Fresneda &amp; Hernando, 1988; Fresneda &amp; Salgado, 2016). According to Fresneda &amp; Escolà (2000), the type series also has a further 14 specimens and 15 remains, also identified as paratypes on the record card of Escolà’s data file.Although this material should be deposited in the MCNB collection (Fresneda &amp; Escolà, 2000; Fresneda &amp; Salgado, 2016), its location currently remains unknown. The MCNB collection has 36 ♂♂ (including a holotype) and 31 ♀♀; however, Fresneda &amp; Hernando (1988) report the deposit in this collection of 38 ♂♂ (including a holotype) and 33 ♀♀. Paratypes deposited in other collections: CHC, FLC, MHNT, PMGC and SZC (Fresneda &amp; Hernando, 1988; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA777542DCB63FF6DFF2478C8FA81	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777542DCB6DFF6DF8A67F6FFF37.text	D42CA777542DCB6DFF6DF8A67F6FFF37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus andorranus Comas 1978	<div><p>Speonomus andorranus Comas, 1978</p><p>Speonomus andorranus Comas, 1978a: 73; figs. 1–3.</p><p>Current accepted name</p><p>Speonomites andorranus (Comas, 1978): Fresneda, 1998: 67.</p><p>Type locality</p><p>Cova de la Gorga de la Margineda [original citation: “Cova de la Gorga de la Margineda, Sant Julià, Andorra ”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1079, type loc., 2-XI-1974, J. Comas leg.</p><p>Paratypes: MZB 78-1080 / MZB 78-1082 (1 ♂, 2 ♀♀), same data as holotype. │ MZB 2021-0834, MZB 2021-0836 (1 ♂, 1 ♀), same data as holotype (ex coll. X. Bellés) . │ MZB 2021-0835, MZB 2021-0837 (5 ♀♀), Cova de la Margineda, 17-XII-1974, J. Comas leg. (ex coll. X. Bellés) .</p><p>Remarks</p><p>Type series consisting of 53 ♂♂ and 66 ♀♀ from two caves in the Andorran Pyrenees (Comas, 1978a).The specimens deposited in the MCNB are labelled with the collection date 2 October 1974 rather than 2 November 1974, according to Comas (1978a). The author of the description refers to paratypes originally deposited in the MCNB collected on 17 December 1974 from both caves, and to paratypes from the type locality collected on 7 October 1978, although we have been unable to locate such material. The rest of the paratypes deposited in the JCC; the paratypes originally deposited in the XBC (Comas, 1978a) have been donated to the MCNB collection.</p></div>	https://treatment.plazi.org/id/D42CA777542DCB6DFF6DF8A67F6FFF37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775423CB6DFF6DF93E7E7FF81C.text	D42CA7775423CB6DFF6DF93E7E7FF81C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Speonomus) aurouxi Espanol 1966	<div><p>Speonomus (Speonomus) aurouxi Español, 1966 (Fig. 56)</p><p>Speonomus (s. str.) aurouxi Español, 1966a: 2; plate 1, figs. 1–3.</p><p>Current accepted name</p><p>Speonomites aurouxi (Español, 1966): Fresneda, 1998: 67.</p><p>Type locality</p><p>Escletxa Minguera; Escletxa de la Vinya d’en Jaufré; Cova de Sant Gervàs [original citation: “Lérida: Escletxa Minguera, Sant Miquel de la Vall … escletxa de la Vinya d’en Jaufré, Sant Miquel de la Vall … cova de Sant Gervás, Sant Miquel de la Vall”].</p><p>Type material</p><p>Syntypes: MZB 78-1054 / MZB 78-1059 (5 ♂♂, 7 ♀♀), Escletxa Minguera, 8-XII-1964, Auroux &amp; Amat leg. │ MZB 78-1060 / MZB 78-1063 (5 ♂♂, 5 ♀♀), Escletxa Minguera, 26-XII-1964, González, Auroux, Amat &amp; Español leg. │ MZB 78-1069 (1 ♀), Escletxa de la Vinya d’en Jaufré, 27-XII-1964, Auroux &amp; Amat leg. │ MZB 78-1064 / MZB 78-1068, MZB 75-9330, 75-9331 (7 ♂♂, 9 ♀♀), Cova de Sant Gervàs, 27-XII-1964, González, Auroux, Amat &amp; Español leg.</p><p>Remarks</p><p>Español (1966a) neither establishes a holotype in his original publication nor states how many specimens there are in the type series; we are unaware of any other specimens in any other collection. The specimen MZB 78-1054 has a red label with the inscription “Typus” hand-written by F. Español; the remaining specimens have red labels with the inscription “ Paratypus ” either hand-written or printed. The collection date on the labels of the specimens from Escletxa de la Vinya d’en Jaufré and Cova de Sant Gervàs is 26 December 1964 , whilst on the labels of the specimens from Escletxa Minguera (González, Auroux, Amat &amp; Español leg.) the date is given 27 December 1964.</p></div>	https://treatment.plazi.org/id/D42CA7775423CB6DFF6DF93E7E7FF81C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775423CB6DFF6DFC4C7916FC29.text	D42CA7775423CB6DFF6DFC4C7916FC29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus antemi Escola 1972	<div><p>Speonomus antemi Escolà, 1972</p><p>Speonomus antemi Escolà, 1972: 42; fig. 1.</p><p>Current accepted name</p><p>Speonomites antemi (Escolà, 1972): Fresneda, 1998: 67.</p><p>Type locality</p><p>Cova de Porredons [original citation: “Bahent. Cova dels Porredons ... situada en la vertiente oriental del río Noguera Pallaresa, en el paralelo de Gerri de la Sal”].</p><p>Type material</p><p>Syntypes: 78-1104 / MZB 78-1110 (2 ♂♂, 5 ♀♀), type loc., 1-III-1970, J. M. Victoria leg.</p><p>Remarks</p><p>Escolà (1972) does not specify the number of specimens in the type series, which, according to the introduction to the description, are deposited in the MCNB. Neither does he establish the holotype, although specimen MZB 78-1104 has a red label printed with the inscription “Type” (the remaining specimens have red labels with the inscription “ Paratype ” or “ Paratypus ”).</p></div>	https://treatment.plazi.org/id/D42CA7775423CB6DFF6DFC4C7916FC29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775422CB6FFF6DFBAE7A55FDAF.text	D42CA7775422CB6FFF6DFBAE7A55FDAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus bolivari subsp. altimontanus Belles 1975	<div><p>Speonomus bolivari altimontanus Bellés, 1975</p><p>Speonomus bolivari altimontanus Bellés, 1975: 49; figs. 1, 3.</p><p>Current accepted name</p><p>Trapezodirus altimontanus (Bellés, 1975): Fresneda, 1998: 59.</p><p>Type locality</p><p>Cueva de Garcés [original citation: “Cova de Garsés, a la Vall d’Ordesa, sobre la línia divisòria dels termes municipals de Torla i Fanlo, província d’Osca”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1179, type loc., 14-VIII-1972, X. Bellés leg.</p><p>Paratypes: MZB 78-1180 / MZB 78-1182 (1 ♂, 2 ♀♀), type loc., 14-VIII-1972, T. Cuñe, J. Serrano &amp; X. Bellés leg. │ MZB 2021-0838, MZB 2021-0839, MZB 2021-0841 (2 ♂♂, 3 ♀♀), type loc., 14-VIII-1972, T. Cuñe, J. Serrano &amp; X. Bellés leg. (ex coll. X. Bellés). │ MZB 2021-0840 (1 ♀), type loc., 16-VII-1972, J. Serrano &amp; X. Bellés leg. (ex coll. X. Bellés).</p><p>Remarks</p><p>Type series consisting of the ♂ holotype (referred to as “Typus”, Article 73.1.1.) and 11 paratypes (4 ♂♂ and 7 ♀♀) without details of where they are deposited (Bellés, 1975). On the hand-written labels of the paratypes originally deposited in the MCNB collection date is given as 14 July 1972 rather than the published date of 14 August 1972. The paratypes originally deposited in the XBC (Fresneda &amp; Salgado, 2016) have been donated to the MCNB collection.</p></div>	https://treatment.plazi.org/id/D42CA7775422CB6FFF6DFBAE7A55FDAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775421CB6FFF6DFFB47876FA72.text	D42CA7775421CB6FFF6DFFB47876FA72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus bolivari subsp. luciani Jeannel 1911	<div><p>Speonomus bolivari luciani Jeannel, 1911</p><p>Speonomus Bolivari subsp. Luciani Jeannel, 1911a: XCII; figs. 12, 13.</p><p>Current accepted name</p><p>Trapezodirus bolivari (Martínez de la Escalera, 1898): Fresneda, 1998: 59.</p><p>Type locality</p><p>Forau de la Drolica [original citation: “grotte dite « Forau de la Drolica », près de Sarsa-de-Surta, partido de Boltaña, provincia de Huesca (Aragon)”] .</p><p>Type material</p><p>Syntypes: 78-2628 (1 ♂, 1 ♀), type loc., 26-VI-1911, R. Jeannel &amp; E. G. Racovitza leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Type series consisting of around 40 specimens (“40 exemplaires environ”) originally deposited in the Biospeologica collection (Jeannel, 1911a, 1924a). The specimens in the MCNB (mounted on the same pin) have a label with the inscription “ TYPE ” printed in red and a hand-written collection label. The MNHNP general collection (box 70) contains the two specimens from the type series (1♂ and 1♀) whose labels are transcribed by Fresneda &amp; Salgado (2016). In the same part of this collection we have also located another four 4 ♂♂ from the type series with white labels and the inscription “ TYPE ” printed in red (material examined by J. Fresneda). We have located in the MNCNM collection (box 21) a male identified as a cotype.</p></div>	https://treatment.plazi.org/id/D42CA7775421CB6FFF6DFFB47876FA72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775421CB6EFF6DFB0A7B87FE63.text	D42CA7775421CB6EFF6DFB0A7B87FE63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Euryspeonomus) breuili subsp. kilixketai Espanol 1945	<div><p>Speonomus (Euryspeonomus) breuili kilixketai Español, 1945</p><p>Speonomus (Euryspeonomus) breuili ssp. kilixketai Español, 1945: 269; pl. XII, fig. 1b.</p><p>Current accepted name</p><p>Euryspeonomus (Euryspeonomus) breuili (Jeannel, 1919): Bellés et al., 1978: 65.</p><p>Type locality</p><p>Cueva de Kilixketa [original citation: “cueva de Kilixketa (Aralar)”].</p><p>Type material</p><p>Paratypes: MZB 78-1183 (1 ♀), type loc., campaign VIII-1945, J. M. Thomas, R. Margalef, J. Mateu &amp; F. Español leg.</p><p>Remarks</p><p>Español (1945) established as a “Tipo” (= holotype, Article 73.1.1.) a ♂ but neither establishes how many specimens there are in the type series nor where it is deposited. Nevertheless, with regard the description of the male antennae, the author mentions “los tres ejemplares examinados de ese sexo” [the three specimens of this sex examined], the remaining specimens being females. The holotype, identified by a red label, is deposited in the MNCNM collection, together with seven other specimens from the type locality. All but one (with date 18 August 1945) have a collection date of 21 August 1945 (the same as the paratype in the MCNB) but none are identified as belonging to the type series .</p></div>	https://treatment.plazi.org/id/D42CA7775421CB6EFF6DFB0A7B87FE63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775420CB6EFF6DFCF87F76FB55.text	D42CA7775420CB6EFF6DFCF87F76FB55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus canyellesi Lagar 1974	<div><p>Speonomus canyellesi Lagar, 1974</p><p>Speonomus canyellesi Lagar, 1974: 28; fig. 2.</p><p>Current accepted name</p><p>Parvospeonomus canyellesi (Lagar, 1974): Fresneda, 1998: 74.</p><p>Type locality</p><p>Avenc de les Pedreres [original citation: “avenc de Les Pedreres Gualba-Vallès Oriental”].</p><p>Type material</p><p>Paratypes: MZB 78-1013 (1 ♀), type loc., 17-II-1973, J. Cañelles &amp; A. Lagar leg.</p><p>Remarks</p><p>Lagar (1974) does not indicate the number of paratypes deposited originally in the author’s collection. On the collection label of the specimen deposited in the MCNB, the date 17 February 1971 is given by error. The holotype and paratypes were collected by Cañelles and Lagar on 4 and 17 February 1973 (Lagar, 1974), and a total of 32 specimens are deposited in the FLC (Fresneda &amp; Salgado, 2016). These authors cite as a paratype a specimen dated 11 March 1973 (Sabater, Cañelles &amp; Lagar leg.), deposited in the same collection.</p></div>	https://treatment.plazi.org/id/D42CA7775420CB6EFF6DFCF87F76FB55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775420CB6EFF6DF9EA7A74F81C.text	D42CA7775420CB6EFF6DF9EA7A74F81C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus cerberus subsp. arcticollis Jeannel 1911	<div><p>Speonomus cerberus arcticollis Jeannel, 1911 (Fig. 14)</p><p>Speonomus Cerberus subsp. arcticollis Jeannel, 1911a: XC; figs. 9, 10.</p><p>Current accepted name</p><p>Trapezodirus arcticollis (Jeannel, 1911): Fresneda, 1998: 59.</p><p>Type locality</p><p>Cova de Sant Salvador [original citation: “cueva de San Salvador, située près du pueblo de Bibils, districto mun. de Bonansa, partido de Benabarre, provincia de Huesca (Aragon)”] .</p><p>Type material</p><p>Syntypes: 78-2627 (1 ♂, 1 ♀), type loc., 20-VI-1911, R. Jeannel &amp; E. G. Racovitza leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>The type series consists of around 20 specimens (“20 exemplaires environ”) originally deposited in the Biospeologica collection (Jeannel, 1911a). The specimens in the MCNB collection (mounted on the same pin) have a label with the inscription “ TYPE ” printed in red letters and hand-written collection labels. These labels are of the same type as those transcribed by Fresneda &amp; Salgado (2016) for various specimens from the type series currently placed in box 70 in the MNHNP general collection. In the MNCNM collection (box 21) a female identified as a cotype has been located.</p></div>	https://treatment.plazi.org/id/D42CA7775420CB6EFF6DF9EA7A74F81C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775426CB68FF6DF9597AD5F88C.text	D42CA7775426CB68FF6DF9597AD5F88C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Speonomidius) ciaurrizi subsp. igaratzai Espanol 1945	<div><p>Speonomus (Speonomidius) ciaurrizi igaratzai Español, 1945</p><p>Speonomus (Speonomidius) ciaurrizi ssp. igaratzai Español, 1945: 270; pl. XII, fig. 2.</p><p>Current accepted name</p><p>Euryspeonomus (Urbasolus) ciaurrizi igaratzai (Español, 1945): Perreau, 2000: 304.</p><p>Type locality</p><p>Cueva de Basolo [original citation: “cueva de Basolo (Aralar)”].</p><p>Type material</p><p>Paratypes: MZB 78-1037, MZB 78-1038 (1 ♂, 1 ♀), type loc., campaign VIII-1945, J. M. Thomas, R. Margalef, J. Mateu &amp; F. Español leg.</p><p>Remarks</p><p>Español (1945) establishes as a “Tipo” (= holotype, article 73.1.1.) a male and notes “varios specimens” [various specimens] from the type locality without indicating where they are deposited. The two specimens in the MCNB collection have red labels with the inscription “ Paratypus ” hand-written by Español. The holotype (identified by a red label) is deposited in the MNCNM, together with three specimens from Cueva de Basolo, two more from Sima de Leizeta (Español, 1945), all with a collection date of 18 August 1945, and probably two other paratypes that are not identified as such.</p></div>	https://treatment.plazi.org/id/D42CA7775426CB68FF6DF9597AD5F88C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775426CB68FF6DFD907998FBC2.text	D42CA7775426CB68FF6DFD907998FBC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Speonomus) ciaurrizi Bolivar 1921	<div><p>Speonomus (Speonomus) ciaurrizi Bolívar, 1921 (Fig. 55)</p><p>Speonomus (s. str.) ciaurrizi Bolívar, 1921: 530; figs. 21–24.</p><p>Current accepted name</p><p>Euryspeonomus (Urbasolus) ciaurrizi ciaurrizi (Bolívar, 1921): Perreau, 2000: 304.</p><p>Type locality</p><p>Cueva de Malkorraundi [original citation: “cueva de Malkorraundi, situada cerca de Gorriti (término municipal de Larraun, partido de Pamplona, provincia de Navarra)”] .</p><p>Type material</p><p>Paratypes: MZB 78-1481, MZB 78-2623 / MZB 78-2625 (3 ♂♂, 1 ♀), type loc., 23-VIII-1919, C. Bolívar &amp; R. Jeannel leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Bolívar (1921) does not specify the number of specimens in the type series (“numerosos ejemplares” [numerous specimens]) and merely designates a single “Tipo” ♂ (= holotype, article 73.1.1.). The specimens deposited in the MCNB collection have red labels with the hand-written inscription “Cotypus”, the exception being MZB 78-1481 (with genitalia mounted on the same pin) that has a label hand-written by O. Escolà placed at a subsequent date with the inscription “paratipus”. The specimen MZB 78-2623 also has a determination label hand-written by Bolívar with the inscription “cotipus”. The ♂ holotype is deposited in the MNCNM (Bolívar, 1923), in box 19, together with a long series of specimens of the same origin and with identical collection labels, probably also belonging to the type series (but not identified as such). According to Jeannel (1924a), there is also material corresponding to the type series in the Biospeologica collection (MNHNP) .</p></div>	https://treatment.plazi.org/id/D42CA7775426CB68FF6DFD907998FBC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775426CB6BFF6DFA937FA2FC4F.text	D42CA7775426CB6BFF6DFA937FA2FC4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Speonomites) colominasi Zariquiey 1924	<div><p>Speonomus (Speonomites) colominasi Zariquiey, 1924 (Fig. 57)</p><p>Speonomus (Speonomites) Colominasi Zariquiey, 1924: 6; figs. 1, 2.</p><p>Current accepted name</p><p>Lagariella colominasi (Zariquiey, 1924): Fresneda, 2000: 147.</p><p>Type locality</p><p>Cova de Joan d’Os [original citation: “Cova de Joan de l’Os, llamada también Cova d’Ullet, abierta a poniente en la Serra de Mora y a unos 6 u 8 kms. de Tartareu; entre Tragó de Noguera y Avellanes, partido de Balaguer, provincia de Lérida”].</p><p>Type material</p><p>Syntypes: MZB 70-1578 / MZB 70-1583 (29 ♀♀), MZB 78-2632 / MZB 78-2645 (35 ♂♂, 32 ♀♀), type loc., R. Zariquiey leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Type series consisting of numerous specimens but no further details (Zariquiey, 1924). The specimens deposited in the MCNB collection have red labels with the printed inscription “ Typus ”, almost all of which have a collection date of October 1923 on their collection labels. There are specimens from the type series in other collections: Biospeologica (MNHNP) (Jeannel, 1924a) and FLC (two specimens according to Fresneda &amp; Salgado, 2016). In box 72 in the MNHNP general collection we have found at least eight specimens with the same red label stating “ Typus ” that probably correspond to the above-mentioned material from the Biospeologica collection. In the MNCNM collection (box 26) we have found around 20 specimens from the same locality with similar labels (but with a capture date of November 1923), none of which are identified as belonging to the type series.</p></div>	https://treatment.plazi.org/id/D42CA7775426CB6BFF6DFA937FA2FC4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775425CB6BFF6DFED47AEAF931.text	D42CA7775425CB6BFF6DFED47AEAF931.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Urbasolus) eloseguii Espanol 1948	<div><p>Speonomus (Urbasolus) eloseguii Español, 1948</p><p>Speonomus (Urbasolus) eloseguii Español, 1948: 240; figs. 1, 2b, 3.</p><p>Current accepted name</p><p>Euryspeonomus (Urbasolus) eloseguii (Español, 1948): Bellés et al., 1978: 65.</p><p>Type locality</p><p>Cueva de Ostalaza [original citation: “cueva de Ostalaza … en Urbasa occidental”].</p><p>Type material</p><p>Paratypes: MZB 78-1184 (1 ♂), type loc., 18-IX-1947, J. ElÓsegui &amp; F. Español leg. │ MZB 78-1185 (1 ♂), Cueva de Txintxaleze, 18-IX-1947, J. ElÓsegui &amp; F. Español leg. │ MZB 78-1186 (1 ♂), Cueva de Iniriturri, 18-IX-1947, J. ElÓsegui &amp; F. Español leg. │ MZB 78-1187 (1 ♂), Cueva de Arleze, 20-IX-1947, J. ElÓsegui &amp; F. Español leg. │ MZB 78-1188 (1 ♀), Cueva de Erbeltz, 20-IX-1947, J. ElÓsegui &amp; F. Español leg.</p><p>Remarks</p><p>Español (1945) established as “Tipo” (= holotype, article 73.1.1.) a male from Cueva de Ostalaza and also examined an unknown quantity of specimens from this and the other quoted caves. He does not specify where the specimens of the type series are deposited but does state that the holotype and ♀ allotype are deposited in the collection of the former Instituto Español de Entomología, today known as the MNCNM. Along with these specimens (box 24), we have found 22 paratypes. The specimens in the MCNB collection have red labels with the inscription “ Paratype ” hand-written by Español .</p></div>	https://treatment.plazi.org/id/D42CA7775425CB6BFF6DFED47AEAF931	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775425CB6AFF6DFA567930FD03.text	D42CA7775425CB6AFF6DFA567930FD03.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus ere Escola & Fresneda 2000	<div><p>Speonomus ere Escolà &amp; Fresneda, 2000</p><p>Speonomus ere Escolà &amp; Fresneda, 2000: 36; figs. 1–4.</p><p>Current accepted name</p><p>Vanderberghia ere (Escolà &amp; Fresneda, 2000): Fresneda et al., 2024: 716.</p><p>Type locality</p><p>Sima F-168 [original citation: “ Sima F-168 (1.800 m aprox.) en el valle de Hecho, Huesca”].</p><p>Type material</p><p>Holotype ♂: MZB 98-0852, type loc., 7-VIII-1997 / 12-VIII-1998 (trap), O. Escolà leg.</p><p>Paratypes: MZB 98-0851, MZB 98-1478 / MZB 98-1507 (14 ♂♂, 17 ♀♀), same data as holotype. │ MZB 98-1508 (1 ♀), Avenc Petit de la Reclusa, 8-VIII-1965, O. Escolà leg.</p><p>Remarks</p><p>Type series consisting of 18 ♂♂ and 20 ♀♀ from two cavities in the province of Huesca (Escolà &amp; Fresneda, 2000). Holotype seriously damaged. The rest of the paratypes are deposited in the EDC (1 ♂) and FLC (2 ♂♂ and 2 ♀♀) collections (Escolà &amp; Fresneda, 2000; Fresneda &amp; Salgado, 2016). On the labels of the trapped material, the collection date appears as 2 August 1998.</p></div>	https://treatment.plazi.org/id/D42CA7775425CB6AFF6DFA567930FD03	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775424CB6AFF6DFE1879DCFA11.text	D42CA7775424CB6AFF6DFE1879DCFA11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus escollae Fresneda & Hernando 1994	<div><p>Speonomus escollae Fresneda &amp; Hernando, 1994</p><p>Speonomus escollae Fresneda &amp; Hernando, 1994a: 63; figs. 1, 2, 6–13.</p><p>Current accepted name</p><p>Trapezodirus escollae (Fresneda &amp; Hernando, 1994): Fresneda, 1998: 60.</p><p>Type locality</p><p>Seira, MSS 002 [original citation: “ España, Huesca, Seira M.S.S. 002”].</p><p>Type material</p><p>Paratypes: MZB 2021-0842, MZB 2021-0843 (1 ♂, 1 ♀), type loc., 4-I-1992 / 4-IV-1992 (trap), J. Fresneda, Mª T. Escoll &amp; C. Hernando leg. (ex coll. X. Bellés) .</p><p>Remarks</p><p>Type series consisting of 41 ♂♂ and 38 ♀♀ from the superficial subterranean medium (SSM) of two localities in the province of Huesca (Fresneda &amp; Hernando, 1994a). Specimens of the type series deposited in other collections: ♂ holotype in the FLC (Fresneda &amp; Hernando, 1994a); paratypes in the CBC, CZULE, FLC, JCC, MGC, NMB, PMGC, (Fresneda &amp; Salgado, 2016). The paratypes originally deposited in the XBC (Fresneda &amp; Salgado, 2016) have been donated to the MCNB collection.</p></div>	https://treatment.plazi.org/id/D42CA7775424CB6AFF6DFE1879DCFA11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775424CB15FF6DFB3678CBFE63.text	D42CA7775424CB15FF6DFB3678CBFE63.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus espanyoli Auroux & Belles 1974	<div><p>Speonomus espanyoli Auroux &amp; Bellés, 1974</p><p>Speonomus espanyoli Auroux &amp; Bellés, 1974: 86; figs. 1–5.</p><p>Current accepted name</p><p>Bellesia espanyoli (Auroux &amp; Bellés, 1974): Fresneda &amp; Hernando, 1994b: 58.</p><p>Type locality</p><p>Cueva de Aso [original citation: “ Cueva de Aso, junto al Molino de Aso, cercano a Sercué, en el Valle de Añisclo, provincia de Huesca”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1070, type loc., 9-VIII-1974, Pallarés &amp; Auroux leg.</p><p>Paratypes: MZB 78-1071 / MZB 78-1077 (3 ♂♂, 4 ♀♀), same data as holotype. │ MZB 2021-0844 / MZB 2021- 0846 (1 ♂, 2 ♀♀), same data as holotype (ex coll. X. Bellés). │ MZB 78-1078 (remains 1 ex.), type loc., 1-IX-1973, Auroux &amp; Bellés leg.</p><p>Remarks</p><p>Auroux &amp; Bellés (1974) do not indicate where the specimens of the type series are deposited: of the 11 paratypes cited by these authors (5 ♂♂ and 6 ♀♀) collected together with the holotype, 10 are deposited in the MCNB collection (three of which have recently been donated by X. Bellés) .</p></div>	https://treatment.plazi.org/id/D42CA7775424CB15FF6DFB3678CBFE63	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545BCB15FF6DFCF878C3FB71.text	D42CA777545BCB15FF6DFCF878C3FB71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus espinosai Belles 1983	<div><p>Speonomus espinosai Bellés, 1983</p><p>Speonomus espinosai Bellés, 1983: 105; figs. 1, 3, 4, 7.</p><p>Current accepted name</p><p>Stygiophyes espinosai (Bellés, 1983): Fresneda, 1998: 61.</p><p>Type locality</p><p>Coves de Berganui [original citation: “Coves de Berganui, cerca del pueblo del mismo nombre, situado a pocos Km. de Areny de Noguera, provincia de Huesca, España ”].</p><p>Type material</p><p>Holotype ♂: MZB 2021-0817, type loc., 7-II-1982, X. Bellés &amp; J. Comas leg. (ex coll. X. Bellés).</p><p>Paratypes: MZB 2021-0818 / MZB 2021-0828 (4 ♂♂, 7 ♀♀), same data as holotype (ex coll. X. Bellés) │ MZB 2021-0829, MZB 2021-0830 (1 ♂, 1 ♀), type loc., 4-IV-1980, X. Bellés &amp; J. Espinosa leg. (ex coll. X. Bellés) │ MZB 2021-0831 / MZB 2021-0833 (2 ♂♂, 1 ♀), type loc., 2-I-1982, X. Bellés &amp; J. Comas leg. (ex coll. X. Bellés) .</p><p>Remarks</p><p>The whole type series donated by X. Bellés to the MCNB collection .</p></div>	https://treatment.plazi.org/id/D42CA777545BCB15FF6DFCF878C3FB71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545BCB14FF6DF8167913FF1B.text	D42CA777545BCB14FF6DF8167913FF1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus faurai subsp. esponellai Zariquiey 1940	<div><p>Speonomus faurai esponellai Zariquiey, 1940 (Fig. 15)</p><p>Speonomus faurai ssp. esponellai Zariquiey, 1940: 533 .</p><p>Current accepted name</p><p>Parvospeonomus delarouzeei guimjuani (Zariquiey, 1919): Fresneda et al., 2024: 695.</p><p>Type locality</p><p>Bauma de les Encantades [original citation: “Bauma de les Encantades, término municipal de Esponellá, provincia de Gerona”].</p><p>Type material</p><p>Syntypes: MZB 70-0345 / MZB 70-0355 (11 ♂♂, 25 ♀♀), type loc. (trap), R. Zariquiey leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Zariquiey (1940) mentions as “Tipos” a number of specimens deposited in his private collection. At a later date he reports the collection of numerous specimens in traps. He mentions that he examined over a hundred specimens to compare this new subspecies with Speonomus faurai Jeannel from the type locality: the material from Bauma de les Encantades was originally attributed to the later species (Jeannel, 1924a; Zariquiey, 1940). No collection date for this material is given. We have found 36 specimens in the MCNB collection from the type locality for esponellai, with collection labels that are either printed or hand-written by Zariquiey. None of these specimens have labels that identify them as part of the type series.</p></div>	https://treatment.plazi.org/id/D42CA777545BCB14FF6DF8167913FF1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545ACB14FF6DF9CE7814F844.text	D42CA777545ACB14FF6DF9CE7814F844.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Speonomus) lopezsellesi Espanol 1950	<div><p>Speonomus (Speonomus) lopezsellesi Español, 1950</p><p>Speonomus (s. str.) lopez-sellesi Español, 1950a: 108; fig. 6.</p><p>Current accepted name</p><p>Vanderberghia lopezsellesi Español, 1950: Fresneda et al., 2024: 716.</p><p>Type locality</p><p>Cueva de Espoz [original citation: “ Navarra: cueva de Espoz, en la vertiente noroeste de la sierra de Labia y a unos 2 Km. y medio del caserío de Espoz”].</p><p>Type material</p><p>Holotype ♂: MZB 98-1510, type loc., X-1949, Español leg.</p><p>Paratypes: MZB 98-1511 (allotype ♀), MZB 98-1512 (1 ♂), same data as holotype .</p><p>Remarks</p><p>Type series consisting of 2 ♂♂ and 2 ♀♀, (Español, 1950a); ♂ holotype originally designated as “Tipo” (article 73.1.1.). The paratype that completes the series is deposited in box 70 in the MNHNP’s general collection (Fresneda &amp; Salgado, 2016). These authors have transcribed the labels of this paratype. The collection labels of all the specimens in the type series are identical (with date “XI-49”).</p></div>	https://treatment.plazi.org/id/D42CA777545ACB14FF6DF9CE7814F844	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545ACB14FF6DFC207850FBB9.text	D42CA777545ACB14FF6DFC207850FBB9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus guimjuani Zariquiey 1919	<div><p>Speonomus guimjuani Zariquiey, 1919</p><p>Speonomus Guimjuani Zariquiey, 1919: 46; figs. 1–5</p><p>Current accepted name</p><p>Parvospeonomus delarouzeei guimjuani (Zariquiey, 1919): Fresneda et al., 2024: 695.</p><p>Type locality</p><p>Cova Bora Major [original citation: “Cova de Vora Major, en la montaña de Santa Magdalena, cerca del pueblo de Terrades, partido de Figueras, provincia de Gerona”].</p><p>Type material</p><p>Syntypes: MZB 78-2242 (1 ♂, 1 ♀), MZB 78-2549 / MZB 78-2551 (5 ♂♂, 2 ♀♀), type loc., VIII-1917, R. Zariquiey &amp; A. Guimjuan leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Type series consisting of 35 specimens (Zariquiey, 1919); however, the author neither designates a holotype nor indicates where the material is deposited. All of the specimens found in the MCNB collection have red labels printed with the inscription “Typus”, the only exceptions being two specimens (MZB 78-2242) with red hand-written labels with the inscription “COTIPO”. These latter two specimens have no collection labels, just an additional identification label, being regarded for the time as part of the type series. In the MNCNM collection (box 17) we have found a long series of specimens from the type locality, of which at least one has an identical collection label to the specimens in the MCNB collection (including the same date, August 1917). None of the above-mentioned specimens are identified as belonging to the type series .</p></div>	https://treatment.plazi.org/id/D42CA777545ACB14FF6DFC207850FBB9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775458CB16FF6DFD907FF3FC0A.text	D42CA7775458CB16FF6DFD907FF3FC0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Speonomus) mengeli Jeannel 1910	<div><p>Speonomus (Speonomus) mengeli Jeannel, 1910 (Fig. 58)</p><p>Speonomus (s. str.) Mengeli Jeannel, 1910a: CLIV; fig. 5.</p><p>Current accepted name</p><p>Troglocharinus mengeli (Jeannel, 1910): Fresneda et al., 2021: 169.</p><p>Type locality</p><p>Cova de Vinyoles [original citation: “... Cova de Vinyoles, située dans le termino municipal de Cava (partido de Seo de Urgel, provincia de Lerida)”].</p><p>Type material</p><p>Syntypes: MZB 78-2626 (1 ♂), type loc., 23-VIII-1910, R. Jeannel, E. G. Racovitza &amp; M. O. Mengel leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Type series consisting of 60 specimens originally deposited in the Biospeologica collection (Jeannel, 1910). The specimen in the MCNB collection has a label with the inscription “ TYPE ” printed in red and hand-written collection labels. We have located specimens belonging to the type series deposited in the Biospeologica collection (Jeannel, 1910, 1924a) in box 70 in the MNHNP general collection; their labels have been transcribed by Fresneda &amp; Salgado (2016) and Fresneda et al. (2021). In the MNCNM collection (box 20) we have found a few specimens that, given the associated data and collection labels, could belong to the type series, although they do not bear any labels that identify them as such (one specimen is indicated as a new species on its determination label).</p></div>	https://treatment.plazi.org/id/D42CA7775458CB16FF6DFD907FF3FC0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775458CB16FF6DF9117876F8A9.text	D42CA7775458CB16FF6DF9117876F8A9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus mercedesi Zariquiey 1922	<div><p>Speonomus mercedesi Zariquiey, 1922</p><p>Speonomus mercedesi Zariquiey, 1922b: 161; fig. pl. XI.</p><p>Current accepted name</p><p>Troglocharinus mercedesi (Zariquiey, 1922): Fresneda et al. (2021): 176.</p><p>Type locality</p><p>Bòfia de Sant Jaume [original citation: “ Bofia de S. Jaume, cerca de la masía llamada Boixadera dels Banchs, ya dentro de la prov. de Lérida y a unas cuatro horas y media de Berga”].</p><p>Type material</p><p>Syntypes: MZB 78-1451 / MZB 78-1453, MZB 78-2245, MZB 78-2581 / MZB 78-2583 (2 ♂♂, 5 ♀♀), type loc., 11-VII-1922, R. Zariquiey leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Type series consisting of 24 specimens, without indications regarding where they are deposited nor designation of holotype (Zariquiey, 1922b). The seven specimens deposited in the MCNB collection have red labels with the printed inscription “Typus”, the exception being MZB 78-2245, identified as a cotype on its red hand-written label. Jeannel (1924a) reports the deposit in the Biospeologica collection of part of the type series, without stating how many specimens. Fresneda &amp; Salgado (2016) and Fresneda et al. (2021) transcribe the labels associated with three specimens found in box 70 in the MNHNP collection, all identified as belonging to the type series. We have found in the MNCNM collection (box 17) a specimen identified as a cotype on its red label, whose collection labels are identical to most of the specimens deposited in the MCNB.</p></div>	https://treatment.plazi.org/id/D42CA7775458CB16FF6DF9117876F8A9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545ECB13FF6DF9E97F51FF1B.text	D42CA777545ECB13FF6DF9E97F51FF1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Speonomus) puncticollis Jeannel 1910	<div><p>Speonomus (Speonomus) puncticollis Jeannel, 1910</p><p>Speonomus (s. str.) puncticollis Jeannel, 1910a: CLVI; fig. 8.</p><p>Current accepted name</p><p>Stygiophyes puncticollis (Jeannel, 1910): Fresneda, 1998: 61.</p><p>Type locality</p><p>Forat de l’Or [original citation: “... «Forat del Or», situé dans le «Paso» de Tarradets (termino municipal de Llimiana, partido de Tremp, provincia de Lerida) ...”.</p><p>Type material</p><p>Syntypes: MZB 78-2558 (2 ♂♂), type loc., 31-VIII-1910, R. Jeannel &amp; E. G. Racovitza leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Jeannel (1910) does not specify how many specimens there are in the type series (“nombreux exemplaires” [numerous specimens]) originally deposited in the Biospeologica collection. The specimens in the MCNB (mounted on the same pin) have a label with the inscription “ TYPE ” printed in red letters with capture labels hand-written by Jeannel. Of the specimens of the type series deposited in the Biospeologica collection (Jeannel, 1910, 1924a), we have located a number of specimens in box 70 of the MNHNP’s general collection, whose labels have been transcribed by Fresneda &amp; Salgado (2016). In the MNCNM collection (box 20), we have located at least one specimen whose characteristics and collection date suggest that it could correspond to the type series (it has no label identifying it as such, but its determination label classifies it as a new species) .</p></div>	https://treatment.plazi.org/id/D42CA777545ECB13FF6DF9E97F51FF1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545ECB10FF6DFD9078DCFB52.text	D42CA777545ECB10FF6DFD9078DCFB52.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus porroiensis Escola & Comas 1983	<div><p>Speonomus porroiensis Escolà &amp; Comas, 1983</p><p>Speonomus porroiensis Escolà &amp; Comas, 1983: 259; figs. 2, 3.</p><p>Current accepted name</p><p>Lagariella porroiensis (Escolà &amp; Comas, 1983): Fresneda, 2000: 147.</p><p>Type locality</p><p>Sia del Tossal de Feliç [original citation: “Sia del Tossal de Felis, Porroi de la Solana (Baixa Ribagorça, province de Huesca)”].</p><p>Type material</p><p>Holotype ♂: MZB 73-2418, type loc., 13-XII-1980, J. Comas &amp; O. Escolà leg.</p><p>Paratypes: MZB 73-2419 / MZB 73-2429 (21 ♂♂, 30 ♀♀), same data as holotype. │ MZB 73-2437 (4 ♀♀), Sia de Santa Margarida (as “ Sia de Stª Marguerida ”), 19-I-1981, O. Escolà leg. │ MZB 73-2430 / MZB 73-2432, MZB 73- 2435, MZB 73-2436 (15 ♂♂, 10 ♀♀), Sia de Santa Margarida, 19-I-1981 / 26-IV-1981 (trap), C. Picañol &amp; O. Escolà leg. │ MZB 73-2433 (3 ♂♂), Sia de Santa Margarida, 26-IV-1981, C. Picañol &amp; O. Escolà leg. │ MZB 73-2438 / 73-2442 (14 ♂♂, 7 ♀♀), Pou Mina de Purroi (as “ Pou-Mina de Porroi ”), 20-I-1981, C. Picañol &amp; O. Escolà leg. │ MZB 73-2443 / MZB 73-2449, MZB 75-9332, MZB 75-9333 (25 ♂♂, 8 ♀♀), Pou Mina de Purroi, 19-I-1981 / 26- IV-1981 (trap), C. Picañol &amp; O. Escolà leg. │ MZB 73-2450, MZB 73-2451 (2 ♂♂, 1 ♀), Pou Mina de Purroi, 25- IV-1981, C. Picañol &amp; O. Escolà leg.</p><p>Remarks</p><p>Type series consisting of 98 ♂♂, 88 ♀♀ and an indeterminate number of remains from three caves in the Huescan Pyrenees (Escolà &amp; Comas, 1883). These authors refer to the ♂ holotype as “Types” (article 73.1.1.). The locality “ Av. Sia ” written on the label of the four females MZB 73-2437 corresponds to Sia de Santa Margarida. The remaining 45 paratypes are deposited in the JCC (Fresneda &amp; Salgado, 2016). Along with the type series, there are three remains with accession number MZB 73-2434, collected on 26 April 1981 in Sia de Santa Margarida (the remains belonging to the type series are dated 13 December 1980) .</p></div>	https://treatment.plazi.org/id/D42CA777545ECB10FF6DFD9078DCFB52	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545DCB13FF6DFC207E78FC29.text	D42CA777545DCB13FF6DFC207E78FC29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus saforensis Escola, Belles & Comas 1985	<div><p>Speonomus saforensis Escolà, Bellés &amp; Comas, 1985 (Fig. 59)</p><p>Speonomus saforensis Escolà, Bellés &amp; Comas, 1985: 225; figs. 4, 6, 9, 11, 12.</p><p>Current accepted name</p><p>Stygiophyes saforensis (Escolà, Bellés &amp; Comas, 1985): Fresneda, 1998: 61.</p><p>Type locality</p><p>Forat de Safor [original citation: “Avenc de Safor, Llastarri, t. m. Tremp (Pallars Jussà, Lleida)”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1019, type loc., 25-XI-1979 / 6-VII-1980 (trap), J. Comas &amp; O. Escolà leg.</p><p>Paratypes: MZB 78-1020 / MZB 78-1022, MZB 78-1024 / MZB 78-1028 (10 ♂♂, 18 ♀♀), same data as holotype. │ MZB 78-1023, MZB 78-1029 / MZB 78-1036 (20 ♂♂, 13 ♀♀), type loc., 28-XII-1982 / 20-IV-1985 (trap), C. Picañol &amp; O. Escolà leg.</p><p>Remarks</p><p>Escolà et al. (1985) do not indicate where the specimens of the type series (46 ♂♂ and 62 ♀♀) are deposited. Only some of the specimens have their original red paratype labels, although all unquestionably belong to the type series referred to by the authors of the description. There are 17 paratypes in the JCC (J. Comas pers. comm.) .</p></div>	https://treatment.plazi.org/id/D42CA777545DCB13FF6DFC207E78FC29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545DCB13FF6DF93E782EF8D0.text	D42CA777545DCB13FF6DF93E782EF8D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus sanctigervasi Jeannel 1911	<div><p>Speonomus sanctigervasi Jeannel, 1911 (Fig. 16)</p><p>Speonomus Sancti-Gervasi Jeannel, 1911a: LXXXIV; figs. 3, 4.</p><p>Current accepted name</p><p>Stygiophyes sanctigervasi (Jeannel, 1911): Fresneda, 1998: 61.</p><p>Type locality</p><p>Mines de la Canal [original citation: “« Minas de Canal », situées dans la sierra de San Gervas, distr. mun. de Llastarri, partido de Tremp, provincia de Lérida (Catalogne)”] .</p><p>Type material</p><p>Syntypes: MZB 78-2580 (1 ♂), type loc., 18-VI-1911, R. Jeannel &amp; E. G. Racovitza leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Type series consisting of three specimens originally deposited in the Biospeologica collection (Jeannel, 1911a, 1924a). The specimen in the MCNB has hand-written capture labels, a white label with the inscription “ TYPE ” hand-written in red letters, a red label with the printed inscription “ Typus ”, and a determination label hand-written by Jeannel. We have located the two specimens that complete the type series in the MNHNP’s general collection (box 69). Both have identical collection labels, as well as labels with the inscription “ TYPE ” (printed in black on a red label on one, and printed in red on a white label on the other).</p></div>	https://treatment.plazi.org/id/D42CA777545DCB13FF6DF93E782EF8D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545DCB12FF6DFA77788EFC4F.text	D42CA777545DCB12FF6DFA77788EFC4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus tincatincensis Escola, Belles & Comas 1985	<div><p>Speonomus tincatincensis Escolà, Bellés &amp; Comas, 1985</p><p>Speonomus tincatincensis Escolà, Bellés &amp; Comas, 1985: 223; figs. 1, 2, 5, 7, 10.</p><p>Current accepted name</p><p>Speonomites tincatincensis (Escolà, Bellés &amp; Comas, 1985): Fresneda, 1998: 68.</p><p>Type locality</p><p>Avenc del Tinc-ca-tinc [original citation: “Avenc del Tinc-ca-tinc, Altron (Pallars Sobirà, Lleida)”].</p><p>Type material</p><p>Holotype ♂ and its genitalia: MZB 78-1039, type loc., 12-X-1978, O. Escolà leg.</p><p>Paratypes: MZB 78-1040 (remains of 5 exs. including 1 genitalia ♂), MZB 78-1041 (2 ♀♀), same data as holotype. │ MZB 78-1042 / MZB 78-1053 (8 ♂♂ including 2 genitalia, 12 ♀♀), type loc., 17-XII-1978, X. Bellés, T. Cuñé &amp; O. Escolà leg.</p><p>Remarks</p><p>Although Escolà et al. (1985) only mentions the deposit of the holotype in the MCNB collection, we can confirm that the whole type series is deposited in the MCNB. These authors do not mention the remains of five specimens dated 12 October 1978 (MZB 78-1040), although probably they must have been examined when describing this new species (aedeagus, including its apex and internal sac are illustrated). These specimens have the same red paratypes labels as the rest of the series and the same collection data hand-written by Escolà. The information on the labels of the male genitalia preparations seems to indicate that they originate from the holotype, from one of the above-mentioned remains, and from two of the paratypes captured on 17 December 1978. The genitalia are mounted separately in microscopic preparations and preserved in Canadian balsam .</p></div>	https://treatment.plazi.org/id/D42CA777545DCB12FF6DFA77788EFC4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545CCB12FF6DFED47940F95D.text	D42CA777545CCB12FF6DFED47940F95D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus torresi Fresneda & Hernando 1990	<div><p>Speonomus torresi Fresneda &amp; Hernando, 1990</p><p>Speonomus torresi Fresneda &amp; Hernando, 1990: 86; figs. 1, 6, 8, 10, 12, 13, 15.</p><p>Current accepted name</p><p>Speonomites torresi (Fresneda &amp; Hernando, 1990): Fresneda, 1998: 67.</p><p>Type locality</p><p>Mines de Peramea [original citation: “Mines de Peramea (Gerri de la Sal, Pallars Sobirà, Lleida)”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1014, type loc., 10-VII-1985, J. Fresneda leg.</p><p>Paratypes: MZB 78-1015 / MZB 78-1018 (7 ♂♂, 6 ♀♀), type loc., 25-VII-1986 / 29-VIII-1986 (trap), J. Fresneda leg.</p><p>Remarks</p><p>Type series consisting of 142 ♂♂ and 184 ♀♀ (Fresneda &amp; Hernado, 1990). The authors indicate that the holotype and 20 paratypes are deposited in the MCNB, although we have only located thirteen of the paratypes. Paratypes deposited in other collections: CBC, CHC, FLC, JILCC, MBC, MPC, NMB and PMGC (Fresneda &amp; Hernando, 1990; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA777545CCB12FF6DFED47940F95D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777545CCB1DFF6DFBE2786FFD3F.text	D42CA777545CCB1DFF6DFBE2786FFD3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Speonomus) urgellesi Espanol 1964	<div><p>Speonomus (Speonomus) urgellesi Español, 1964 (Fig. 60)</p><p>Speonomus (s. str.) urgellesi Español, 1964: 55; figs. 2–3, 5–7.</p><p>Current accepted name</p><p>Parvospeonomus urgellesi (Español, 1964): Fresneda, 1998: 74.</p><p>Type locality</p><p>Avenc de les Aranyes [original citation: “Avenc de les Aranyes, Orsavinyà, en las inmediaciones de Pineda (prov. Barcelona)”].</p><p>Type material</p><p>Syntypes: MZB 78-0974, MZB 78-0975, MZB 78-0977 / MZB 78-0997 (42 ♂♂, 43 ♀♀, 2 exs), type loc., 1-XII-1963, I. González leg. │ MZB 75-9328, MZB 78-0976, MZB 78-0998 / MZB 78-1012 (34 ♂♂, 43 ♀♀), type loc., 26-I-1964, I. González, F. Sas &amp; F. Español leg.</p><p>Remarks</p><p>Español (1964) neither establishes a holotype nor the number of specimens in the type series, which, as far as we can ascertain, is deposited in its entirety in the MCNB. Specimen MZB 78-0974 has a red label hand-written by F. Español with the inscription “ Holotypus ”; the remaining specimens have red labels with the inscription “ Paratypus ”, either written by J. Comas or printed (“ Paratipo ” in the case of de MZB 78-0975). There is a discrepancy between one of the collection dates indicated by the author in his description (26 January 1964) and the date on the labels of the specimens deposited in the MCNB (19 January 1964). In the case of specimen MZB 78-0976, separated to extract its genitalia, the transcribed date is 19 January 1968 .</p></div>	https://treatment.plazi.org/id/D42CA777545CCB1DFF6DFBE2786FFD3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775453CB1DFF6DFE447A67F979.text	D42CA7775453CB1DFF6DFE447A67F979.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus vilarrubiasi Zariquiey 1940	<div><p>Speonomus vilarrubiasi Zariquiey, 1940 (Figs. 17A–B)</p><p>Speonomus vilarrubiasi Zariquiey, 1940: 532; fig. 7.</p><p>Current accepted name</p><p>Parvospeonomus vilarrubiasi (Zariquiey, 1940): Fresneda, 1998: 75.</p><p>Type locality</p><p>Cova de les Feixasses [original citation: “… cova de les Feixases, junto al Santuario de la Salud, sobre San Feliu de Pallarols, en el límite de las provincias de Barcelona y Gerona”].</p><p>Type material</p><p>Syntypes: MZB 70-0460 (1 ♀), type loc., 17-VI-1934, A. Vilarrubia leg. │ MZB 70-0458, MZB 70-0459, MZB 70-0461 (5 ♂♂, 7 ♀♀), type loc., 10-VIII-1934 (trap), R. Zariquiey leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Zariquiey (1940) cites as “Tipos” various specimens (without specifying the number) deposited in the collection of the Barcelona Museum and in his private collection. Subsequently, he states that one was collected by A. Vilarrubia (without indicating the date) and that a number of specimens were also trapped on 10 August 1934. With this date and origin we have found 12 specimens in the MCNB collection, five of which are mounted on the same pin (MZB 70-0461) with their original labels hand-written by Zariquiey (the others have more recent labels with the data copied by hand by O. Escolà). Likewise, we have found a specimen captured by A. Vilarrubia (MZB 70-0460) on 17 June 1934, which must corresponds to the specimen cited by Zariquiey (1940): Escolà &amp; Bellés (1984) refer to it as the first record (capture) from the type locality. None of the specimens have labels that identify them as part of the type series.</p></div>	https://treatment.plazi.org/id/D42CA7775453CB1DFF6DFE447A67F979	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775453CB1CFF6DFA0E7F59F85B.text	D42CA7775453CB1CFF6DFA0E7F59F85B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus vinyasi Escola 1971	<div><p>Speonomus vinyasi Escolà, 1971</p><p>Speonomus vinyasi Escolà, 1971: 32; figs. 1, 2.</p><p>Current accepted name</p><p>Troglocharinus vinyasi (Escolà, 1971): Fresneda &amp; Salgado, 2016: 208.</p><p>Type locality</p><p>Avenc del Mort (by designation of a lectotype (Fresneda et al., 2021: 181, fig. 70)) [original citation: “Avenc del «Mort» (Peramola)” (Escolà, 1971)].</p><p>Type material</p><p>Lectotype ♂: MZB 78-1084, type loc., 25-X-1970, C. Ribera leg.</p><p>Paralectotypes: MZB 78-1085 / MZB 78-1090 (3 ♂♂, 4 ♀♀), same data as holotype. │ MZB 78-1083, MZB 78- 1091 (1 ♂, 1 ♀), type loc., 1-XI-1969, C. Ribera, Mª D. Romero &amp; E. Roca leg. │ MZB 78-1092 (1 ♂), Avenc de Pau Esteve, 18-I-1970, M. Ubach &amp; O. Escolà leg. │ MZB 78-1093 (1 ♂), Avenc de Pau Esteve, 18-I-1970, O. Escolà leg. │ MZB 78-1094 (1 ♂), Forat de les Casanoves (as “ Avenc de Can Sala ”), 25-I-1970, C. Ribera &amp; Mª D. Romero leg. │ MZB 78-1095, MZB 78-1096 (1 ♂, 1 ♀), Forat de les Casanoves (as “ Coves de Can Sala ”), 12-III-1972, J. Bley &amp; L. Auroux leg. │ MZB 78-1097 / MZB 78-1103 (3 ♂♂, 10 ♀♀, 7 remains), Forat de les Casanoves (as “ Forat de les Cases Noves ”), 27-II-1972, J. Bley &amp; A. Meseguer leg.</p><p>Remarks</p><p>Specimen MZB 78-1083 has a red label with the printed inscription “Tipo”, while the other specimens have red labels with the printed inscriptions “ Paratypus ” or “ Paratipo ”. In the original publication, Escolà (1971) does not establish a holotype, although in the introduction he states that these are “los dos primeros representantes de esta nueva especie” [the first two examples of this new species], whose details correspond to the specimens MZB 78- 1083 and MZB 78-1091 (1 ♂ and 1 ♀, respectively). Escolà then describes the species and at the end of the article provides a list of specimens captured in each of the caves he refers to (a total of 15 ♂♂ and 12 ♀♀ from six caves). In a subsequent addendum, where more data are provided (Escolà, 1973), the author treats as paratypes all the specimens from all six localities mentioned in the original description, and specifies that the species was discovered in Avenc del Mort.</p><p>The MCNB holds 12 ♂♂, 16 ♀♀ and 7 remains from three caves (the author of the description used five different toponyms) labelled as part of the type series. These numbers do not coincide with those cited by Escolà (1971) but do match the collection data associated with the labels hand-written by the author himself. Specifically, Escolà (1971) cites 6 ♂♂ and 2 ♀♀ from Avenc del Mort on 25 October 1970, whilst in the collection there are 4 ♂♂ and 4 ♀♀ labelled with the same capture information. A ♂ specimen from Avenc del Pau Esteve with the accession number MZB 78-1092 and labelled “ Paratipo ” has the same information as MZB 78-1093 (apart from the names of the collectors, M. Ubach and O. Escolà) and is not included in the material mentioned by Escolà (1971). Nevertheless, both specimens appear on the record card of the author’s data file as “ paratypus ” and both are cited in a subsequent publication (Escolà, 1980b).</p><p>Escolà (1971) cites 4 ♂♂ and 7 ♀♀ from “Forat de les Cases Noves”, although in the MCNB collection there are 3 ♂♂ and 10 ♀♀ labelled as paratypes, along with the remains of a further seven specimens. For all these specimens the same capture information is given (both in the description and on the labels the collection date is subsequent to the publication date). On the record card of the Escolà’s data file, these “ paratypes ” correspond to the four previously mentioned males and 11 females, along with other remains preserved in ethanol. Subsequently, the author indicates on the same record card that one of the males does not belong to the type series and in fact corresponds to a Troglocharinus ludovici Bellés &amp; Déliot, 1983 . This specimen has been localized in the general collection, mounted with its genitalia .</p><p>The labels of the two specimens from “ Coves de Can Sala ” contain the date of capture, not mentioned by Escolà (1971) .</p><p>Some specimens of the type series of Speonomus vinyasi have turned out to belong to another species, Troglocharinus fadriquei Fresneda, Rizzo, Comas &amp; Ribera, 2021, recently described (Fresneda et al., 2021). Consequently, the aforementioned authors have designated lectotype (MZB 78-1084) and paralectotypes of S. vinyasi (the rest of the series); specimens MZB 78-1094 / MZB 78-1103 (5 ♂♂, 11 ♀♀ and 7 remains) have been designated at the same time as paratypes of T. fadriquei (see below).</p><p>All specimens of the type series of S. vinyasi are deposited in the MCNB collection, except one paralectotype from Avenc de la Roca del Corb, deposited in the JCC (Fresneda et al., 2021) .</p></div>	https://treatment.plazi.org/id/D42CA7775453CB1CFF6DFA0E7F59F85B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775450CB1EFF6DFD907E31FBC2.text	D42CA7775450CB1EFF6DFD907E31FBC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speonomus (Speonomus) zariquieyi Jeannel 1924	<div><p>Speonomus (Speonomus) zariquieyi Jeannel, 1924 (Fig. 61)</p><p>Speonomus (s. str.) Zariquieyi Jeannel, 1924a: 159 .</p><p>Current accepted name</p><p>Stygiophyes zariquieyi (Jeannel, 1924): Fresneda &amp; Escolà, 2000: 62.</p><p>Type locality</p><p>Cova Negra de Corçà [original citation: “Lerida: cova negra de Corsá, au sud du Montsech de Ager, termino mun. de Ager ...”] .</p><p>Type material</p><p>Syntypes: MZB 70-0941 / MZB 70-0948, MZB 78-2559 / MZB 78-2579 (48 ♂♂, 76 ♀♀), type loc., R. Zariquiey leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Type series consisting of numerous specimens (with no more details) deposited originally in Zariquiey’s collection and in the Biospeologica collection (Jeannel, 1924a). Most of the specimens deposited in the MCNB collection have red labels with the printed inscription “Cotypus” or “ Typus ”. The two labels with collection data are identical throughout the whole series, January 1924 being given as a capture date for most of the series. Three specimens (MZB 70-0945) are separate and have been re-labelled by hand by O. Escolà. In the MNHNP general collection (box 69) we have found at least five specimens from the type series (one with the inscription “ TYPE ” printed in black letters on a red label, and the rest with the same inscription printed in red characters on white labels). In the MNCNM collection (box 17) there is a long series of specimens with collection labels that are identical to the specimens in the MCNB, although none of them have labels that identify them as belonging to the type series .</p></div>	https://treatment.plazi.org/id/D42CA7775450CB1EFF6DFD907E31FBC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775450CB1EFF6DF95A7A00F845.text	D42CA7775450CB1EFF6DF95A7A00F845.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus carrodillae Jeannel 1911	<div><p>Speophilus carrodillae Jeannel, 1911 (Fig. 18)</p><p>Speophilus Carrodillae Jeannel, 1911a: XCIV; fig. 14.</p><p>Current accepted name</p><p>Trapezodirus carrodillae (Jeannel, 1911): Fresneda, 1998: 59.</p><p>Type locality</p><p>Grallera de Estadilla [original citation: “l’aven dit « la Grallera de Estadilla », situé dans la sierra de la Carrodilla, près d’Estadilla, partido de Tamarite-de-Litera, provincia de Huesca (Aragon). Cette grotte dépend du bassin du rio Cinca”] .</p><p>Type material</p><p>Syntypes: MZB 78-1316 (1 ♂), type loc., 2-VII-1911, R. Jeannel &amp; E. G. Racovitza leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>The type series consists of 18 specimens originally deposited in the Biospeologica collection (Jeannel, 1911a). We have found in the MNHNP general collection (box 73) the four specimens of the type series cited by Fresneda &amp; Salgado (2016) who transcribe the inscriptions on their labels. Specimen MZB 78-1316 has almost identical collection labels, as well as a label with the inscription “ TYPE ” printed in red letters on one label. This specimen also has a label with the determination “ Speophilus (Trapezodirus) Estadillae Jeann., R. JEANNEL det.”. In the MNCNM collection (box 27) we have found a ♂ labelled as a cotype, which also has identical labels to the specimen from the MCNB .</p></div>	https://treatment.plazi.org/id/D42CA7775450CB1EFF6DF95A7A00F845	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775457CB19FF6DF93E7A3FF845.text	D42CA7775457CB19FF6DF93E7A3FF845.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus (Speophilus) espanoli Jeannel 1930	<div><p>Speophilus (Speophilus) españoli Jeannel, 1930 (Figs. 19, 63)</p><p>Speophilus (s. str.) Españoli Jeannel, 1930: 91 .</p><p>Current accepted name</p><p>Troglocharinus espanoli (Jeannel, 1930): Fresneda &amp; Salgado, 2016: 191.</p><p>Type locality</p><p>Cova del Traça [original citation: “une grotte des environs de Tarragona, sur le Pla de Cabra”].</p><p>Type material</p><p>Syntypes: MZB 78-1214 (1 ♀), type loc., F. Español leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>The cave mentioned by Jeannel (1930) is Cova del Traça, near the hamlet of Fontscaldetes, Pla de Cabra, Tarragona province (Español, 1934, 1950b). This species is only known from this locality (Salgado et al., 2008; Fresneda &amp; Salgado, 2016). According to Jeannel (1930), the type series consists of three females deposited in the collections of Jeannel (MNHNP) and Zariquiey (MCNB). The specimen located (MZB 78-1214) has a red hand-written label with the inscription “COTIPO”. According to the collection label, this specimen was captured in 1928. The date, which is hard to read, was probably July 1928, as cited by Español (1950b) and, more precisely, 16 July 1928, as reported by Escolà (1977, 1982). In the general collection of the MNHNP (box 73) we have found a specimen with a red label and the inscription “ TYPE ” printed in black. The determination, attributed to Zariquiey (1929), on one of its labels is of a new genus and species (Espagnolia roseti) that was never formally described, which is also mentioned by Jeannel (1930).</p></div>	https://treatment.plazi.org/id/D42CA7775457CB19FF6DF93E7A3FF845	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775457CB19FF6DFD907A90FC26.text	D42CA7775457CB19FF6DFD907A90FC26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus cenarroi Espanol 1955	<div><p>Speophilus cenarroi Español, 1955 (Fig. 62)</p><p>Speophilus cenarroi Español, 1955: 263; pl. VIII, figs. 1, 2a, 3.</p><p>Current accepted name</p><p>Ceretophyes cenarroi (Español, 1955): Fresneda, 1998: 79.</p><p>Type locality</p><p>Forat de les Gralles [original citation: “ Sima «Forat de les Gralles» (Bellver de Cerdaña)”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2067, type loc., 17-VI-1950, F. Español leg.</p><p>Paratypes: MZB 78-2068 / MZB 78-2075 (8 ♀♀), type loc., 17-VI-1950 and 24-IX-1950, F. Español leg.</p><p>Remarks</p><p>According to Español (1955), the type series includes 11 paratypes (all female), eight of which have been located in the MCNB collection. On the collection label of the holotype (the only male, cited as “Tipo”, article 73.1.1.) appears the date September 1950, which does not correspond to published capture date (17 June 1950). The same occurs in the case of the paratypes except for MZB 78-2075 whose label does include the date 17 June 1950 (according to Español, 1955, five paratypes were collected along with a male, the rest being collected on 24 September 1950). Español (1953b) mentions the new subspecies in the section “Índice cronológico de los trabajos publicados (1952)” [Chronological index of articles published (1952)] in which he reports the on-going study of the corresponding material, and indicates the title and journal of publication. The name becomes available with the publication of the description (article 13.1.1).</p></div>	https://treatment.plazi.org/id/D42CA7775457CB19FF6DFD907A90FC26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775456CB18FF6DFD9079CCFC6E.text	D42CA7775456CB18FF6DFD9079CCFC6E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus (Trapezodirus) fonti subsp. schuttei Espanol 1955	<div><p>Speophilus (Trapezodirus) fonti schüttei Español, 1955</p><p>Speophilus (Trapezodirus) fonti ssp. schüttei Español, 1955: 268; pl. VIII, fig. 4b.</p><p>Current accepted name</p><p>Troglocharinus schuttei (Español, 1955): Fresneda et al., 2024: 776.</p><p>Type locality</p><p>Cova de la Font Mentidora [original citation: “ Cueva «Font Mentidora» (Sierra del Bou Mort)”].</p><p>Type material</p><p>Syntypes: MZB 78-1298 / MZB 78-1309 (6 ♂♂, 6 ♀♀), type loc., 30-IX-1950, F. Español leg.</p><p>Remarks</p><p>Español (1955) mentions “unos pocos ejemplares” [a few specimens], males and females, without specifying either their number or place of deposit. The specimens found in the MCNB collection have red hand-written labels with the inscription “Serie típica”. According to Español (1955: 262), the type locality was prospected on 30 September [1950] during the second of the biospeleological campaigns described. On the hand-written collection labels appears the date October 1952. Español (1953b) reports the new subspecies in the section “Índice cronológico de los trabajos publicados (1952)” [Chronological index of articles published (1952)] in which he reports the on-going study of the corresponding material, and indicates the title and journal of publication. The name becomes available with the publication of the description (article 13.1.1).</p></div>	https://treatment.plazi.org/id/D42CA7775456CB18FF6DFD9079CCFC6E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775456CB18FF6DFEF67AC1F8D0.text	D42CA7775456CB18FF6DFEF67AC1F8D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus (Trapezodirus) fonti subsp. zariquieyi Jeannel 1924	<div><p>Speophilus (Trapezodirus) fonti zariquieyi Jeannel, 1924</p><p>Speophilus (Trapezodirus) Fonti ssp. Zariquieyi Jeannel, 1924a: 172 .</p><p>Current accepted name</p><p>Troglocharinus zariquieyanus Salgado, Blas &amp; Fresneda, 2008: Fresneda et al., 2024: 777.</p><p>Type locality</p><p>Cova de Guils [original citation: “Lerida: cova de Guils, à Mata Rossa, près de Guils del Cantó, au nord-ouest de Novés”].</p><p>Type material</p><p>Syntypes: MZB 70-4098, MZB 78-1310 / MZB 78-1314, MZB 78-2647 / MZB 78-2652 (5 ♂♂, 9 ♀♀), type loc., R. Zariquiey leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Jeannel (1924a) does not specify the exact number of specimens in the type series, and indicates that it is deposited in the collections of R. Zariquiey (MCNB) and Bioespeologica (MNHNP). The specimens found in the MCNB collection have red labels with the printed inscription “ Typus ” or “Cotypus”. In the MNHNP general collection (box 73) are deposited three specimens whose labels are transcribed by Fresneda &amp; Salgado (2016). In the MNCNM collection (box 27) we have found a specimen with the same collection data but not identified as belonging to the type series. The labels indicating the localities of origin of the specimens deposited in the three above-mentioned museums are identical.</p></div>	https://treatment.plazi.org/id/D42CA7775456CB18FF6DFEF67AC1F8D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775456CB1BFF6DFA7779DBF973.text	D42CA7775456CB1BFF6DFA7779DBF973.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus jacasi Lagar 1966	<div><p>Speophilus jacasi Lagar, 1966 (Figs. 20A–B, 64)</p><p>Speophilus jacasi Lagar, 1966: 29; figs. 1, 2a, 3.</p><p>Current accepted name</p><p>Troglocharinus jacasi (Lagar, 1966): Fresneda &amp; Salgado, 2016: 199.</p><p>Type locality</p><p>Cova de les Rondes [original citation: “cueva de «Les Rondes», cavidad situada sobre «Cal Marquet», en la cabecera del barranco de la «Roca del Frare», en las inmediaciones del castillo de Vilademager, municipio de La Llacuna, provincia de Barcelona”].</p><p>Type material</p><p>Paratypes and possible paratypes: MZB 78-1215 / MZB 78-1225, MZB 78-1242 / MZB 78-1249 (12 ♂♂, 51 ♀♀), type loc., 1-X-1964 O. Escolà leg. [cited by Lagar (1966) as O. Escolà leg. XI-1965?] │ MZB 78-1226 / MZB 78- 1241, MZB 78-1250 / MZB 78-1266 (52 ♂♂, 50 ♀♀), type loc., X-1965, F. Rodríguez leg. [cited by Lagar (1966) as O. Escolà leg. XI-1965?] │ MZB 2021-0847, MZB 2021-0848 (1 ♂, 1 ♀), type loc., 10-XII-1965, J. Mª Jacas, F. Raventós &amp; A. Lagar (ex coll. X. Bellés).</p><p>Remarks</p><p>The type series cited by Lagar (1966) consists of the following material: 1 ♂ (the holotype) and “numerosos paratipos de ambos sexos” [numerous paratypes of both sexes] collected on 10 December 1965 (Jacas, Raventós and Lagar leg.); “una serie importante” [a long series] of specimens collected in November 1965 by O. Escolà and deposited in the MCNB collection; and, lastly, “algunos ejemplares” [a few specimens] collected in December 1958 and November 1959 (L. Mª Udina-A.Poyo leg.) deposited in Lagar’s personal collection. All specimens originate from the type locality. Fresneda &amp; Salgado (2016) give the number of specimens deposited in the FLC collection, which include the holotype and paratypes of the material cited by Lagar (1966) except the specimens collected by O. Escolà. There are two paratypes deposited in the CJC “Lagar-Jacas leg.”, donated by Lagar (J. Comas, pers. comm.). Other two paratypes deposited in the XBC (collection date of 10 December 1965) have recently been donated to the MCNB collection.</p><p>The MCNB collection has numerous specimens from the type locality (64 ♂♂ and 101 ♀♀) labelled as paratypes, with their corresponding red labels, some of which also have labels with the inscription “ Speophilus jacasi nov. sp. ”. Nevertheless, the dates and/or collectors associated with this material do not correspond to the information given by Lagar (1966). A total of 102 specimens (52 ♂♂ and 50 ♀♀) were collected by F. Rodríguez in October 1965 (data cited by Escolà, 1977). The labels of most of these specimens seem to have been hand-written by Lagar, according to the opinion of J. Fresneda (pers. comm.); the rest were hand-written by O. Escolà. Rodríguez was a collaborator who donated the specimens he collected to the Museum; this material was added to the series collected by Escolà (O. Escolà, pers. comm.). No specimens originating from F. Rodríguez is to be found in the CFL collection, according to Fresneda &amp; Salgado (2016). On the labels written by Escolà corresponding to the remaining specimens (O. Escolà leg.) the collection date is given as 1 October 1964 (information cited by Escolà, 1977) rather than November 1965, the date that Lagar (1966) attributes to the paratypes deposited in the MCNB. We believe that it is likely that this discrepancy is due to an error in the publication date, given that on the record card of the Escolà’s data file, the entry associated with the collection date of 1 October 1964 has the inscription “ Speophilus jacasi (paratipus) (prep) 45 ex Bathysciinae”. In this record card there is no record of any material being entered in November 1965 (nor any specimen in the general collection).</p><p>In light of the material examined, and taking into account the information provided by J. Fresneda and O. Escolà, it seems likely that all the material deposited in the MCNB is part of the type series and was attributed by Lagar (1966) to the collector and date cited .</p></div>	https://treatment.plazi.org/id/D42CA7775456CB1BFF6DFA7779DBF973	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775455CB1AFF6DFA097FABFBDF.text	D42CA7775455CB1AFF6DFA097FABFBDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus kiesenwetteri subsp. andresi Escola 1966	<div><p>Speophilus kiesenwetteri andresi Escolà, 1966</p><p>Speophilus kiesenwetteri ssp. andresi Escolà, 1966: 200; fig. 1a</p><p>Current accepted name</p><p>Troglocharinus andresi (Escolà, 1966): Fresneda et al., 2024: 774.</p><p>Type locality</p><p>Avenc Montserrat [original citation: “Avenc Montserrat ... a 420 m. de altura, en las proximidades de Coll Cassot, no muy lejos de la conocida ermita de St. Salvador ”] .</p><p>Type material</p><p>Syntypes: MZB 78-1204 (1 ♂), type loc., XII-1962, O. Andrés leg. │ MZB 78-1197 / MZB 78-1199 (8 ♀♀), type loc., 31-XII-1962, O. Escolà leg. │ MZB 78-1195, MZB 78-1196, MZB 78-1200 / MZB 78-1203, MZB 78-1206 / MZB 78-1213 (17 ♂♂, 27 ♀♀), type loc., 31-XII-1962 / 4-I-1963 (trap), O. Escolà leg. │ MZB 78-1205 (1 ♂), type loc., 7-IV-1963, O. Escolà leg.</p><p>Remarks</p><p>Escolà (1966) neither establishes a holotype nor states exactly how many specimens there are in the type series. In the introduction to the description, the author mentions the collection (directly or by trapping) of around 20 specimens during fieldwork performed between 30 December 1962 and 4 January 1963 (a number of specimens that he regarded as sufficient for study). Subsequently, however, in the body of the description, he indicates the choice of 10- 15 specimens of each sex to be measured (chosen at random, which implies that the original series must have had a larger number of specimens). The specimens located in the MCNB collection (19 ♂♂ and 35 ♀♀) have similar hand-written capture labels (mostly by O. Escolà himself) and red labels inscribed with “ Paratipo ” or “ Paratypus ”. Aside from the specimens captured in the above-mentioned period, the specimen MZB 78-1205 is also present, with a subsequent capture date (7 April 1963, O. Escolà leg.) that predates the publication of the description. This specimen also has a label with the inscription “org. cop. ♂ ” that indicates that its genitalia was studied, as well as a label indicating “Speoph. Kiesen. / andresi nov. / Escolá det.”, hand-written by F. Español. Additionally, this latter specimen is identified as a paratype on the record card of the Escolà’s data file. We have been unable to locate in the MCNB collection the two specimens that Escolà (1966) cites with the capture date of 1 January 1962 (O. Andrés leg.). Escolà (1977) provides the capture data (date and collector) of all the material discussed here.</p></div>	https://treatment.plazi.org/id/D42CA7775455CB1AFF6DFA097FABFBDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775454CB05FF6DF9657A4DFE3B.text	D42CA7775454CB05FF6DF9657A4DFE3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus kiesenwetteri subsp. castellsaperai Jeannel 1924	<div><p>Speophilus kiesenwetteri castellsaperai Jeannel, 1924 (Figs. 21, 65)</p><p>Speophilus Kiesenwetteri subraza Castellsaperai Zariquiey, 1924: 10; fig. 4. (unavailable)</p><p>Speophilus (Speophilus) Kiesenwetteri subsp. Castellsaperai Jeannel, 1924a: 171.</p><p>Current accepted name</p><p>Troglocharinus sanllorensi (Zariquiey, 1924): Fresneda et al., 2024: 776.</p><p>Type locality</p><p>Avenc de Castellsapera [original citation: “Avenc de Castellsapera, Sant Llorenç de Munt”].</p><p>Type material</p><p>Syntypes: MZB 78-1194, MZB 78-1455 / MZB 78-1461, MZB 78-2708, MZB 78-2709 (5 ♂♂, 5 ♀♀), type loc., 25-XI-1922, S. Novellas leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1922a) in a previous publication announced the discovery of a new taxon (a subspecies of Speophilus kiesenwetteri), based on four specimens collected by A. Codina on 12 October 1913, but provided no description. Subsequently, Zariquiey (1924) proposed the name castellsaperai as a infrasubspecific entity (“subraza”) as is shown in the publication (article 45.6.1.) in which he also described another new entities explicitly as subspecies of Speophilus kiesenwetteri . The name castellsaperai was used at a later date by Jeannel (1924a), which made it available for a subspecies of S. kiesenwetteri, Jeannel thereby becoming the author of this new subspecies (article 45.5.1.).</p><p>Zariquiey (1924) mentions “bastantes ejemplares” [plenty of specimens] collected by S. Novellas, of which we have located 10 in the MCNB collection, bearing red labels printed with “ Typus ” (the only exception is one labelled as “COTIPO”). Zariquiey also cites an indeterminate number of specimens collected by Faura i Sans and the four above-mentioned specimens collected by Codina. The specimens of the type series collected by Faura i Sans were deposited in the Biospeologica collection (Zariquiey, 1924; Jeannel, 1924a) that is currently in the MNHNP. In fact, in box 73 of the general collection of the MNHNP we have found two of these specimens, labelled as cotypes, as well as one of the specimens of the series captured by Novelles (with identical capture labels to those of the specimens deposited in the MCNB and with the same red printed labelled indicating “ Typus ”). After Zariquiey’s description (1924), Codina (1924) cites three “Tipus” along with nine other specimens from the type locality with a capture date of 12 October 1913 deposited in his private collection. However, we have failed to find in the MCNB collection any of the specimens captured by Codina. In the MNCNM collection (box 27), we have found a specimen with an identical capture label to those from the series in the MCNB and with a red label printed with the inscription “Cotypus”.</p></div>	https://treatment.plazi.org/id/D42CA7775454CB05FF6DF9657A4DFE3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544BCB05FF6DFF41785FFAE2.text	D42CA777544BCB05FF6DFF41785FFAE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus kiesenwetteri subsp. patracoi Zariquiey 1922	<div><p>Speophilus kiesenwetteri patracoi Zariquiey, 1922</p><p>Speophilus Kiesenwetteri ssp. Patracoi Zariquiey, 1922b: 162; pl. XI.</p><p>Current accepted name</p><p>Troglocharinus patracoi (Zariquiey, 1922): Fresneda &amp; Salgado, 2016: 205.</p><p>Type locality</p><p>Cova del Patracó [original citation: “Cueva del PATRACÓ, en el término municipal de Esparraguera, (Prov. de Barcelona)”].</p><p>Type material</p><p>Syntypes: MZB 78-1189 / MZB 78-1193, MZB 78-2713 / MZB 78-2723 (5 ♂♂, 11 ♀♀), type loc., 6-IX-1922, R. Zariquiey leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1922b) refers to the collection of “varios ejemplares” [various specimens] but gives no further details nor indicates where they are deposited. We have located 16 specimens of the type series in the MCNB collection, all of which have red labels printed with the inscription “ Typus ”. There are an unknown number of specimens of the type series deposited in the Bioespeologica collection (Jeannel, 1924a). In box 73 of the MNHNP’s general collection we have found at least one specimen identified as a cotype.</p></div>	https://treatment.plazi.org/id/D42CA777544BCB05FF6DFF41785FFAE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544BCB04FF6DF87A7A2AFE47.text	D42CA777544BCB04FF6DF87A7A2AFE47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus kiesenwetteri subsp. sanllorensi Zariquiey 1924	<div><p>Speophilus kiesenwetteri sanllorensi Zariquiey, 1924 (Fig. 67)</p><p>Speophilus Kiesenwetteri ssp. San Llorensi Zariquiey, 1924: 10; fig. 3.</p><p>Current accepted name</p><p>Troglocharinus sanllorensi (Zariquiey, 1924): Fresneda et al., 2024: 776.</p><p>Type locality</p><p>Cova de Santa Agnès [original citation: “Cova de Sta. Agnés, Sant Llorenç de Munt, prov. Barcelona”].</p><p>Type material</p><p>Syntypes: MZB 78-1462 / MZB 78-1466, MZB 78-2247, MZB 78-2248, MZB 78-2696 / MZB 78-2707 (20 ♂♂, 13 ♀♀), type loc., 3-XII-1922, S. Novellas leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1924) mentions “numerosos ejemplares” [numerous specimens] but provides no more information and does not indicate where they are deposited. Part of the type series is deposited in the Bioespeologica collection (Jeannel, 1924a). Most of the 33 specimens located in the MCNB collection have red labels with the inscription “ Typus ”; those that do not, are labelled as cotypes. In box 73 of the MNHNP general collection we have found at least five specimens with the same red label “ Typus ” as most of the specimens in the type series deposited in the MCNB. In the MNCNM collection (box 27) we have found two specimens mounted on the same pin with a red label “ Typus ”, and two others with a red cotype label. All these specimens have identical —or very similar— collection labels to the specimens in the type series deposited in the MCNB. Zariquiey (1924) indicates that material pertaining to this new taxon from caves near the type locality (Faura i Sans leg.) is deposited in the Jeannel and Museo de Madrid collections.</p></div>	https://treatment.plazi.org/id/D42CA777544BCB04FF6DF87A7A2AFE47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544ACB04FF6DFCDD7ABBFABA.text	D42CA777544ACB04FF6DFCDD7ABBFABA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus quadricollis Jeannel 1911	<div><p>Speophilus quadricollis Jeannel, 1911</p><p>Speophilus quadricollis Jeannel, 1911a: XCV; figs. 15, 16.</p><p>Current accepted name</p><p>Troglocharinus quadricollis (Jeannel, 1911): Fresneda &amp; Salgado, 2016: 206.</p><p>Type locality</p><p>Graller de Castellet [original citation: “grotte dite «Lo Grallé», située près de Castellet, districto mun. d’Espluga de Serra, partido de Tremp, provincia de Lérida (Catalogne). Elle se trouve dans la sierra de Lleras, sur le versant de la Noguera Ribagorzana”].</p><p>Type material</p><p>Syntypes: MZB 78-1315 (1 ♂), type loc., 17-VI-1911, R. Jeannel &amp; E. G. Racovitza leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>The type series consist of 20 specimens originally deposited in the Biospeologica collection (Jeannel, 1911a). Alongside MZB 78-1315, we have found in the MCNB collection two other males (MZB 78-1482) with identical hand-written collection labels but not identified as belonging to the type series (nevertheless, they may in fact be a part), bearing a circular acetate label. We have also found, with identical collection dates and labels as MZB 78-1315 a large part of the type series in the MNHNP general collection (box 73): two specimens with red labels with the inscription “ TYPE ” printed in black letters, and 10 specimens with the inscription “ TYPE ” in red letters on a white background. Fresneda &amp; Salgado (2016) transcribe the inscription on the labels of these 12 specimens belonging to the type series in the MNHNP. In the MNCNM collection (box 27), we have found three specimens with similar collection labels: two have the same white label with the inscription “ TYPE ” in red letters, while the third is identified as a cotype.</p></div>	https://treatment.plazi.org/id/D42CA777544ACB04FF6DFCDD7ABBFABA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544ACB07FF6DF8C27904FF1B.text	D42CA777544ACB07FF6DF8C27904FF1B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus schibii Espanol 1972	<div><p>Speophilus schibii Español, 1972</p><p>Speophilus schibii Español, 1972b: 56; figs. 1a–c.</p><p>Current accepted name</p><p>Troglocharinus schibii (Español, 1972): Fresneda &amp; Salgado, 2016: 207.</p><p>Type locality</p><p>Avenc de la Solana [original citation: “Avenc de La Solana, Serra d’Ancosa, 900 m. de altitud, La Llacuna, prov. Barcelona”] .</p><p>Type material</p><p>Holotype ♂: MZB 78-1267, type loc., 8-VIII-1971, Schibi &amp; Nubiola leg.</p><p>Paratypes: MZB 78-1278, MZB 78-1279 (2 ♀♀), same data as holotype. │ MZB 78-1268 / MZB 78-1277, MZB 78-1280 / MZB 78-1285 (10 ♂♂, 6 ♀♀), type loc., 12-X-1971, Schibi, Cantons, Ribes &amp; Español leg.</p><p>Remarks</p><p>Full type series deposited in the MCNB.</p></div>	https://treatment.plazi.org/id/D42CA777544ACB07FF6DF8C27904FF1B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775449CB07FF6DFC2179FDFC29.text	D42CA7775449CB07FF6DFC2179FDFC29.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Speophilus subilsi Espanol 1966	<div><p>Speophilus subilsi Español, 1966 (Fig. 66)</p><p>Speophilus subilsi Español, 1966b: 5; pl. 1, figs. 4, 5</p><p>Current accepted name</p><p>Troglocharinus subilsi (Español, 1966): Fresneda &amp; Salgado, 2016: 207.</p><p>Type locality</p><p>Grallera de Cambrils; Avenc de l’Obaga del Grau [original citation: “Lérida: Grallera de Cambrils, Llinás, Oliana, en la Sª de Odén … avenc de l’Obaga del Grau, Alsina d’Alinyá, estribaciones septentrionales de la Sª de Odén”].</p><p>Type material</p><p>Syntypes: MZB 78-1317 / MZB 78-1324 (5 ♂♂, 4 ♀♀), Avenc de l’Obaga del Grau, Subils leg. │ MZB 78-1325 / MZB 78-1329 (5 ♂♂), Grallera de Cambrils, Escolà &amp; Senent leg.</p><p>Remarks</p><p>Español (1966b) does not establish a holotype nor the number of specimens in the type series (all of which we believe are deposited in the MCNB collection). We have found 14 specimens (10 ♂♂ and 4 ♀♀) with red labels, hand-written by Español with the inscription “ Paratypus ”; the only exception is the male specimen MZB 78-1317, which has a label with the inscription “Typus” .</p></div>	https://treatment.plazi.org/id/D42CA7775449CB07FF6DFC2179FDFC29	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775449CB07FF6DF93E7E5AF918.text	D42CA7775449CB07FF6DF93E7E5AF918.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stygiophyes aldomai subsp. allomorphus Fresneda & Escola 2000	<div><p>Stygiophyes aldomai allomorphus Fresneda &amp; Escolà, 2000</p><p>Stygiophyes aldomai allomorphus Fresneda &amp; Escolà, 2000: 56; figs. 2, 4.</p><p>Current accepted name</p><p>Stygiophyes aldomai allomorphus Fresneda &amp; Escolà, 2000</p><p>Type locality</p><p>Forat d’Irgo [original citation: “Lleida. El Pont de Suert, Forat d’Irgo”].</p><p>Type material</p><p>Holotype ♂: MZB 98-1509, type loc., 7-X-1985, J. Fresneda leg.</p><p>Paratypes: MZB 73-2411 / MZB 73-2413 (3 ♂♂, 9 ♀♀), type loc., 27-VII-1986, O. Escolá, C. Picañol &amp; J. Fresneda leg. │ MZB 73-2414 / MZB 73-2417 (3 ♂♂, 15 ♀♀), Forat de Sarroqueta, VIII-1966, L. Auroux leg.</p><p>Remarks</p><p>Type series consisting of numerous specimens collected from three localities in the province of Lleida (Fresneda &amp; Escolà, 2000). Specimens of the type series deposited in other collections: CBC, CZULE, FLC, JMLC, MBC, MGC, MNCNM, MNHNP, PMGC and XBC (Fresneda &amp; Escolà, 2000; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775449CB07FF6DF93E7E5AF918	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775449CB06FF6DFA207BB8FCDB.text	D42CA7775449CB06FF6DFA207BB8FCDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stygiophyes hansferyi Fresneda & Escola 2001	<div><p>Stygiophyes hansferyi Fresneda &amp; Escolà, 2001 (Fig. 68)</p><p>Stygiophyes hansferyi Fresneda &amp; Escolà, 2001: 71; figs. 1–4.</p><p>Current accepted name</p><p>Stygiophyes hansferyi Fresneda &amp; Escolà, 2001</p><p>Type locality</p><p>Santoréns, MSS 018 [original citation: “Huesca, Betesa, MSS 018, CN 230, PK 117”].</p><p>Type material</p><p>Holotype ♂: MZB 2004-0950, type loc., 23-V-1992 / 21-VII-1992 (trap), J. Fresneda leg.</p><p>Paratypes: MZB 2004-0951 (1 ♂), same data as holotype. │ MZB 72-4963 / MZB 72-4967 (3 ♂♂, 2 ♀♀), Cova d’Espluga Fonda (as “ Esplugafonda ”), 16-IV-1967, F. Español, L. Auroux, M. Ubach &amp; O. Escolà leg. │ MZB 72- 4968 (2 remains), Cova de la Carretera, 16-IV-1967, F. Español, L. Auroux, M. Ubach &amp; O. Escolà leg. │ MZB 72-4969 (1 ♂), Cova de la Carretera, 1-VI-1974, L. Auroux leg. │ MZB 2004-0952 (1 ♂), Graller Gran del Sodo, 11-XI-1973, J. Comas leg.</p><p>Remarks</p><p>The collection date indicated by Fresneda &amp; Escolà (2001) of specimen MZB 72-4969 is 1 July 1974, although both on the label and in publications that refer to this specimen (Escolà, 1975 -1976; Fresneda &amp; Escolà, 2000; Fresneda &amp; Salgado, 2016) the date is given as 1 June 1974. The type series consists of 114 specimens and the remains of two more from six localities in the provinces of Huesca and Lleida (Fresneda &amp; Escolà, 2001). Paratypes deposited in other collections: SZC, CZULE, FLC, HFC, MNCNM, MNHNP (Fresneda &amp; Escolà, 2001; Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA7775449CB06FF6DFA207BB8FCDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775448CB06FF6DFE617831F8CE.text	D42CA7775448CB06FF6DFE617831F8CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus espanoli subsp. arlai Zariquiey 1950	<div><p>Troglocharinus españoli arlai Zariquiey, 1950</p><p>Troglocharinus españoli ssp. arlai Zariquiey, 1950: 199; figs. 2d, 4d.</p><p>Current accepted name</p><p>Troglocharinus arlai Zariquiey, 1950: Fresneda et al., 2024: 774.</p><p>Type locality</p><p>Avenc de l’Arlà [original citation: “Prov. de Tarragona: Avenc de l’Arlá, barrio de les Pesses, término municipal de Albinyana, part. judicial de Vendrell”].</p><p>Type material</p><p>Syntypes: MZB 71-2387, MZB 71-2388 (1 ♂, 2 ♀♀), type loc., 1934, F. Español leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Zariquiey (1950) mentions “unos pocos ejemplares” [a few specimens] from the type locality without designating the holotype (all specimens originally deposited in the author’s collection). In the MCNB collection we have found a male and two females without labels identifying them as belonging to the type series. However, the information on the collection labels and the hand-writing (belonging to F. Español and R. Zariquiey), as well as the original presence of this material in Zariquiey’s collection, allows us to attribute this material to the type series. Specifically, the labels do not indicate the collection date; Zariquiey (1950) likewise fails to mention it when citing the material from the type locality (other than in the introduction, in which he indicates that Español visited the caves in 1934). Escolà (1969) mentions these three specimens and dates them from 1934. Subsequently, Lagar (1980) and Escolà (1982) mention a precise date (28 January 1934) for the material collected by Español from Avenc de l’Arlà. This date coincides with that of the collection of material from another cave reported by Zariquiey (1950), Cova de Vallmajor, which is not, however, the type locality for ssp. arlai .</p></div>	https://treatment.plazi.org/id/D42CA7775448CB06FF6DFE617831F8CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775448CB01FF6DFA567F65FD3F.text	D42CA7775448CB01FF6DFA567F65FD3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus espanoli subsp. elongatus Zariquiey 1950	<div><p>Troglocharinus españoli elongatus Zariquiey, 1950</p><p>Troglocharinus españoli ssp. elongatus Zariquiey, 1950: 200; figs. 1c, 2e, 4e.</p><p>Current accepted name</p><p>Troglocharinus elongatus elongatus Zariquiey, 1950: Fresneda &amp; Salgado, 2016: 188.</p><p>Type locality</p><p>Cova del Mandil [original citation: “Prov. de Tarragona: Cova del Mandil, término municipal de Querol part. Judicial de Montblanch”].</p><p>Type material</p><p>Syntypes: MZB 71-2389 / MZB 71-2400 (8 ♂♂, 9 ♀♀), type loc., 24-IX-1934, F. Español leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1950) neither specifies the number of specimens in the type series nor establishes a holotype. He indicates that the “Tipos” are deposited in his own personal collection. In the MCNB collection we have found eight males and nine females, without labels indicating that they belong to the type series, which correspond to the 17 specimens cited by Escolà (1969). The collection data (mostly hand-written by Zariquiey), well as the original deposit of the material in his collection, nevertheless, allow us to attribute them to the type series.</p></div>	https://treatment.plazi.org/id/D42CA7775448CB01FF6DFA567F65FD3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544FCB00FF6DFE457A16FEAB.text	D42CA777544FCB00FF6DFE457A16FEAB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus espanoli subsp. espanoli Zariquiey 1950	<div><p>Troglocharinus españoli españoli Zariquiey, 1950 (Figs. 22A–B, 69)</p><p>Troglocharinus españoli ssp. españoli Zariquiey, 1950: 195; figs. 1a, 2a, 3a,b, 4a, 5a.</p><p>Current accepted name</p><p>Troglocharinus elongatus elongatus Zariquiey, 1950: Fresneda &amp; Salgado, 2016: 188.</p><p>Type locality</p><p>Avenc d’Ancosa [original citation: “Prov. de Barcelona: Avenc d’Ancosa, situado en la Plana d’Ancosa, entre La Llacuna y Pontons”].</p><p>Type material</p><p>Syntypes: MZB 71-2415 / MZB 71-2437 (4 ♂♂, 19 ♀♀), MZB 72-9786 / MZB 72-9795 (10 ♀♀), MZB 78-2041 / MZB 78-2056 (16 ♂♂), MZB 78-2919 (1 ♀), type loc., IX-1934, F. Español leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1950) cites “numerosos ejemplares” [many specimens] from the type locality but without stating the exact number of specimens in the type series or establishing the holotype (all were originally deposited in the author’s collection). Subsequently, Zariquiey refers to having examined 25 males and 69 females. Escolà (1969) refers to 18 specimens with the same data as mentioned by Zariquiey (1950). In the MCNB we have located a series of 20 males from Zariquiey’s collection, with the same collection dates as the original description (hand-written by Zariquiey). Only some of these specimens have a label numbered by Zariquiey: specimens MZB 78-2041 to MZB 78-2056 also have another label with the symbol ♂ followed by a correlative number from 1 to 17. We have also found 30 females of the same origin. The type and the inscription on the collection label are identical in all the males in the series and in the majority of females (with a just few variations in some cases). The collection label associated with the females has the date 16 September 1934; Zariquiey (1950), however, does not state the exact day. Only two of these specimens are identified as being part of the type series: the male MZB 78-2041 has a red label with the inscription “ Holotipo ” (possibly hand-written by Lagar) and the numbered labels “16” (hand-written by Zariquiey) and “ ♂ 1”; one of the females, MZB 78-2919, has a similar red label with the inscription “ Alotipo ” in the same hand-writing. It is worth pointing out that Fresneda &amp; Salgado (2016) refer to 18 specimens in this type series, with five “ paratipos ” in the FLC (J. Fresneda, pers. comm.). These five specimens probably correspond to those that, according to Lagar (1980), were lent by Zariquiey. In the MNHNP general collection we have located at least two specimens with red labels inscribed “ Paratipo ” that seem to have been written by the same person as wrote the “ Holotipo ” and “ Alotipo ” on the labels mentioned above.</p><p>Zariquiey (1950) provides the morphometric size of the antennal segments of the male numbered “16” in this series, which corresponds to the number on the label written by Zariquiey associated with specimen MZB 78-2041. For this reason, it is likely that this specimen was chosen and labelled as the “ holotype ”. Unfortunately, this specimen is missing both its pronotum and head. The labels with the symbol ♂ followed by a number seem to correspond to the measurements made by O. Escolà (1966, 1969, 1983; Escolà &amp; Cuadras, 1977) of various species of Bathysciinae.</p></div>	https://treatment.plazi.org/id/D42CA777544FCB00FF6DFE457A16FEAB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544ECB00FF6DFCAE794FFB07.text	D42CA777544ECB00FF6DFCAE794FFB07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus espanoli subsp. mateui Zariquiey 1950	<div><p>Troglocharinus españoli mateui Zariquiey, 1950</p><p>Troglocharinus españoli ssp. mateui Zariquiey, 1950: 200; figs. 2f, 4f.</p><p>Current accepted name</p><p>Troglocharinus elongatus mateui Zariquiey, 1950: Fresneda &amp; Salgado, 2016: 189.</p><p>Type locality</p><p>Cova del Garrofet [original citation: “Prov. de Tarragona: Cova del Garrofet, Santa Agnés, term. municipal de Querol part. Judicial de Montblanch”].</p><p>Type material</p><p>Syntypes: MZB 71-2367 / MZB 71-2379 (10 ♂♂, 17 ♀♀), type loc., VIII-1934, F. Español leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1950) refers to “bastantes ejemplares” [quite a few specimens] from the type locality without specifying the number of specimens in the type series nor establishing the holotype (all originally deposited in the author’s collection). Escolà (1969) mentions a total of 37 specimens with the same collection dates as given by Zariquiey (1950). In the MCNB collection we have found a series of 10 males and 17 females without labels that would identify them as belonging to the type series. The collection dates on their labels, the hand-writing (mostly in the hands of F. Español and R. Zariquiey), and their original deposit in Zariquiey’s collection, nevertheless, indicates that they belong to the type series.</p></div>	https://treatment.plazi.org/id/D42CA777544ECB00FF6DFCAE794FFB07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544ECB00FF6DF81D7FB4F837.text	D42CA777544ECB00FF6DF81D7FB4F837.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus espanoli subsp. olerdolai Lagar 1952	<div><p>Troglocharinus españoli olerdolai Lagar, 1952 (Fig. 70)</p><p>Troglocharinus españoli ssp. olerdolai Lagar, 1952: 79; figs. 2a, 3a, 4b.</p><p>Current accepted name</p><p>Troglocharinus elongatus olerdolai Lagar, 1952: Fresneda et al., 2024: 775.</p><p>Type locality</p><p>Avenc d’Olèrdola [original citation: “Prov. de Barcelona:Avenc d’Olérdola, situado junto al Castillo de Sant Miquel, término municipal de Olérdola, part. judicial de Villafranca del Panadés”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2039, type loc., 11-II-1951, F. Español &amp; A. Lagar leg.</p><p>Paratype (Allotype) ♀: MZB 78-2040, same data as holotype .</p><p>Remarks</p><p>Lagar (1952) wrote of the type series: “Tipos del Avenc d’Olérdola en el Museo de Ciencias Naturales de Barcelona; un paratipo hembra, en mi colección” [Types from Avenc d’Olérdola in the MCNB; one female paratype in my collection]. We believe that the author used the expression “Tipos” to designate the ♂ holotype and ♀ allotype, as their respective labels suggest. The two types deposited in the MCNB are referred to by Escolà (1969). The type series is completed by the ♀ paratype, currently deposited in the FLC (Fresneda &amp; Salgado, 2016).</p></div>	https://treatment.plazi.org/id/D42CA777544ECB00FF6DF81D7FB4F837	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544DCB03FF6DFD907A8FFC0A.text	D42CA777544DCB03FF6DFD907A8FFC0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus espanoli subsp. ollai Zariquiey 1950	<div><p>Troglocharinus españoli ollai Zariquiey, 1950</p><p>Troglocharinus españoli ssp. ollai Zariquiey, 1950: 199; figs. 2c, 4c.</p><p>Current accepted name</p><p>Troglocharinus elongatus ollai Zariquiey, 1950: Fresneda &amp; Salgado, 2016: 189.</p><p>Type locality</p><p>Cova de l’Olla [original citation: “Prov. de Tarragona: Cova de l’Olla, en Cal Sumoy, term. municipal de Montmell, Part. judicial de Vendrell”].</p><p>Type material</p><p>Syntypes: MZB 71-2380 / MZB 71-2386 (7 ♂♂, 10 ♀♀), type loc., 27-VIII-1934, F. Español leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1950) mentions “varios ejemplares” [a number of specimens] but does not indicate how many specimens there are in the type series nor establishes a holotype (all were originally deposited in his personal collection). Escolà (1969) cites a total of 15 specimens with the collection dates given by Zariquiey (1950). In the MCNB collection we have found a series of seven males and ten females without labels identifying them as belonging to the type series. The information on the collection labels and its hand-writing (mostly in the hand of F. Español), as well as the original placement of this material in Zariquiey’s collection, enables us, nevertheless, to attribute these specimens to the type series.</p></div>	https://treatment.plazi.org/id/D42CA777544DCB03FF6DFD907A8FFC0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544DCB03FF6DF912794DF934.text	D42CA777544DCB03FF6DF912794DF934.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus espanoli subsp. pinyareti Zariquiey 1950	<div><p>Troglocharinus españoli pinyareti Zariquiey, 1950 (Fig. 71)</p><p>Troglocharinus españoli ssp. pinyareti Zariquiey, 1950: 201; figs. 2g, 4g, 5c.</p><p>Current accepted name</p><p>Troglocharinus pinyareti Zariquiey, 1950: Fresneda et al., 2024: 776.</p><p>Type locality</p><p>Avenc dels Pinyerets [original citation: “Prov. de Tarragona: avenc de Pinyarets, term. municipal de Montmell part. Judicial de Vendrell”].</p><p>Type material</p><p>Syntypes: MZB 71-2401 / MZB 71-2403 (3 ♂♂, 1 ♀), type loc., 26-VIII-1934, F. Español leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1950) did not establish a holotype, merely indicating that the “Tipos” were deposited in his collection (he then refers to six specimens from the type locality). In the MCNB collection we have found three males and a female. The collection dates on their labels and the hand-writing (mostly in the hand of F. Español), as well as their original deposit in Zariquiey’s collection, nevertheless, indicates that they belong to the type series. Escolà (1969) refers to three of these specimens.</p></div>	https://treatment.plazi.org/id/D42CA777544DCB03FF6DF912794DF934	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544DCB02FF6DFA4B7999FD02.text	D42CA777544DCB02FF6DFA4B7999FD02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus espanoli subsp. portai Zariquiey 1950	<div><p>Troglocharinus españoli portai Zariquiey, 1950</p><p>Troglocharinus españoli ssp. portai Zariquiey, 1950: 197; fig. 5b.</p><p>Current accepted name</p><p>Troglocharinus elongatus portai Zariquiey, 1950: Fresneda &amp; Salgado, 2016: 190.</p><p>Type locality</p><p>Cova del Bolet [original citation: “Prov. de Barcelona: Cova del Bolet, term. municipal de San Quintín de Mediona, part. de Villafranca”].</p><p>Type material</p><p>Syntypes: MZB 71-2404 (1 ♀), type loc., 24-XII-1934, F. Español leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1950) did not establish a holotype and instead indicates that the “Tipos” are deposited in his collection (he refers to a male and a female from the type locality). In the MCNB collection (Zariquiey’s collection) we have located a female with the collection dates given by Zariquiey (1950), with labels written in his own hand. This specimen, moreover, has a red label with the inscription “Typus” printed in black letters. We were unable to locate the male. Escolà (1969) refers to both specimens.</p></div>	https://treatment.plazi.org/id/D42CA777544DCB02FF6DFA4B7999FD02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA777544CCB02FF6DFE197AE2F87D.text	D42CA777544CCB02FF6DFE197AE2F87D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus espanoli subsp. roselli Lagar 1952	<div><p>Troglocharinus españoli roselli Lagar, 1952 (Fig. 72)</p><p>Troglocharinus españoli ssp. roselli Lagar, 1952: 77; figs. 1, 2c, 3b.</p><p>Current accepted name</p><p>Troglocharinus elongatus roselli Lagar, 1952: Fresneda et al., 2024: 775.</p><p>Type locality</p><p>Avenc de Foix [original citation: “Prov. de Barcelona: Avenc de Foix, situado cerca de la antigua iglesia de Foix, término municipal de Torrellas de Foix, part. judicial de Villafranca del Panadés”].</p><p>Type material</p><p>Syntypes: MZB 78-2033 / MZB 78-2035, MZB 78-2037, MZB 78-2038 (4 ♂♂, 8 ♀♀), type loc., 30-XII-1951, A. Lagar &amp; J. Rosell leg. │ MZB 78-2036 (1 ♂), type loc., 3-II-1952, A. Lagar &amp; J. Rosell leg. │ MZB 2021-0870, MZB 2021-0871 (1 ♂, 2 ♀♀), same data. (ex coll. X. Bellés) .│ MZB 2023-0483 (1 ♂), same data (ex coll. A. Lagar) .</p><p>Remarks</p><p>Lagar (1952) indicates the deposit of the “Tipos” in his private collection, reporting the collection of 24 specimens from the type locality on 30 December 1951. Ten specimens of the type series are deposited in the FLC (Fresneda &amp; Salgado, 2016) and a further 12 have been located in the entomological collection of the MCNB (corresponding to the specimens cited by Escolà, 1969). All these specimens have red hand-written paratype labels, although no holotype was designed in the original publication (Lagar, 1952).</p><p>In the MCNB collection there are deposited five specimens (four of which recently donated, three by X. Bellés and another one from the Lagar collection itself) also labelled as paratypes, with the same collection information except for the date (3 February 1952). In the FLC there are 73 specimens with the same information cited as paratypes by Fresneda &amp; Salgado (2016). Belonging to this series and with a red hand-written label with the indication “ Paratipo ” we have located a single male in the MNCNM collection, as well as two other specimens deposited in the CJC (J. Comas, pers. comm.). According to J. Fresneda (pers. comm.), before the publication of the description only specimens from the two dates indicated existed. All the specimens dated 3 February 1952 that we have located bear paratype labels, most are deposited in the FLC and those from the JCC and XBC were donated to them by Lagar himself; all of which indicates that they probably form part of the type series, despite not being cited as such in the description .</p></div>	https://treatment.plazi.org/id/D42CA777544CCB02FF6DFE197AE2F87D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775443CB0DFF6DFD9078D0F9E6.text	D42CA7775443CB0DFF6DFD9078D0F9E6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus fadriquei Fresneda, Rizzo, Comas & Ribera 2021	<div><p>Troglocharinus fadriquei Fresneda, Rizzo, Comas &amp; Ribera, 2021</p><p>Troglocharinus fadriquei Fresneda, Rizzo, Comas &amp; Ribera, 2021: 166; figs. 4, 8, 12, 16, 21, 26, 29, 30, 43, 48, 53, 59.</p><p>Current accepted name</p><p>Troglocharinus fadriquei Fresneda, Rizzo, Comas &amp; Ribera, 2021</p><p>Type locality</p><p>Forat de les Casanoves [original citation: “ Spain, Lleida, Figols d’Organyà, Forat de Casanova, UTM (ETRS89) 31N 3637 46744, 905 m ”] .</p><p>Type material</p><p>Holotype ♂: MZB 2021-0039, type loc., 22-VIII-1992, M. Escoll &amp; J. Fresneda leg.</p><p>Paratypes: MZB 70-1388 / MZB 70-1393 (12 ♂♂, 10 ♀♀), type loc., 2-V-1982, Escolà, Lista &amp; Hernández leg. │ MZB 78-1094 (1 ♂), type loc., 25-I-1970, C. Ribera &amp; Mª D. Romero leg. │ MZB 78-1095, MZB 78-1096 (1 ♂, 1 ♀), type loc., 12-III-1972, J. Bley &amp; L. Auroux leg. │ MZB 78-1097 / MZB 78-1103 (3 ♂♂, 10 ♀♀, 7 remains), type loc., 27-II-1972, J. Bley &amp; A. Meseguer leg. │ MZB 2021-0875, MZB 2021-0876 (1 ♂, 1 ♀), type loc., 23- XII-2018, R. Sendra &amp; J. Pallisé leg. (ex coll. X. Bellés) .</p><p>Remarks</p><p>The paratypes with register numbers MZB 78-1094 / MZB 78-1103 were included by Escolà (1971) as part of the type series of Speonomus vinyasi, and subsequently designated by Fresneda et al. (2021) as paralectotypes of such taxon (see remarks for Speonomus vinyasi Escolà 1971).</p><p>Fresneda et al. (2021), designate 42 specimens from “Forat de Casa Nova ”, dated 2 May 1982 (Escolà, Lista &amp; Hernández leg.), all of them deposited in the MCNB collection, according to the authors (without indicating the register numbers). However, only 22 specimens have been located with these data (12 ♂♂ and 10 ♀♀, with register numbers MZB 70-1388 / MZB 70-1393), placed on together in the same entomological box, and originally attributed to Speonomus vinyasi Escolà, 1971 . This discrepancy could be explained taking into account the information we have obtained from the record card of the Escolà’s data file. The entry associated with the above-mentioned date and collectors, indicates 14 specimens and 7 remains (in alcohol) collected by Escolà, as well as 28 more specimens collected by Lista-Hernández (just 42 specimens in all, if we exclude the remains). Sixteen of those specimens were sent two days later for cytological analysis, with negative results. Escolà then refers the preparation during the three days following the collection date of 11 ♂♂ and 13 ♀♀, as well as 2 ♂♂ of those sent to be analyzed, without specifying a date. The whereabouts of the rest of the specimens analyzed are not indicated, but from this circumstance, as well as the documented number of specimens prepared (similar to the specimens we have located), it is deduced that either they were not returned to the MCNB collection or they were lost (destructive analysis?) shortly after their collection.</p><p>Type series of T. fadriquei consisting of 227 specimens; paratypes deposited in other collections (Fresneda et al., 2021): AFC, CBC, DVC, FLC, IBE, JCC, MNHNP and PMGC. The paratypes originally deposited in the XBC (Fresneda et al., 2021) have been donated to the MCNB collection .</p></div>	https://treatment.plazi.org/id/D42CA7775443CB0DFF6DFD9078D0F9E6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775443CB0CFF6DFB7E78EEFBB2.text	D42CA7775443CB0CFF6DFB7E78EEFBB2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus ferreri subsp. codinai Zariquiey 1917	<div><p>Troglocharinus ferreri codinai Zariquiey, 1917 (Figs. 23A–C, 73)</p><p>Troglocharinus Ferreri ssp. Codinai Zariquiey, 1917: 291; fig. 5–7.</p><p>Current accepted name</p><p>Troglocharinus ferreri ferreri (Reitter, 1908): Fresneda &amp; Salgado, 2016: 192.</p><p>Type locality</p><p>Cova Fosca [original citation: “Cova Fosca de Gavá, en el valle de Joan, cerca de Gavá, partido municipal de San Feliu de Llobregat ”].</p><p>Type material</p><p>Possible syntypes: MZB 71-4742, MZB 78-1357 / MZB 78-1364, MZB 78-1467 / MZB 78-1469 (5 ♂♂, 7 ♀♀), type loc., 30-IX-1917, 14-X-1917, 21-X-1917, A. Guimjuan leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1917) mentions as types two males and two females deposited in his collection but gives no other details. Subsequently, he notes the collection (A. Guimjuan leg.) of 35 specimens on 30 September and 14 and 21 October [1917], without details of where they are deposited.</p><p>Seven specimens in the MCNB collection have red labels printed with the inscription “ Typus ”. A further two specimens are identified as co-types: one has a red label printed with “Cotipus” and the other a hand-written red label stating “CO-TIPO”. Apart from one specimen (MZB 78-1362), all the capture labels of these specimens indicate in the author’s hand-writing one of the three capture dates cited by Zariquiey (1917). We have located three further specimens not identified as belonging to the type series: MZB 71-4742 and MZB 78-1467, dated 14 October 1917 with the rest of the capture label identical to MZB 78-1357 (one with an inscription hand-written by Zariquiey stating “ Troglocharinus Ferreri ssp. Codinai nov”); and MZB 78-1364, with no collection date but with a capture label identical to MZB 78-1362 .</p><p>Escolà (1969) mentions a total of 10 specimens dated 21 October and two from 14 October (we have only been able to find five and three, respectively). He does not mention any of the specimens collected in September (two females, as the author of the description cites in the introduction), which would correspond to specimens MZB 78- 1361 and 78-1468.</p><p>In box 74 of the MNHNP general collection we have located at least three specimens with identical collection labels, but without labels that identify them as belonging to the type series. Jeannel (1924a) indicates textually “… type: cova fosca de Gava (coll. R. Zariquiey et coll. Biospeologica)”. In the MNCNM collection (box 28) we have also located two specimens (dated 14 and 17 October 1917) that lack labels that identify them as belonging to the type series .</p><p>In light of the information provided by Zariquiey (1917), including the introductory details, it is possible that the description was based on the series of 35 specimens mentioned at the end of the description, despite citing explicitly as “Tipos” only two undated males and two undated females.</p></div>	https://treatment.plazi.org/id/D42CA7775443CB0CFF6DFB7E78EEFBB2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775442CB0CFF6DF9C97A9BF835.text	D42CA7775442CB0CFF6DF9C97A9BF835.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus ferreri subsp. fonti Zariquiey 1924	<div><p>Troglocharinus ferreri fonti Zariquiey, 1924</p><p>Troglocharinus Ferreri ssp. Fonti Zariquiey, 1924: 17; fig. 9d.</p><p>Current accepted name</p><p>Troglocharinus ferreri ferreri (Reitter, 1908): Fresneda &amp; Salgado, 2016: 192.</p><p>Type locality</p><p>Avenc del Vermell [original citation: “Avenc del Vermell ... Alrededores de Begues, Prov. Barcelona”].</p><p>Type material</p><p>Syntypes: MZB 78-1330 / MZB 78-1347, MZB 78-1925 (4 ♂♂, 15 ♀♀), type loc., 16-XII-1922 (trap), R. Zariquiey leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>Zariquiey (1924) cites the capture of a number of specimens but does not mention exactly how many. In the MCNB collection we have found 19 specimens with red labels printed with “ Typus ”, the exceptions being MZB 78-1345 and MZB 78-1925, which have red labels with the hand-written inscriptions “TIPO” and “COTIPO”, respectively. Escolà (1969) reports the same number of specimens. In the MNHNP general collection (box 74) we have located at least four specimens with identical labels of origin that also have a red label inscribed with “ Typus ”. According to Jeannel (1924a), this material is part of the type series deposited in the Biospeologica collection. As well, we have located in the MNCNM collection (box 28) a specimen with identical information and a red hand-written label inscribed with “COTIPO”.</p></div>	https://treatment.plazi.org/id/D42CA7775442CB0CFF6DF9C97A9BF835	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775441CB0FFF6DFD907E5AFBFE.text	D42CA7775441CB0FFF6DFD907E5AFBFE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus ferreri subsp. jeanneli Zariquiey 1917	<div><p>Troglocharinus ferreri jeanneli Zariquiey, 1917</p><p>Troglocharinus Ferreri ssp. Jeanneli Zariquiey, 1917: 289; fig. 3, 4.</p><p>Current accepted name</p><p>Troglocharinus ferreri ferreri (Reitter, 1908): Fresneda &amp; Salgado, 2016: 192.</p><p>Type locality</p><p>Cova de la Fou de Montaner [original citation: “cueva de la Fou Montaner, cerca de Vallirana, partido de San Feliu de Llobregat ”].</p><p>Type material</p><p>Syntypes: MZB 78-1348, MZB 78-1349, MZB 78-1352 / MZB 78-1355 (3 ♂♂, 3 ♀♀), type loc., 28-X-1917, R. Zariquiey leg. (ex coll. R. Zariquiey).</p><p>Remarks</p><p>Zariquiey (1917) mentions as types a number of specimens but notes subsequently that he collected 13 specimens on 28 October [1917]. He also adds that Jeannel (along with E. G. Racovitza) collected this taxon for the first time in a cave in October 1910. This material is cited by Jeannel (1910b, 1911b, 1924a) and is deposited in the Biospeologica collection (MNHNP) (Fresneda &amp; Salgado, 2016). In MCNB we have found red printed labels inscribed with “ Typus ” (four specimens) and “Cotypus” (one specimen). We have also found a specimen (MZB 78-1355) with an identical collection label in Zariquiey’s hand-writing that must belong to the type series, although it is not labelled as such. Likewise, we have located in the MNCNM collection (box 28) another specimen with the same collection information. Escolà (1969) mentions a total of 11 specimens whose information corresponds to that of the type series.Another specimen in the MCNB collection (MZB 78-1356), collected by Jeannel in October 1910, according to the label, corresponds to the first specimens that were collected (Jeannel, 1910b; Zariquiey, 1917).</p></div>	https://treatment.plazi.org/id/D42CA7775441CB0FFF6DFD907E5AFBFE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775441CB0FFF6DF9737825F837.text	D42CA7775441CB0FFF6DF9737825F837.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus ferreri subsp. pallaresi Belles 1973	<div><p>Troglocharinus ferreri pallaresi Bellés, 1973</p><p>Troglocharinus ferreri ssp. pallaresi Bellés, 1973: 45; figs. 1, 2.</p><p>Current accepted name</p><p>Troglocharinus ferreri pallaresi Bellés, 1973</p><p>Type locality</p><p>Avenc de Can Montmany [original citation: “Avenc de Can Montmany, próximo a La Palma de Cervelló, término municipal de Cervelló, provincia de Barcelona”].</p><p>Type material</p><p>Holotype ♂: MZB 2021-0859, type loc., 23-XII-1972, J. Pallarès &amp; X. Bellés leg. (ex coll. X. Bellés).</p><p>Paratypes: MZB 2021-0860 / MZB 2021-0867 (3 ♂♂, 5 ♀♀), same data as holotype (ex coll. X. Bellés) │ MZB 78-1928, MZB 78-1929 (4 ♂♂, 4 ♀♀), same data as holotype. │ MZB 78-1470 / MZB 78-1474 (4 ♂♂, 3 ♀♀), type loc., 24-X-1971, J. Bley leg. │ MZB 78-1475 / MZB 78-1478 (2 ♂♂, 4 ♀♀), Cova de la Carretera, 14-I-1973, L. Auroux, J. Bley &amp; O. Escolà leg. │ MZB 2021-0869 (2 ♀♀), Cova de la Carretera, 14-I-1973, L. Auroux, J. Bley &amp; O. Escolà leg. (ex coll. X. Bellés) │ MZB 78-1479, MZB 78-1480, MZB 78-1926, MZB 78-1927 (2 ♂♂, 3 ♀♀), Avenc de la Cabra, 14-I-1973, L. Auroux, J. Bley &amp; O. Escolà leg. │ MZB 2021-0868 (1 ♂), Avenc de la Cabra, 23-XII-1972, X. Bellés leg. (ex coll. X. Bellés) .</p><p>Remarks</p><p>The type series consists of a ♂ holotype and 43 paratypes (Bellés, 1973) from three cavities in the province of Barcelona; where they are deposited is not indicated. From the Cova de la Carretera, the author cites 1♂ and 7 ♀♀ (of the eight paratypes from this locality deposited in the MCNB, two are males). The holotype and 11 paratypes deposited in the XBC have been donated to the MCNB collection .</p></div>	https://treatment.plazi.org/id/D42CA7775441CB0FFF6DF9737825F837	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775440CB0EFF6DFD907F1CFCFA.text	D42CA7775440CB0EFF6DFD907F1CFCFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus ferreri subsp. zariquieyi Jeannel 1924	<div><p>Troglocharinus ferreri zariquieyi Jeannel, 1924 (Figs. 24, 74)</p><p>Troglocharinus Ferreri ssp. Zariquieyi Jeannel, 1924b: 17 .</p><p>Current accepted name</p><p>Troglocharinus ferreri ferreri (Reitter, 1908): Fresneda &amp; Salgado, 2016: 192.</p><p>Type locality</p><p>Avenc de la Funiosa [original citation: “avench de la Funiosa, dans le massif du Panadés”].</p><p>Type material</p><p>Syntypes: MZB 78-1350, MZB 78-1351 (2 ♀♀), type loc., R. Zariquiey leg. (ex coll. R. Zariquiey) .</p><p>Remarks</p><p>The type series consists of three females (Jeannel, 1924b), two of which are deposited in the MCNB collection and a third in the Biospeologica collection (MNHNP) (Jeannel, 1924a). The cave, in the municipality of Gavà (Barcelona province), currently lies buried under the Garraf rubbish dump (Fresneda &amp; Salgado, 2016). In the MNHNP’s general collection (box 74) we have located the third specimen that completes the type series (identical collection labels, as well as a label with “ TYPE ” written in red letters on a white background).</p></div>	https://treatment.plazi.org/id/D42CA7775440CB0EFF6DFD907F1CFCFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775440CB0EFF6DFE827F94F81C.text	D42CA7775440CB0EFF6DFE827F94F81C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus hustachei Jeannel 1911	<div><p>Troglocharinus hustachei Jeannel, 1911 (Figs. 25A–B)</p><p>Troglocharinus Hustachei Jeannel, 1911a: XCVI; fig. 17.</p><p>Current accepted name</p><p>Troglocharinus hustachei Jeannel, 1911</p><p>Type locality</p><p>Cova del Gel [original citation: “Cova del Gel, située près du sommet de la sierra de Montsech, districto mun. de Llimiana, partido de Tremp, provincia de Lérida (Catalogne)”] .</p><p>Type material</p><p>Syntypes: MZB 78-1930 (1 ♂), type loc., 19-VIII-1910, M. A. Hustache leg. (ex coll. R. Zariquiey).</p><p>Possible syntypes: MZB 78-1931 (1 ♀), same data as MZB 78-1930 .</p><p>Remarks</p><p>Jeannel (1911a) cites an unspecified number of specimens collected by Hustache in August 1910, from which he selected three type specimens (19 August 1910), which were originally deposited in the MNHNP biospeleological collection. The author does not state the sex of the specimens. In Zariquiey’s collection we have located an incomplete male (MZB 78-1930) with the same collection dates and a white label inscribed with “ TYPE ” in red letters. In the MNHNP general collection (box 74) we have found two specimens, each with labels printed with “ TYPE ”, one in black letters on a red background, the other red letters on a white background, identical to the label of MZB 78- 1930. The information on and format of the two hand-written collection labels of these three specimens are identical and so suggest a common origin. Zariquiey (1917) cites a female captured by Hustache deposited in his collection that, given the information on the capture label, must correspond to the female with accession number MZB 78- 1931 found with MZB 78-1930. This specimen seems to correspond to the series collected by Hustache referred to by Jeannel (1911a). It is possible that the description of this new taxon was based on the study of more specimens and not simply on the examination of the three specimens designated as types, which we believe correspond to the three specimens with the labels inscribed with “ TYPE ” (MZB 78-1930 and the two from the MNHNP). Jeannel (1911a), in fact, uses the expressions “des mâles…” [males] and “Chez les femelles…” [in the females], both in the plural, and so it is possible that specimen MZB 78-1931 was indeed part of the studied material.</p></div>	https://treatment.plazi.org/id/D42CA7775440CB0EFF6DFE827F94F81C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775447CB09FF6DFD9079E7FBFF.text	D42CA7775447CB09FF6DFD9079E7FBFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus hustachei subsp. impellitierii Espanol 1955	<div><p>Troglocharinus hustachei impellitierii Español, 1955 (Fig. 75)</p><p>Troglocharinus hustachei ssp. impellitierii Español, 1955: 270; pl. VIII, fig. 5b, 6b.</p><p>Current accepted name</p><p>Troglocharinus impellitierii Español, 1955: Español, 1966b: 21.</p><p>Type locality</p><p>Cova Palomera [original citation: “ Cueva “La Palomera” (Tahús) ... abierta a unos 5 metros sobre el torrente de Tahús, junto a su confluencia con el de Castells y a unos 500 m de este último pueblo”].</p><p>Type material</p><p>Syntypes: MZB 78-1932 / MZB 78-1942 (7 ♂♂, 6 ♀♀), type loc., 1-X-1950, F. Español leg.</p><p>Remarks</p><p>Español (1955) does not establish any holotype but mentions “un cierto número de ejemplares (♂♂ y ♀♀)” [a certain number of specimens (♂♂ and ♀♀)] without providing any more details. In the MCNB collection we have found 13 specimens. All have red, hand-written labels with the inscription “Serie típica”. Specimen ♂ MZB 78- 1932 was designated incorrectly by Escolà (2000) as a holotype (article 73.1.3.); this author added a red label on which he wrote “Tipus / Troglocharinus / impellitierii sp. n. ♂ / ESPAÑOL det. 52”. On the collection labels hand-written by Español appears the date October 1952. Español (1953b) cites this new subspecies in the section “Índice cronológico de los trabajos publicados” [Chronological index of published works] (1952), in which he mentions an on-going study of the corresponding material and gives its title and the journal of publication. The name becomes available with the publication of the description (article 13.1.1.). The 13 specimens belonging to the type series seem to be those that Escolà (1969) mentions with a collection date of 1 October 1955 (the collection campaigns that Español (1955) refers to took place in 1950).</p></div>	https://treatment.plazi.org/id/D42CA7775447CB09FF6DFD9079E7FBFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775447CB09FF6DF9867F5FF8D1.text	D42CA7775447CB09FF6DF9867F5FF8D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus pallisei Rizzo & Comas 2015	<div><p>Troglocharinus pallisei Rizzo &amp; Comas, 2015</p><p>Troglocharinus pallisei Rizzo &amp; Comas, 2015: 107; figs. 2-9.</p><p>Current accepted name</p><p>Troglocharinus pallisei Rizzo &amp; Comas, 2015</p><p>Type locality</p><p>Cova de la Riba [original citation: “COUNTRY: Spain: Province: Tarragona: ♂, labelled “ Cv. de la Riba o / del Talc - La Riba- / (Alt Camp) II-2012 / R. Sendra leg.; X 34895.7 / Y 4575816.3 / Z 240 / ETRS89”] .</p><p>Type material</p><p>Paratypes: MZB 74-3360 / MZB 74-3363 (3 ♂♂, 1 ♀), type loc., 17-VII-1971, J. Pallisé leg.</p><p>Remarks</p><p>Type series consisting of 12 ♂♂ and 17 ♀♀ (Rizzo &amp; Comas, 2015). On the capture label of specimen MZB 74- 3363 the collection date is given erroneously as “17-I-71”; this specimen shows some disarticulated appendages (“damaged”, according to the authors of the description). Specimens of the type series deposited in other collections: ♂ holotype in the JCC; paratypes in the IBE and JCC (Rizzo &amp; Comas, 2015).</p></div>	https://treatment.plazi.org/id/D42CA7775447CB09FF6DF9867F5FF8D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775447CB08FF6DFA777A53FD02.text	D42CA7775447CB08FF6DFA777A53FD02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus sendrai Fresneda, Rizzo, Comas & Ribera 2021	<div><p>Troglocharinus sendrai Fresneda, Rizzo, Comas &amp; Ribera, 2021</p><p>Troglocharinus sendrai Fresneda, Rizzo, Comas &amp; Ribera, 2021: 165; figs. 3, 7, 15, 20, 27, 28, 42, 52, 58.</p><p>Current accepted name</p><p>Troglocharinus sendrai Fresneda, Rizzo, Comas &amp; Ribera, 2021</p><p>Type locality</p><p>Mina de Coll d’Arnat [original citation: “ Spain, Lleida, La Vansa i Fórnols, Mina de Coll d’Arnat, UTM (ETRS 89) 31N 3690 46791, 1342 m ”].</p><p>Type material</p><p>Holotype ♂: MZB 2021-0038, type loc., 16-X-2015, V. Rizzo leg.</p><p>Paratypes: MZB 70-1327 / MZB 70-1330 (2 ♂♂, 5 ♀♀), type loc., 19-X-1969, L. Auroux leg. │ MZB 2021-0873, MZB 2021-0874 (1 ♂, 1 ♀), type loc., 12-XI-2016 / 11-V-2017, J. Comas leg. (ex coll. X. Bellés, ex coll. J. Comas) │ MZB 70-1331 (1 ♂), Baborell de Montsec, 26-X-1969, L. Auroux leg.</p><p>Remarks</p><p>Type series consisting of 27 ♂♂ and 40 ♀♀; paratypes deposited in other collections: FLC, IBE, JCC (Fresneda et al., 2021) .</p></div>	https://treatment.plazi.org/id/D42CA7775447CB08FF6DFA777A53FD02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775446CB08FF6DFE197A38F8A5.text	D42CA7775446CB08FF6DFE197A38F8A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus senenti Escola 1967	<div><p>Troglocharinus senenti Escolà, 1967 (Fig. 76)</p><p>Troglocharinus senenti Escolà, 1967: 85 .</p><p>Current accepted name</p><p>Troglocharinus senenti Escolà, 1967</p><p>Type locality</p><p>Querant Gran de Paús [original citation: “«Querant Gran de Paús», Vilanova de Meià, Montsec de Rubíes, Lérida”].</p><p>Type material</p><p>Holotype ♂: MZB 78-1943, type loc., 1-I-1964, J. Senent &amp; O.Escolà leg.</p><p>Paratypes: MZB 75-9326, MZB 75-9327, MZB 78-1944 / MZB 78-1969, MZB 78-2025 / MZB 78-2032 (45 ♂♂, 40 ♀♀, 1 remain), same data as holotype. │ MZB 2021-0872 (2 ♂♂), same data as holotype (ex coll. X. Bellés) .</p><p>Remarks</p><p>In the original description, Escolà (1967) designates a holotype ♂ and 50 paratypes (Senent &amp; Escolà leg.). Subsequently, Escolà (2000) modified this data regarding the composition of the type series, and indicates instead a total of 93 paratypes (40 ♂♂ and 51 ♀♀), all deposited in the MCNB except for two specimens in other museums (without specifying which) used as part of an exchange. It is likely that the two specimens that Escolà (2000) refers to never in fact left the Museum and correspond to specimens MZB 75-9326 and MZB 75-9327 (1 ♂ and 1 ♀), separated from the rest of the material, which were recently found in one of Escolà’s entomological boxes labelled as material to be used in exchanges. Fresneda &amp; Salgado (2016) indicate that a part (without specifying how many) of the type series mentioned by Escolà (1967) is deposited in the Canció collection. On the other hand, two males from the XBC have recently been donated to the MCNB collection.</p><p>In all, 87 specimens (47 ♂♂ and 40 ♀♀) are labelled as paratypes in the MCNB collection, a total that includes more males than those cited by Escolà (2000), as well as the remains of one specimen of unknown sex. Escolà (1969, 1981) cites a total of 104 specimens (52 of each sex), all with the same data as the type series. Thus, given the available information, we believe that it is justifiable to regard all these specimens as part of the type series (article 72.4.1.1.).</p></div>	https://treatment.plazi.org/id/D42CA7775446CB08FF6DFE197A38F8A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775445CB0BFF6DFD907BB8FCDE.text	D42CA7775445CB0BFF6DFD907BB8FCDE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglocharinus vivesi Comas 2022	<div><p>Troglocharinus vivesi Comas, 2022</p><p>Troglocharinus vivesi Comas, 2022: 143; figs. 1a–b, 2a, 3.</p><p>Current accepted name</p><p>Troglocharinus vivesi Comas, 2022</p><p>Type locality</p><p>Bòfia de la Valldan [original citation: « Bòfia de la Valldan / -la Valldan- Odèn (Solsonès) / 7-VII-2022 / Meseguer &amp; Pallisé leg.» «ETRS89 Fus: 31N / 364350, 4661571 / 905 m. s.n.m»].</p><p>Type material</p><p>Holotype ♂: MZB 2024-0818, type loc., 7-VII-2022, A. Meseguer &amp; J. Pallisé leg.</p><p>Paratypes: MZB 2024-0819 / MZB 2024-0828 (5 ♂♂, 5 ♀♀), same data as holotype .</p><p>Remarks</p><p>Type series consisting of 17 ♂♂ and 22 ♀♀. The rest of the paratypes deposited in the JCC and FLC (Comas, 2022) .</p></div>	https://treatment.plazi.org/id/D42CA7775445CB0BFF6DFD907BB8FCDE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775445CB0BFF6DFE667A59F9A5.text	D42CA7775445CB0BFF6DFE667A59F9A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Troglophyes riberai Espanol 1967	<div><p>Troglophyes riberai Español, 1967 (Fig. 77)</p><p>Troglophyes riberai Español, 1967: 81; figs. 1–4.</p><p>Current accepted name</p><p>Ceretophyes riberai (Español, 1967): Fresneda, 1998: 79.</p><p>Type locality</p><p>Cova d’en Manent [original citation: “Cova Manent, 1.200 m. altitud, a una media hora del pueblo de Olopte, no lejos de Puigcerdá”].</p><p>Type material</p><p>Holotype ♂: MZB 78-2076, type loc., 28-VIII-1966, F. Español, C. Ribera, R. Viñas &amp; I. González leg. Paratypes: MZB 78-2077 / MZB 78-2124 (84 ♂♂, 158 ♀♀), same data as holotype. │ MZB 78-2125 / MZB 78- 2132 (18 ♂♂, 25 ♀♀), type loc., VIII-1966, C. Ribera, R. Viñas, A. Miravitlles, A. Traver &amp; D. Pérez leg.</p><p>Remarks</p><p>Type series consisting of the holotype ♂ (referred to as “Tipo”, article 73.1.1.) and numerous paratypes (with no details) from three caves in the Girona Pyrenees (Español, 1967). We have only been able to find in the MCNB collection the specimens from the type locality (holotype and paratypes; a total of 103 ♂♂ and 183 ♀♀). Nevertheless, Español (1967) mentions paratypes from “Cova de Olopte n.º 1” and “Cova de Olopte n.º 4”, also deposited in the Museum.</p></div>	https://treatment.plazi.org/id/D42CA7775445CB0BFF6DFE667A59F9A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775445CB0AFF6DFBBB7904FEAA.text	D42CA7775445CB0AFF6DFBBB7904FEAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Typhlochlamys bardisai Espanol 1975	<div><p>Typhlochlamys bardisai Español, 1975 (Fig. 78)</p><p>Typhlochlamys bardisai Español, 1975: 123; figs. 1, 2.</p><p>Current accepted name</p><p>Spelaeochlamys bardisai (Español, 1975): Salgado &amp; Fresneda, 2004a: 379.</p><p>Type locality</p><p>Avenc de Barratxina [original citation: “Cova-avenc Barratxina, Jijona, Alicante”].</p><p>Type material</p><p>Holotype ♂: MZB 78-0869, type loc., V-1969, F. Bardisa leg.</p><p>Paratypes: MZB 78-0852 (1 ♀), same data as holotype .</p><p>Remarks</p><p>Full type series deposited in the MCNB.</p></div>	https://treatment.plazi.org/id/D42CA7775445CB0AFF6DFBBB7904FEAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
D42CA7775444CB0AFF6DFCB17893FB55.text	D42CA7775444CB0AFF6DFCB17893FB55.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Typhlochlamys escolai Comas 1978	<div><p>Typhlochlamys escolai Comas, 1978 (Fig. 79)</p><p>Typhlochlamys escolai Comas, 1978b: 161; figs. 1–3.</p><p>Current accepted name</p><p>Spelaeochlamys bardisai (Español, 1975): Salgado &amp; Fresneda, 2004a: 379.</p><p>Type locality</p><p>Avenc del Caldero [original citation: “Avenc del Caldero, Tibi, Alicante”].</p><p>Type material</p><p>Holotype ♂: MZB 78-0853, type loc., 30-X-1977, O. Escolà leg.</p><p>Paratypes: MZB 78-0854 / MZB 78-0855, MZB 78-1914 / MZB 78-1915 (5 ♂♂, 8 ♀♀), same data as holotype .</p><p>Remarks</p><p>The type series referred to by Comas (1978b) consists of the ♂ holotype and 13 paratypes (6 ♂♂ and 7 ♀♀), the latter shared between the MCNB and JCC collections. Nevertheless, there are 5 ♂♂ and 8 ♀♀ deposited in the MCNB that are labelled as paratypes. In the JCC collection there is 1 ♂ and 2 ♀♀ (J. Comas, pers. comm.), which means that the total number of female paratypes exceeds the number indicated in the description. Given the type of labels and the information they contain, we believe that the whole series was studied when making the original description of the species and so all should be included in the type series .</p></div>	https://treatment.plazi.org/id/D42CA7775444CB0AFF6DFCB17893FB55	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Prieto-Manzanares, Miguel;Masó-Ros, Glòria	Prieto-Manzanares, Miguel, Masó-Ros, Glòria (2025): Annotated catalogue of type specimens of Iberian Leptodirini Lacordaire, 1854 (Coleoptera: Leiodidae: Cholevinae) deposited in the Natural Science Museum of Barcelona (Spain). Zootaxa 5682 (1): 1-102, DOI: 10.11646/zootaxa.5682.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
