identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
D477D357D22AA53B7E93C2D2FBB00326.text	D477D357D22AA53B7E93C2D2FBB00326.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atyidae De Haan 1849	<div><p>Family Atyidae De Haan, 1849</p><p>Diagnosis (from [7]): freshwater shrimps from small to large size (4–120 mm). Body usually pigmented, except for some subterranean species. Ocular peduncle without projection on the dorsal margin near the cornea. Last two articles of antennular peduncle longer or as long as wide; stylocerite without dorsal process, or vestigial one. Mandibles without palp but with a well-developed molar process. Exopodite of maxillae with a posterior tuft of long setae. Exopodite of third maxillipeds prolonged by a transverse or oblique projection on its antero-internal angle; epipodites of third maxilliped very reduced or vestigial, without posterior lobe. Dactylus of second maxilliped longer or as long as propodus. Dactylus of third maxilliped armed with a rather strong spine, arched and never styliform. Chelipeds with simple carpus and ending in a chela. Dactylus of walking legs moderately long, flattened or conical, armed with a terminal spine and a series of spinules, and numerous and contiguous on the fifth pereiopods. Epipods always lack a projection. Telson without median stylus on its posterior margin.</p></div>	https://treatment.plazi.org/id/D477D357D22AA53B7E93C2D2FBB00326	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D22AA5387E93C03AFCCB0669.text	D477D357D22AA5387E93C03AFCCB0669.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atyopsis Chace 1983	<div><p>Genus Atyopsis Chace, 1983</p><p>Diagnosis (from [44]): body pigmented, eyes well developed; rostrum not strongly compressed laterally, median dorsal carina typically unarmed, ventral keel with 2–16 teeth; anterior margin of carapace armed with antennal spine, pteygostomian margin sharply acute; supraorbital spines absent; telson with posterolateral angles overreaching setigerous posterior margin; third maxilliped not terminating in single apical spine; pereiopods without exopods; first and second pereiopods with chelae monomorphic (without palm), fingers tipped with brushes of long setae adapted for filter feeding, carpus of both appendages excavate distally, little if at all longer than broad; third pereiopod of large males with prominent spur on merus; branchial complement consisting of five pleurobranchs, three arthrobranchs, one podobranch, five epipods (reduced posteriorly), no mastigobranchs; first pleopod of male with endopod rigid, rhomboidally oval, submarginally spinose; second pleopod of male with appendix masculina subcylindrical, spinose over entire length distal to base of appendix interna.</p></div>	https://treatment.plazi.org/id/D477D357D22AA5387E93C03AFCCB0669	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D229A5387E93C572FB7503AE.text	D477D357D229A5387E93C572FB7503AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atyopsis spinipes (Newport 1847)	<div><p>Atyopsis spinipes (Newport, 1847)</p><p>Figures 3A and 9A</p><p>- Atya spinipes Newport 1847: 159 [type locality: Philippine Islands];</p><p>- Atyopsis spinipes —Chace 1983 [44]: 35, Figures 20–22.</p><p>Material examined</p><p>PALAU: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.52913&amp;materialsCitation.latitude=7.4528165" title="Search Plazi for locations around (long 134.52913/lat 7.4528165)">Babeldaob Island</a>. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.52913&amp;materialsCitation.latitude=7.4528165" title="Search Plazi for locations around (long 134.52913/lat 7.4528165)">Tabechedingriver</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.52913&amp;materialsCitation.latitude=7.4528165" title="Search Plazi for locations around (long 134.52913/lat 7.4528165)">station 1</a>, 27 February 2011, 07 ◦ 27.169 ′ N, 134 ◦ 31.748 ′ E, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.61597&amp;materialsCitation.latitude=7.6087832" title="Search Plazi for locations around (long 134.61597/lat 7.6087832)">Ngerchokl river</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.61597&amp;materialsCitation.latitude=7.6087832" title="Search Plazi for locations around (long 134.61597/lat 7.6087832)">station 2</a>, 27 February 2011, 07 ◦ 36.527 ′ N, 134 ◦ 36.958 ′ E, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.62923&amp;materialsCitation.latitude=7.6902833" title="Search Plazi for locations around (long 134.62923/lat 7.6902833)">Tireloich river</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.62923&amp;materialsCitation.latitude=7.6902833" title="Search Plazi for locations around (long 134.62923/lat 7.6902833)">station 3</a>, 07 ◦ 41.417 ′ N, 134 ◦ 37.754 ′ E, 28 February 2011, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.59288&amp;materialsCitation.latitude=7.591367" title="Search Plazi for locations around (long 134.59288/lat 7.591367)">Ngardmau waterfall</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.59288&amp;materialsCitation.latitude=7.591367" title="Search Plazi for locations around (long 134.59288/lat 7.591367)">station 7</a>, 2 March 2011, 07 ◦ 35.482 ′ N, 134 ◦ 35.573 ′ E, observed only; USNM 105280: 4 ♀, cl 7.3 mm –11.0 mm, south fork of Arakitaoch stream, station 170A, 26 November 1956, coll. H.A. Fehlmann; USNM 190993: 1 ♀, cl 12.0 mm, Metengakakumer river, station 2-MR- 4-05, 13 February 1973, coll. G. Bright. Tissue only: CA1861 and CA1862, no locality data.</p><p><a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.49403&amp;materialsCitation.latitude=7.3404" title="Search Plazi for locations around (long 134.49403/lat 7.3404)">Koror Island</a>. USNM 104899: 1 ♀, cl 8.3 mm, station 216, 7 ◦ 20.424 ′ N, 134 ◦ 29.642 ′ E, 11 October 1955, coll. H.A. Fehlmann.</p><p>FEDERATED STATES OF MICRONESIA: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.20442&amp;materialsCitation.latitude=6.9207" title="Search Plazi for locations around (long 158.20442/lat 6.9207)">Pohnpei Island</a>. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.20442&amp;materialsCitation.latitude=6.9207" title="Search Plazi for locations around (long 158.20442/lat 6.9207)">Nanpil river</a> <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.20442&amp;materialsCitation.latitude=6.9207" title="Search Plazi for locations around (long 158.20442/lat 6.9207)">station 1</a>, 9 March 2012, 127 m a.s.l., 06 ◦ 55.242 ′ N, 158 ◦ 12.265 ′ E; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22116&amp;materialsCitation.latitude=6.9083" title="Search Plazi for locations around (long 158.22116/lat 6.9083)">Nanpil river station 2</a>, 9 March 2012, 93 m a.s.l., 06 ◦ 54.498 ′ N, 158 ◦ 13.270E, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.21463&amp;materialsCitation.latitude=6.9185166" title="Search Plazi for locations around (long 158.21463/lat 6.9185166)">Nanpil river station 3</a>, 9 March 2012, 52 m a.s.l., 06 ◦ 55.111 ′ N, 158 ◦ 12.878 ′ E, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.1815&amp;materialsCitation.latitude=6.850917" title="Search Plazi for locations around (long 158.1815/lat 6.850917)">Lehn Mesi river station 1</a>, 10 March 2012, 197 m a.s.l., 06 ◦ 51.055 ′ N, 158 ◦ 10.890 ′ E, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.18993&amp;materialsCitation.latitude=6.8517833" title="Search Plazi for locations around (long 158.18993/lat 6.8517833)">Lehn Mesi river station 2</a>, 10 March 2012, 139 m a.s.l., 06 ◦ 51.107 ′ N, 158 ◦ 11.396 ′ E, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.26683&amp;materialsCitation.latitude=6.8651" title="Search Plazi for locations around (long 158.26683/lat 6.8651)">Senipehn river station 2</a>, 12 March 2012, 95 m a.s.l., 06 ◦ 51.906 ′ N, 158 ◦ 16.010 ′ E, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22583&amp;materialsCitation.latitude=6.9447165" title="Search Plazi for locations around (long 158.22583/lat 6.9447165)">Nanpil river station 5</a>, 5 m a.s.l., 06 ◦ 56.683 ′ N, 158 ◦ 13.550 ′ E, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.28687&amp;materialsCitation.latitude=6.8434834" title="Search Plazi for locations around (long 158.28687/lat 6.8434834)">Mahnd river</a>, 14 March 2012, 107 m a.s.l., 06 ◦ 50.609 ′ N, 158 ◦ 17.212 ′ E, observed only; unnamed river 1 estuary, 5 m a.s.l., 06 ◦ 51.120 ′ N, 158 ◦ 17.854 ′ E, observed only; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.26036&amp;materialsCitation.latitude=6.86325" title="Search Plazi for locations around (long 158.26036/lat 6.86325)">Senipehn river station 3</a>, 119 m a.s.l., 06 ◦ 51.795 ′ N, 158 ◦ 15.622 ′ E, observed only, M. Castelin, P. Keith, P. Gerbeaux, G. Marquet and L. Taillebois. Tissue only: CA1873 CA1872 and CA1874, no locality data, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois.</p><p>Short diagnosis</p><p>Body pigmented, eyes well-developed. Rostrum not strongly compressed laterally, dorsal carina typically unarmed, ventral margin with two–six discrete teeth, and anterior margin of carapace armed with antennal spine. Pterygostomian margin sharply acute. first and second pereopods with chelae monomorphic (without palm); fingers tipped with brushes of long setae adapted for filter feeding. Third pereopods of large males with prominent spur on merus.</p><p>Distribution</p><p>Pacific, extending from the Philippines and Indonesia to Samoa.</p><p>Habitat</p><p>This species prefers fresh and fast-flowing waters. Its biotope is characterized by rocky walls under waterfalls, where individuals aggregate in great numbers.</p></div>	https://treatment.plazi.org/id/D477D357D229A5387E93C572FB7503AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D228A5397E93C7A7FA8206DA.text	D477D357D228A5397E93C7A7FA8206DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atyoida Randall 1840	<div><p>Genus Atyoida Randall, 1840</p><p>Diagnosis (from [44]): body pigmented, eyes well developed; rostrum not strongly compressed laterally, median dorsal carina typically unarmed, ventral keel with zero–four teeth; anterior margin of carapace armed with antennal spine, pterygostomian margin sharply or bluntly acute; supraorbital spines absent; telson with setigerous posterior margin overreaching posterolateral angles; third maxilliped with uncinate terminal spine; pereiopods without exopods; first and second pereiopods with chelae heteromorphic (with or without palm), fingers tipped with brushes of setae, carpus of both appendages excavate distally, little if at all longer than broad; third pereiopod without meral spur in large males; branchial complement consisting of five pleurobranchs, three arthrobranchs, one podobranch, five epipods, and five mastigobranchs; first pleopod of male with endopod tapering sinuously but rather regularly to slender apex; second pleopod of male with appendix masculina sub-cylindrical, spinose over more than distal half.</p></div>	https://treatment.plazi.org/id/D477D357D228A5397E93C7A7FA8206DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D228A5347E93C5EFFBAF0698.text	D477D357D228A5347E93C5EFFBAF0698.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atyoida chacei Mazancourt & Marquet & Keith 2024	<div><p>Atyoida chacei sp. nov.</p><p>urn:lsid:zoobank.org:act:E409BE2B-D026-46D1-BE44-9CD5B34FC8E0</p><p>Figures 3B, 4 and 9B</p><p>- Atyoida cf. pilipes —Lorang et al., 2020 [45]: 300, Figure 1;</p><p>- Atya serrata —Miyake, 1938 [10]: 111;</p><p>- Atyoida pilipes —Chace, 1983 [44]: 13, Figures 5–8 (part).—Maciolek and Ford, 1987 [13]: 628.—Nelson et al., 1996 [14]: 11.—Buden et al., 2001 [15]: 260.—Leberer and Cai, 2003 [18]: 354.—March et al., 2003 [20]: 126.—Page et al., 2007 [21]: 139, Figure 1 (part); 2008 [46]: 73, Figure 4 (part).—Benstead et al., 2009 [16]: 459.</p><p>Material examined</p><p>Type material:</p><p>PALAU: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.62923&amp;materialsCitation.latitude=7.6902833" title="Search Plazi for locations around (long 134.62923/lat 7.6902833)">Babeldaob Island</a>. Holotype: MNHN-IU-2024-439 (DNA: CA1858), 1 ♂, cl 4.7 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.62923&amp;materialsCitation.latitude=7.6902833" title="Search Plazi for locations around (long 134.62923/lat 7.6902833)">Tireloich river</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.62923&amp;materialsCitation.latitude=7.6902833" title="Search Plazi for locations around (long 134.62923/lat 7.6902833)">Station 3</a>), 07 ◦ 41.417 ′ N, 134 ◦ 37.754 ′ E, 28 February 2011, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet and L. Taillebois. Paratypes: MNHN-IU-2024- 440 (DNA: CA1859), 1 ♂, cl 4.7 mm, MNHN-IU-2024-441 (DNA: CA1863), 1 ♂, cl 4.5 mm, and MNHN-IU-2024-442 (DNA: CA1860), 1 ♀, cl 7.5 mm, same data as holotype. CA1863, no locality data. USNM 104901: 6 ♀ ovig., cl 5.9–7.3 mm and 1 ♀, cl 4.9 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.59636&amp;materialsCitation.latitude=7.5472665" title="Search Plazi for locations around (long 134.59636/lat 7.5472665)">Arakitaoch river</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.59636&amp;materialsCitation.latitude=7.5472665" title="Search Plazi for locations around (long 134.59636/lat 7.5472665)">station 241</a>), 7 ◦ 32.836 ′ N, 134 ◦ 35.782 ′ E, 18 October 1955, coll. H.A. Fehlmann. USNM 104926: 1 ♂, cl 4.2 mm, 1 ♀ ovig., cl 6.0 mm and 4 ♀, cl 4.0 mm– 6.8 mm, Station 170-A, 27 November 1956, coll. H.A. Fehlmann. USNM 105280: 1 ♀, cl 5.8 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.5701&amp;materialsCitation.latitude=7.5259333" title="Search Plazi for locations around (long 134.5701/lat 7.5259333)">South Fork of Arakitaoch river</a> (station 170- A), 7 ◦ 31.556 ′ N, 134 ◦ 34.206 ′ E, 15 November 1956, coll. H.A. Fehlmann. Arakabesan Island: USNM 172586: 1 ♀ ovig., cl 7.0 mm, below reservoir, station 2 A 14-01, 9 August 1976, coll. G. Bright.</p><p>FEDERATED STATES OF MICRONESIA: Pohnpei Island . MNHN-IU-2024-443 (DNA: CA1869): 1 ♂, cl 4.6 mm, MNHN-IU-2024-444: 1 ♀, cl 5.1 mm, MNHN-IU-2024-445 (DNA CA1870): 1 ♀ ovig., cl 6.7 mm and MNHN-IU-2024-446 (DNA CA 1871), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.26683&amp;materialsCitation.latitude=6.8651" title="Search Plazi for locations around (long 158.26683/lat 6.8651)">Senipehn river</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.26683&amp;materialsCitation.latitude=6.8651" title="Search Plazi for locations around (long 158.26683/lat 6.8651)">Station 7</a>), 06 ◦ 51.906 ′ N, 158 ◦ 16.010 ′ E, March 12 2012, 95 m a.s.l., coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. ZMB 8926: 4 ♀, cl 4.7–6.1 mm, February 1880, coll. O. Finsch.</p><p>Comparative material:</p><p>Atyoida pilipes (Newport, 1947): See [45].</p><p>Atyoida tahitensis Stimpson, 1860: See [45].</p><p>Description</p><p>Cephalothorax. Rostrum short (Figure 4o–q), 0.2–0.4 of cl, reaching near to or slightly beyond end of basal segment of antennular peduncle, with dorsal margin convex, ventral margin usually unarmed, sometimes with 1–2 teeth; antennal spine fused with inferior orbit angle, acute; pterygostomian margin acute.</p><p>Eyes well-developed. Antennular peduncle stout (Figure 4o), 0.46 (♀)–0.65 (♂) times as long as carapace; basal segment shorter than half length of antennular peduncle, second segment longer than third segment. Stylocerite reaching near to or slightly beyond end of basal segment of antennular peduncle.</p><p>Pereiopods. Epipods on first four pereiopods. P1 and P2 similar in size and shape. Chelae dimorphic, ortmannioid in shape (palm present, Figure 4b,d) mostly in males and atyoid (palm virtually absent, Figure 4a,c) mostly in females. P1 (Figure 4a,b) chela about 1.9–4.2 times as long as wide, movable finger 3.9–5.9 times as long as wide, 1.7–2.0 times length of palm with tufts of long setae distally, carpus excavated strongly anteriorly, 1.0–1.3 times as long as wide. P2 (Figure 4c,d) chela 2.2–3.9 times as long as wide: dactylus 3.7–6.3 times as long as wide, 2.4 times length of palm with tufts of long setae distally, P2 carpus excavated strongly anteriorly, 1.0–1.8 times as long as wide. P3 (Figure 4e) with row of setae on outer surface of merus, very short dactylus (Figure 4f) ending in a strong claw, 1.7–3.2 times as long as wide (terminal spine included) with 2–6 spines on flexor margin in addition to the terminal spine; propodus 3.0–6.6 times as long as wide, 2.5–3.7 times as long as dactylus. P5 (Figure 4g) with setae only present on carpus; dactylus (Figure 4h) short, 2.1–3.5 times as long as wide with 8–18 spines on flexor margin; propodus 5.9–13.4 times as long as wide, 3.3–5.6 times as long as dactylus.</p><p>Abdomen. Sixth abdominal somite 0.45 times length of carapace, 1.2 times as long as fifth somite, 0.79 times as long as telson.</p><p>Telson (Figure 4k) 2.4–2.8 times as long as wide, with from two to eight pairs of dorsal spinules, often not in pairs, one pair of dorsolateral spinules near dorsal end; posterior margin rounded with 8–11 plumose intermediate setae distinctly longer than lateral ones.</p><p>Male Pl1 (Figure 4l): Endopod subtriangular, 3.4 times as long as wide, reaching 0.59 times length of endopod, with a distinct appendix interna near distal end of endopod.</p><p>Male Pl2 (Figure 4m): Appendix masculina reaching 0.58 times length of endopod, inner and distal surface densely lined with long spines; appendix interna at basal 0.27 times length of appendix masculina, extending to distal 0.33 times length of appendix interna.</p><p>Preanal carina triangular (Figure 4j) without spine.</p><p>Uropodal diaeresis (Figure 4i) with 15–20 spinules.</p><p>Eggs (Figure 4n): undeveloped: 0.31–0.29 mm × 0.52–0.51 mm.</p><p>Etymology</p><p>A. chacei sp. nov. is named in honor of Fenner A. Chace, Jr (1908–2004) as an acknowledgement to his posthumous cooperation with us on the study of Atya -like shrimps of the Indo-Pacific region and for being the first to suggest that the Atyoida from Palau may belong to a distinct species.</p><p>Colour pattern</p><p>Body transparent, flanks whitish ornamented with numerous dark blue-black spots and stripes all over the body.</p><p>Distribution</p><p>Reported only from Babeldaob Island and Pohnpei Island in Micronesia.</p><p>Habitat</p><p>This species is largely rheophile and prefers fresh and fast-flowing waters.</p><p>Remarks</p><p>Atyoida chacei sp. nov. resembles A. pilipes and A. tahitensis (Table 3) by its short, unarmed, bent rostrum, as well as by the proportions between the joints of pereiopods and the presence of ortmannioid and atyid chelae. The absence of caridinoid chelae in males or in females in these three species distinguishes them from A. serrata (Bate, 1888) and A. bisulcata Randall, 1840 . Indeed, in these two species, the chelae are trimorphic: atyoid, caridinoid, and ortmannioid (see [44]).</p><p>Due to their high morphological variability, finding diagnostic characters to separate A. chacei sp. nov. from A. pilipes or A. tahitensis turned out to be a difficult task (see Table 3), so much so that they could be considered to be cryptic species, as they can only be distinguished by the following characters:</p><p>- Frequently, presence of one or more teeth on the ventral margin of the rostrum, vs. always 0 in A. pilipes and A. tahitensis . This character was already highlighted by Chace (1983): “Of 613 specimens with complete rostra from the Palau Islands, 270 have no ventral rostral teeth, 240 have 1, 97 have 2, 5 have 3, and 1 has 4”;</p><p>- Two–eight pairs of dorsal spinules, often not in pairs for A. chacei sp. nov. (vs. one–five pairs of dorsal spinules, often not in pairs for A. pilipes, vs. from two to four pairs of dorsal spinules, often in pairs for A. tahitensis);</p><p>- Posterior margin rounded with from 8 to 11 plumose intermediate setae distinctly longer than lateral ones (vs. 4 to 9 plumose intermediate setae distinctly longer than lateral ones for A. pilipes, vs. with 6 to 10 plumose intermediate setae distinctly longer than lateral ones for A. tahitensis);</p><p>- By the size of undeveloped eggs (0.51–0.52 mm × 0.29–0.31 mm (vs. 0.47–0.60 mm × 0.31–0.39 mm for A. pilipes, vs. 0.55–0.70 mm × 0.29–0.54 mm for A. tahitensis)).</p><p>Nevertheless, following an integrative taxonomy framework, their genetic distinctiveness (16S p-distances: 9–15%) coupled with their discrete geographical distribution provides enough evidence to distinguish the two species as distinct taxa, as it was the case for other species of freshwater shrimps (e.g., [47–49]).</p></div>	https://treatment.plazi.org/id/D477D357D228A5347E93C5EFFBAF0698	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D225A5357E93C01FFB0C05D2.text	D477D357D225A5357E93C01FFB0C05D2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina H. Milne Edwards 1837	<div><p>Genus Caridina H. Milne Edwards, 1837</p><p>Diagnosis: From small- to medium-sized shrimps (cl: 10–40 mm); ocular peduncle with variable corneal dilatation; rostrum strongly compressed laterally, with or without serration on both margins; carapace with antennal spine on the infraorbital angle, protruding or fused with it; pterygostomian margin unarmed, supraorbital spines absent; third maxilliped with a series of unequal spines on the dactylus; pereiopods without exopods; first pereiopods shorter than second, both with palm, fingers tipped with brushes of setae, carpus of P1 sometimes excavated, P2 never, both clearly longer than broad; branchial complement consisting of five pleurobranchs, three arthrobranchs, and one podobranch, with variable number of epipods and five mastigobranchs; first pleopod of male with a flat endopod of diverse shapes, with or without appendix; second pleopod of male with appendix masculina sub-cylindrical, spinose over more than distal half.</p></div>	https://treatment.plazi.org/id/D477D357D225A5357E93C01FFB0C05D2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D224A5337E93C448FCAE0300.text	D477D357D224A5337E93C448FCAE0300.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina ponapensis Mazancourt & Marquet & Keith 2024	<div><p>Caridina ponapensis sp. nov.</p><p>urn:lsid:zoobank.org:act:8165E679-E718-4975-8AF1-9784D16FC9F3</p><p>Figures 3C, 5 and 9C</p><p>- Caridina cf. weberi Pohnpei —de Mazancourt et al., 2019 [47]: 166, Figures 2–5;</p><p>- Non Caridina weberi parvirostris —Maciolek &amp; Ford, 1987 [13]: 628;</p><p>- Non Caridina weberi —Maciolek &amp; Ford, 1987 [13]: 628 (part?).—Nelson et al., 1996 [14]: 11 (part?).—Buden et al., 2001 [15]: 257 (part?).</p><p>Material examined</p><p>Type material</p><p>FEDERATED STATES OF MICRONESIA: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22116&amp;materialsCitation.latitude=6.9083" title="Search Plazi for locations around (long 158.22116/lat 6.9083)">Pohnpei Island</a>. Holotype: MNHN-IU-2024-447: 1 ♂ cl 2.4 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22116&amp;materialsCitation.latitude=6.9083" title="Search Plazi for locations around (long 158.22116/lat 6.9083)">Nanpilriver</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22116&amp;materialsCitation.latitude=6.9083" title="Search Plazi for locations around (long 158.22116/lat 6.9083)">station 2</a>), 9 March 2012, 93 ma.s.l., 06 ◦ 54.498 ′ N, 158 ◦ 13.270 ′ E, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. Paratypes: MNHN-IU-2024-448: 1 ♀ ovig. cl 3.0 mm <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.20442&amp;materialsCitation.latitude=6.9207" title="Search Plazi for locations around (long 158.20442/lat 6.9207)">Nanpilriver</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.20442&amp;materialsCitation.latitude=6.9207" title="Search Plazi for locations around (long 158.20442/lat 6.9207)">station 1</a>), 9 March 2012, 127 ma.s.l., 06 ◦ 55.242 ′ N, 158 ◦ 12.265 ′ E, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. MNHN-IU-2024-449: 1 ♂ cl 2.4 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22116&amp;materialsCitation.latitude=6.9083" title="Search Plazi for locations around (long 158.22116/lat 6.9083)">Nanpilriver</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22116&amp;materialsCitation.latitude=6.9083" title="Search Plazi for locations around (long 158.22116/lat 6.9083)">station 2</a>), 9 March 2012, 93 ma.s.l., 06 ◦ 54.498 ′ N, 158 ◦ 13.270 ′ E, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. MNHN-IU-2024-450 (DNA: CA1390): 1 ♀ cl 3.8 mm and MNHN-IU-2024-451 (DNA: CA1391): 1 ♀ ovig. cl 3.7 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.18993&amp;materialsCitation.latitude=6.8517833" title="Search Plazi for locations around (long 158.18993/lat 6.8517833)">Lehn Mesi river</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.18993&amp;materialsCitation.latitude=6.8517833" title="Search Plazi for locations around (long 158.18993/lat 6.8517833)">station 2</a>), 06 ◦ 51.107 ′ N, 158 ◦ 11.396 ′ E, 10 March 2012, 139 ma.s.l., coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. MNHN-IU-2024-452 (DNA: CA1388): 1 ♀ ovig. cl 3.5 mm and MNHN-IU-2024-453 (DNA: CA1389): 1 ♀ ovig. cl 2.9 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.26683&amp;materialsCitation.latitude=6.8651" title="Search Plazi for locations around (long 158.26683/lat 6.8651)">Senipehn river</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.26683&amp;materialsCitation.latitude=6.8651" title="Search Plazi for locations around (long 158.26683/lat 6.8651)">station 2</a>), 95 ma.s.l., 06 ◦ 51.906 ′ N, 158 ◦ 16.010 ′ E, 12 March 2012, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. MNHN-IU-2024-454: 1 ♀ ovig. cl 3.1 mm; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.28687&amp;materialsCitation.latitude=6.8434834" title="Search Plazi for locations around (long 158.28687/lat 6.8434834)">Mahnd river</a>, 06 ◦ 50.609 ′ N, 158 ◦ 17.212 ′ E, 14 March 2012, 107 m a.s.l., coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois .</p><p>Comparative material</p><p>Caridina weberi De Man, 1892: Syntypes MNHN-IU-2015-1755 2 ♂ cl 4.4–4.5 mm, 1 ♀ ovig. cl 6.1 mm.</p><p>Caridina parvirostris De Man, 1892: Syntypes MNHN-IU-2015-1748 cl. 3.1–3.2 mm, 1 ♀ cl. 4.2 mm.</p><p>Caridina keiensis Roux, 1911: Lectotype NMB 6.IV.b 1 ♀ ovig. cl 5.7 mm; Paralectoptypes NMB 6.IV.a 2 ♂ cl 2.6 mm; NMB 6.IV.b 1 ♀ cl 5.5 mm; one male cl 3.4 mm.</p><p>Description</p><p>Cephalothorax. Suborbital angle indistinguishably fused with antennal spine. Pterygostomian margin rounded. Rostrum (Figure 5m,n): curved down, short, 0.2–0.3 of cl, not overreaching first segment of antennular peduncle, armed with four–six teeth on dorsal margin, zero of them situated on carapace behind orbital margin, ventral margin with zero–one teeth.</p><p>Eyes well-developed, anterior end reaching to 0.7 times length of basal segment of antennular peduncle. Antennular peduncle 0.57 times as long as carapace. Anterolateral angle reaching 0.28 times length of the second segment, second segment distinctly shorter than third segment. Stylocerite reaching to 0.9 times length of basal segment of antennular peduncle.</p><p>Pereiopods. Epipods on first four pereiopods. P1 (Figure 5a): chela about 1.8–2.0 times as long as wide, movable finger 2.0–3.3 times as long as wide, 0.5–1.2 times length of palm; carpus 1.3–1.9 times as longas wide. P2 (Figure 5b) more slender and longer than first pereiopod with chela 2.4–2.6 times as long as wide: movable finger 4.0–4.5 times as long as wide, 1.3–1.8 times length of palm; carpus slender 5.6–6.5 times as longas wide. P3 (Figure 5c): stout, dactylus (Figure 5e) 2.4–3.7 times as long as wide terminal spiniform (seta included) with five–six spines on flexor margin in addition to the terminal spine; propodus 7.9–8.9 times as long as wide, 3.5–4.8 times as long as dactylus. P5 (Figure 5d): dactylus (Figure 5f) 3.0–4.3 times as long as wide with 29–39 spiniform setae on flexor margin; propodus 10.6–15.7 times as long as wide, 4.1–5.2 times as long as dactylus.</p><p>Abdomen. Third abdominal somite with moderately convex dorsal profile. Sixth abdominal somite 0.48 times as long as carapace, 1.5 times as long as fifth somite, 0.86 times as long as telson.</p><p>Telson (Figure 5i) two times as long as wide, with four pairs of dorsal spinules and one pair of dorsolateral spinules; posterior margin with a median process, rounded with five–nine very long intermediate plumose setae longer than lateral ones.</p><p>Male Pl1 (Figure 5k): Endopod subtriangular, 0.53 times length of exopod, appendix interna stout, exceeding end of endopod.</p><p>Male Pl2 (Figure 5l): Long appendix masculina on second pleopod reaching 0.85 times length of endopod; appendix interna reaching about 0.64 times length of appendix masculina.</p><p>Preanal carina high (Figure 5g) and unarmed.</p><p>Uropodal diaeresis (Figure 5h) with 19–24 spinules.</p><p>Eggs sized: 0.43–0.50 mm × 0.25–0.29 mm (Figure 5j).</p><p>Habitat</p><p>This new species is largely rheophile and prefers waters that are fresh and fast-flowing from the lower to the higher courses.</p><p>Etymology</p><p>C. ponapensis sp. nov. is named after the former name of Pohnpei, Ponape, from where this new species is endemic.</p><p>Colour pattern (Figure 9C)</p><p>Body transparent with spots of chromatophores scattered over the body.</p><p>Distribution</p><p>This new species seems endemic to Pohnpei Island.</p><p>Remarks</p><p>Based on the form of the rostrum, the high pre-anal carina, the large number of uropodal teeth, and the shape of the telson, this new species belongs to the C. weberi species group. In this group, it is most similar to C. parvirostris by its short curved-down rostrum and slender P2 carpus: 5.6–6.5 (vs. 6.0–7.4). However, it could be easily separated from C. parvirostris by lower numbers of dorsal and ventral teeth on the rostrum, respectively, 4–6 (vs. 8–10) and 0–1 (vs. 2). This new species is also similar to C. keiensis by its rostrum (lower numbers of dorsal and ventral teeth on the rostrum, respectively, 4–6 (vs. 2–13) and 0–1 (vs. 1–2)) and its relatively long stylocerite reaching to 0.90 times the length of the basal segment of antennular peduncle (vs. 0.92), but it could be easily distinguished by its longer P2 carpus 5.6–6.5 (vs. 3.5–4.5), and the size of the eggs 0.43–0.50 mm × 0.25–0.29 mm (vs. 0.39–0.42 mm × 0.24–0.25 mm).</p></div>	https://treatment.plazi.org/id/D477D357D224A5337E93C448FCAE0300	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D222A52E7E93C0D9FADC0037.text	D477D357D222A52E7E93C0D9FADC0037.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina lobocensis Cai, Choy & Ng 2009	<div><p>Caridina lobocensis Cai, Choy &amp; Ng, 2009</p><p>Figures 3D and 6.</p><p>- Caridina lobocensis Cai, Choy, and Ng, 2009: 68, Figures 2–4 [type locality: Tributary of Loboc River, Loboc (approx. 9 ◦ 38.206 ′ N, 124 ◦ 1.807 ′ E), Bohol Island, Philippines];</p><p>-? Caridina lobocensis —de Mazancourt et al., 2019 [47]: 166, Figures 2–5.</p><p>Material examined</p><p>PALAU: Babeldaob Island . MNHN-IU-2024-455 (DNA: CA1552): 1 ♀ ovig. cl 4.6 mm and MNHN-IU-2024-456 (DNA: CA1553): 1 ♀ ovig. cl 4.2 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.61597&amp;materialsCitation.latitude=7.6087832" title="Search Plazi for locations around (long 134.61597/lat 7.6087832)">Ngerchokl river</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.61597&amp;materialsCitation.latitude=7.6087832" title="Search Plazi for locations around (long 134.61597/lat 7.6087832)">Station</a> 2), 07 ◦ 36.527 ′ N, 134 ◦ 36.958 ′ E, 27 February 2011, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. MNHN-IU-2024-457 (DNA: CA1056): 1 ♀ ovig. cl 4.2 mm and MNHN-IU-2024-458 (DNA: CA1554): 1 ♀ ovig. cl 4.3 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.62923&amp;materialsCitation.latitude=7.6902833" title="Search Plazi for locations around (long 134.62923/lat 7.6902833)">Tireloich river</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.62923&amp;materialsCitation.latitude=7.6902833" title="Search Plazi for locations around (long 134.62923/lat 7.6902833)">Station 3</a>), 07 ◦ 41.417 ′ N, 134 ◦ 37.754 ′ E, 28 February 2011, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. MNHN-IU-2024-459 (DNA: CA1555): 1 ♀ ovig. cl 4.3 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.6154&amp;materialsCitation.latitude=7.7176" title="Search Plazi for locations around (long 134.6154/lat 7.7176)">West Coast stream Northern Tip</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.6154&amp;materialsCitation.latitude=7.7176" title="Search Plazi for locations around (long 134.6154/lat 7.7176)">Station 4</a>), 07 ◦ 43.056 ′ N, 134 ◦ 36.924 ′ E, 28 February 2011, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. MNHN-IU-2024- 460 (DNA: CA1556): 1 ♀ ovig. cl 4.4 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.57784&amp;materialsCitation.latitude=7.5414" title="Search Plazi for locations around (long 134.57784/lat 7.5414)">Ngermeskang river</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.57784&amp;materialsCitation.latitude=7.5414" title="Search Plazi for locations around (long 134.57784/lat 7.5414)">Station 6</a>), 75 m a.s.l., 07 ◦ 32.484 ′ N, 134 ◦ 34.670 ′ E, 1 March 2011, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. USNM 105446: 2 ♀, cl 4.3–4.8 mm and 1 juv. cl 2.3 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.6231&amp;materialsCitation.latitude=7.71475" title="Search Plazi for locations around (long 134.6231/lat 7.71475)">Station 122</a>, 7 ◦ 42.885 ′ N, 134 ◦ 37.386 ′ E, 23 August 1955, Coll. F.M. Bayer. USNM 105447: 2 ♀ ovig., cl 4.4–5.0 mm and a juv. cl 3.0 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.5961&amp;materialsCitation.latitude=7.5472164" title="Search Plazi for locations around (long 134.5961/lat 7.5472164)">Station 241</a>, 7 ◦ 32.833 ′ N, 134 ◦ 35.766 ′ E, 18 October 1955, coll. H.A. Fehlmann. USNM 105448: 1 ♀ ovig., cl 3.0 mm and 2 ♀, cl 3.0– 3.9 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.57004&amp;materialsCitation.latitude=7.52595" title="Search Plazi for locations around (long 134.57004/lat 7.52595)">Arakitaoch river</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.57004&amp;materialsCitation.latitude=7.52595" title="Search Plazi for locations around (long 134.57004/lat 7.52595)">cascade zone</a>, 7 ◦ 31.557 ′ N, 134 ◦ 34.202 ′ E, 4 November 1956, coll. H.A. Fehlmann. Koror Island: USNM 105445: 2 ♀, 4.5–4.9 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.49403&amp;materialsCitation.latitude=7.34035" title="Search Plazi for locations around (long 134.49403/lat 7.34035)">Station 194</a>, 7 ◦ 20.421 ′ N, 134 ◦ 29.642 ′ E, 30 September 1955, coll. H.A. Fehlmann.</p><p>THE PHILIPPINES. RMNH.CRUS.D.54659: 1 ♀ ovig., cl 7.1 mm, Barrio Mainit, Tayabas, Quezon Province, Luzon, 2 November 1986, coll. D.S. Bale.</p><p>Description</p><p>Cephalothorax. Suborbital angle indistinguishably fused with antennal spine. Pterygostomian margin rounded. Rostrum (Figure 6k): straight, short, 0.4–0.5 times length of cl, reaching middle of the third segment of antennular peduncle, armed with 12–17 teeth on dorsal margin, 1–3 of them situated on carapace behind orbital margin, ventral margin with 2–3 teeth.</p><p>Eyes well-developed, anterior end reaching to 0.7 times length of basal segment of antennular peduncle. Antennular peduncle 0.54 times as long as carapace. Anterolateral angle reaching 0.28 times length of the second segment, second segment slightly shorter than third segment. Stylocerite reaching to the basal segment of antennular peduncle.</p><p>Pereiopods. Epipods on first four pereiopods. P1 (Figure 6a): chela about 1.3–1.6 times as long as wide, movable finger 1.9–2.4 times as long as wide, 0.5–0.7 times length of palm; carpus 1.3–1.6 times as longas wide. P2 (Figure 6b) more slender and longer than first pereiopod with chela 2.2–2.8 times as long as wide: movable finger 3.5–5.0 times as long as wide, 1.3–1.7 times length of palm; carpus slender 4.3–5.5 times as longas wide. P3 (Figure 6c): stout, dactylus (Figure 6e) 3.7–4.4 times as longas wide (terminal spiniform seta included) terminating in two very strong claws with four-five spiniform setae on flexor margin; propodus 7.3–9.9 times as longas wide, 3.7–4.4 times as longas dactylus. P5 (Figure 6d): dactylus (Figure 6f) with two strong claws, 2.5–3.7 as long as wide with 31–33 spiniform setae on flexor margin; propodus 11.7–14.3 times as long as wide, 4.3–7.4 times as long as dactylus.</p><p>Abdomen. Third abdominal somite with moderately convex dorsal profile. Sixth abdominal somite 0.45 times as long as carapace, 1.3 times as long as fifth somite, and 0.77 times as long as telson.</p><p>Telson (Figure 6g) two times as long as wide, with four pairs of dorsal spinules and one pair of dorsolateral spinules; posterior margin without a median process, rounded with from seven to nine very long intermediate plumose setae longer than lateral ones.</p><p>Pl1 and Pl2: no males in our collected specimens. According to Cai et al. [50], “Endopod of male first pleopod subtriangular, half-length of exopod, appendix interna stout, exceeding end of endopod. Appendix masculina of male second pleopod 2/3 length of endopod, with appendix interna reaching base of distal one-third of appendix masculina”.</p><p>Uropodal diaeresis (Figure 6i) with 16–21 spinules.</p><p>Pre-anal carina (Figure 6j) elevated, without spine but with a few stiff setae.</p><p>Ovigerous females with eggs sized 0.37–0.43 mm × 0.22–0. 27 mm (Figure 6h).</p><p>Habitat</p><p>This species is largely rheophile and prefers waters that are fresh and fast-flowing waters the lower to the higher courses.</p><p>Colour pattern</p><p>Unknown.</p><p>Distribution</p><p>This species has been found in Palau and the Philippines.</p><p>Remarks</p><p>Our specimens agree well with those described from Bohol Island (Philippines) by Cai et al. [50] in relation to their long stylocerite, their stout first pereiopod, their short fingers, their third to fifth pereiopods ending in two claws, and their egg size. The rostrum in our specimens seems shorter: reaching the middle of the third segment of antennular peduncle (vs. reaching slightly beyond end of antennular peduncle) and its pterygostomian margin is rounded (vs. pterygostomian margin subrectangular), but we consider these differences to be within intra-specific variability. This species is closely related to C. buehleri and C. gueryi, both recently re-described by de Mazancourt et al. [43], but they are clearly separated in our molecular analyses and they can be distinguished by its P1 chela 1.3–1.6 times as long as wide (vs. 2.2–2.3 for C. buehleri and 1.9–2.4 for C. gueryi), and by its P5 dactylus with two strong claws (vs. one strong claw for C. buehleri and C. gueryi). Its P2 carpus is 4.3–5.5 times as long as wide and its P5 propodus is 4.3–7.4 times as long as its dactylus (vs., respectively, 5.5–6.2 and 3.8–4.1 for C. buehleri). Its rostrum is armed with dorsal teeth, one–three of them situated on carapace behind orbital margin (vs. three–five for C. gueryi) and its P5 dactylus has 31–33 spiniform setae on its flexor margin (vs. 26–31 for C. gueryi).</p></div>	https://treatment.plazi.org/id/D477D357D222A52E7E93C0D9FADC0037	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D23FA52F7E93C2EEFCBF0628.text	D477D357D23FA52F7E93C2EEFCBF0628.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina H. Milne Edwards 1837	<div><p>Caridinatypus H. Milne Edwards, 1837</p><p>Figure 3E</p><p>- Caridinatypus H. Milne Edwards, 1837: 363 [type locality unknown].</p><p>Material examined</p><p>PALAU: Babeldaob Island . MNHN-IU-2018-3355 (DNA voucher: CA2015): 1 ♀ ovig. cl 5.2 mm; MNHN-IU-2018-3356 (DNA voucher: CA2016): 1 ♀ ovig. cl 4.5 mm; MNHN- IU-2018-3357 (DNA voucher: CA2017): 1 ♀ cl 5.0 mm and MNHN-IU-2018-3358 (DNA voucher: CA2018): 1 ♀ ovig. cl 5.5 mm, no locality data, February–March 2011, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. Ngerchokl river, 07 ◦ 36.527 ′ N, 134 ◦ 36.958 ′ E, 27 February 2011, M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois, observed only. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.62923&amp;materialsCitation.latitude=7.6902833" title="Search Plazi for locations around (long 134.62923/lat 7.6902833)">Tireloich river</a>, 07 ◦ 41.417 ′ N, 134 ◦ 37.754 ′ E, 28 February 2011, M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois, observed only. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.6154&amp;materialsCitation.latitude=7.7176" title="Search Plazi for locations around (long 134.6154/lat 7.7176)">West coast unnamed stream</a>, 07 ◦ 43.056 ′ N, 134 ◦ 36.924 ′ E, 28 February 2011, M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois, observed only.</p><p>FEDERATED STATES OF MICRONESIA: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.7606&amp;materialsCitation.latitude=13.39315" title="Search Plazi for locations around (long 144.7606/lat 13.39315)">Pohnpei Island</a>. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.7606&amp;materialsCitation.latitude=13.39315" title="Search Plazi for locations around (long 144.7606/lat 13.39315)">Nanpilriver</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.7606&amp;materialsCitation.latitude=13.39315" title="Search Plazi for locations around (long 144.7606/lat 13.39315)">station 5</a>), 06 ◦ 56.683 ′ N, 158 ◦ 13.550 ′ E, 5 m a.s.l.; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.7606&amp;materialsCitation.latitude=13.39315" title="Search Plazi for locations around (long 144.7606/lat 13.39315)">Mahnd river</a>, 14 March 2012, 107m a.s.l., 06 ◦ 50.609 ′ N, 158 ◦ 17.212 ′ E, observed only.</p><p>GUAM. ZMB 34613 (DNA: CA2791 and CA2792): twospecimens, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.7606&amp;materialsCitation.latitude=13.39315" title="Search Plazi for locations around (long 144.7606/lat 13.39315)">Yligriver</a>, 13 ◦ 23.589 ′ N 144 ◦ 45.636 ′ E, 2 ma.s.l., January 2011, coll. D.C. Rogers, A. Farahi, and T. Jones. ZMB 34614 (DNA: CA2794 and CA2795): two specimens, Taguan river, January 2011, coll. D.C. Rogers, A. Farahi, and T. Jones.</p><p>Distribution</p><p>This species has a wide distribution in rivers of the Indo-West Pacific area ranging from South and East Africa to Japan, Fiji, and Australia.</p><p>Remarks</p><p>This species here reported from Palau and Guam is found in most of the Indo-Pacific region (from Madagascar to Fiji).</p></div>	https://treatment.plazi.org/id/D477D357D23FA52F7E93C2EEFCBF0628	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D23EA52C7E93C4B1FA3C05EB.text	D477D357D23EA52C7E93C4B1FA3C05EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina serratirostris De Man 1892	<div><p>Caridina cf. serratirostris</p><p>Figures 3Hand 7.</p><p>Material examined</p><p>FEDERATED STATES OF MICRONESIA: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.21463&amp;materialsCitation.latitude=6.9185166" title="Search Plazi for locations around (long 158.21463/lat 6.9185166)">Pohnpei Island</a>. MNHN-IU-2024-461 (DNA: CA1245): 1 ♀ ovig. cl 2.5 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.21463&amp;materialsCitation.latitude=6.9185166" title="Search Plazi for locations around (long 158.21463/lat 6.9185166)">Nanpilriver</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.21463&amp;materialsCitation.latitude=6.9185166" title="Search Plazi for locations around (long 158.21463/lat 6.9185166)">station 3</a>), 9 March 2012, 52 ma.s.l., 06 ◦ 55.111 ′ N, 158 ◦ 12.878 ′ E, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois.</p><p>Distribution</p><p>Reported only from Pohnpei Island.</p><p>Remarks</p><p>This species is distinct from all the others known in this group based on molecular data (minimum 16S p-distance: 14%), but the only specimen available is too damaged to provide a sufficient morphological description. Rostral formula: 19 (10)/3, rostrum 0.38 times as long as carapace length, egg size 0.40–0.43 mm × 0.23–0.26 mm (Figure 7).</p></div>	https://treatment.plazi.org/id/D477D357D23EA52C7E93C4B1FA3C05EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D23DA52A7E93C6F0FD9D011B.text	D477D357D23DA52A7E93C6F0FD9D011B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina rintelenorum Mazancourt & Marquet & Keith 2024	<div><p>Caridina rintelenorum sp. nov.</p><p>urn:lsid:zoobank.org:act:FF389D46-B517-4D59-9102-34638B8B62C2</p><p>Figures 3G, 8 and 9D.</p><p>Material examined</p><p>Type material</p><p>PALAU: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.59288&amp;materialsCitation.latitude=7.591367" title="Search Plazi for locations around (long 134.59288/lat 7.591367)">Babeldaob Island</a>. Holotype: MNHN-IU-2024-462 (DNA: CA1864): 1 ♀ ovig. cl 3.3 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.59288&amp;materialsCitation.latitude=7.591367" title="Search Plazi for locations around (long 134.59288/lat 7.591367)">Ngardmau river</a>, waterfall, 07 ◦ 35.482 ′ N, 134 ◦ 35.573 ′ E, 2 March 2011, coll. M. Castelin, P. Gerbeaux, P. Keith, G. Marquet, and L. Taillebois.</p><p>Paratypes: RMNH.CRUS.D.16641: 1 ♀ ovig. cl 2.9 mm, stn. 122, Luhet river, Ollei village, Ngarchelong peninsula, 23 August 1955, coll. A. Fehlmann, George Vanderbilt Foundation; USNM 105450: 1 ♀ ovig., cl 3.0 mm and 1 juv. cl 2.0 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.62305&amp;materialsCitation.latitude=7.7148" title="Search Plazi for locations around (long 134.62305/lat 7.7148)">Station 122</a>, 7 ◦ 42.888 ′ N, 134 ◦ 37.383 ′ E, 23 August 1955, coll. F.M. Bayer. USNM 172595: 1 ♂ cl 2.2 mm, 1 ♀ ovig. cl 2.6 mm and 1 juv. cl 1.9 mm, Ngerbekuu river, station 1NBK 4-12, 1 June 1978, coll. G. Bright. USNM 172596: 1 ♀ ovig. cl 2.7 mm, Ngerbekuu river, station 1NBK 4-11, 1 June 1978, coll. G. Bright.</p><p>SOLOMON ISLANDS. <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.54057&amp;materialsCitation.latitude=-7.625167" title="Search Plazi for locations around (long 158.54057/lat -7.625167)">Isabel Island</a>. MNHN-IU-2024-463 (DNA voucher: CA2534): 1 ♀ ovig. cl 2.9 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.54057&amp;materialsCitation.latitude=-7.625167" title="Search Plazi for locations around (long 158.54057/lat -7.625167)">Hobolito river</a>, 7 ◦ 37.510 ′ S, 158 ◦ 32.434 ′ E, 24 October 2019, coll. R. Causse, P. Keith, and C. Lord .</p><p>Comparative material</p><p>Caridina celebensis . INDONESIA. Paralectotype (lectotype designated in [51]), MNHN- IU-2015-1926: 1 ♀ ovig. cl 3.5 mm, Sulawesi, Luwu, river near Palopo, February 1889, coll. M. Weber . SOLOMON ISLANDS: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=157.072&amp;materialsCitation.latitude=-7.84935" title="Search Plazi for locations around (long 157.072/lat -7.84935)">Kolombangara Island</a>. MNHN-IU-2018-2941: 1 ♀ ovig. cl 3.1 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=157.072&amp;materialsCitation.latitude=-7.84935" title="Search Plazi for locations around (long 157.072/lat -7.84935)">Lodumoe river</a>, 07 ◦ 50.961 ′ S, 157 ◦ 04.320 ′ E, 16 November 2015, coll. P. Keith, C. Lord, and G. Marquet. MNHN-IU-2018-2942: 1 ♀ ovig. cl 3.2 mm, same collection data as for preceding. MNHN-IU-2018-2943 (DNA: CA1518): 1 ♀ cl 3.5 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=156.96553&amp;materialsCitation.latitude=-7.907" title="Search Plazi for locations around (long 156.96553/lat -7.907)">Munga river</a>, 07 ◦ 54.420 ′ S, 156 ◦ 57.932 ′ E, 5 ma.s.l., 19 November 2015, coll. P. Keith, C. Lord, and G. Marquet. MNHN-IU-2018-2944: 1 ♀ ovig. cl 3.4 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=157.17722&amp;materialsCitation.latitude=-8.0643835" title="Search Plazi for locations around (long 157.17722/lat -8.0643835)">Liva river</a>, 08 ◦ 03.863 ′ S, 157 ◦ 10.633 ′ E, 14 m a.s.l., 11 November 2015, coll. P. Keith, C. Lord and G. Marquet. Vella Lavella Island . MNHN-IU-2018-2945: 1 ♀ ovig. cl 3.5 mm, Wariassi river, 29 October 2016, coll. P. Keith, and C. Lord . VANUATU: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=168.2856&amp;materialsCitation.latitude=-17.564405" title="Search Plazi for locations around (long 168.2856/lat -17.564405)">Efate Island</a>. MNHN-IU-2018-2946 (DNA: CA1381): 1 ♀ cl 3.0 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=168.2856&amp;materialsCitation.latitude=-17.564405" title="Search Plazi for locations around (long 168.2856/lat -17.564405)">Marona river</a>, 17 ◦ 33 ′ 51.860 ′′ S, 168 ◦ 17 ′ 08.140 ′′ E, 5 m a.s.l., 9 November 2014, coll. A. Acou, G. Marquet, and M. Mennesson. MNHN-IU-2018-2947: 1 ♀ ovig. cl 3.4 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=168.2577&amp;materialsCitation.latitude=-17.6763" title="Search Plazi for locations around (long 168.2577/lat -17.6763)">Mele river</a>, 17 ◦ 40.578 ′ S, 168 ◦ 15.462 ′ E, 9 November 2014, 32 ma.s.l., coll. A. Acou, G. Marquet and M. Mennesson.</p><p>Description</p><p>Cephalothorax. Antennal spine ventral to inferior orbital angle. Pterygostomian margin rounded. Rostrum (Figure 8k): straight, 0.5–0.6 times length of cl, reaching to base of third segment of antennular peduncle, armed with 19–21 teeth on dorsal margin, 7–10 of them situated on carapace behind orbital margin, ventral margin with 5–8 teeth. Eyes developed, anterior end reaching to 0.65 times length of basal segment of antennular peduncle. Long antennular peduncle, 0.69 (♀) times as longas carapace. Second segment distinctly longer than third segment. Long stylocerite reaching beyond end of first segment of antennular peduncle.</p><p>Pereiopods. Epipods on first four pereiopods. P1 (Figure 8a): chela about 2.1–3.1 times as long as wide, movable finger 2.5–5.2 times as long as wide, 0.9–1.6 times length of palm; carpus 3.4–4.9 times as longas wide. P2 (Figure 8b) more slender and longer than first pereiopod, with chela 5.0–6.1 times as long as wide: movable finger 6.2–7.0 times as long as wide, 1.3–1.4 times length of palm; carpus slender, 8.0–12.3 times as longas wide. P3 (Figure 8c): slender, dactylus (Figure 8e) 3.7–4.0 times as long as wide (terminal spiniform seta included), with 6-9 spiniform setae on flexor margin including terminal one; propodus 12.1–14.1 times as long as wide, 3.8–4.5 times as long as dactylus. P5 (Figure 8d): dactylus (Figure 8f) 3.7–4.5 as long as wide, with 7–13 spiniform setae on flexor margin; propodus 15.5–17.4 times as long as wide, 4.1 times as long as dactylus.</p><p>Abdomen. Third abdominal somite with moderately convex dorsal profile. Sixth abdominal somite 0.45 times as long as carapace, 1.6 times as long as fifth somite, shorter than telson.</p><p>Telson (Figure 8h). 2.4 times as long as wide, with four pairs of dorsal spinules and one pair of dorsolateral spinules; posterior margin with a median process, rounded, with 7–12 very long intermediate plumose setae longer than lateral ones.</p><p>Male pleopods. No males.</p><p>Pre-anal carina (Figure 8i). High, with a spine.</p><p>Uropodal diaeresis (Figure 8g). With 14–19 spinules.</p><p>Eggs (Figure 8j). Size: 0.34–0.41 mm × 0.19–0.24 mm.</p><p>Habitat</p><p>This new species occurs in the lower to middle course of the rivers, sometimes in slightly brackish conditions.</p><p>Etymology</p><p>C. rintelenorum sp. nov. is named in honor of Kristina and Thomas von Rintelen (Museum für Naturkunde, Berlin, Germany) as an acknowledgement to their continuous support for the research of the first author and their years of study of atyid shrimps from South-East Asia.</p><p>Colour pattern</p><p>Body (Figure 8d) with more or less wide transverse bands of whitish, brownish, or dark color. Pereiopods with alternating bands of light and dark color.</p><p>Distribution</p><p>This species occurs in Palau and Solomon Islands (Isabel Island) and is expected to have a wider distribution in the region.</p><p>Remarks</p><p>This species is more similar to C. celebensis than to C. serratirostris by its shorter rostrum reaching to the base of the third segment of the antennular peduncle (vs. reaching beyond the end of the antennular peduncle), with fewer dorsal teeth, 19–21 (vs. 22–26); its longer P2 carpus, 8.0–12.3 as long as wide (vs. 8.3–10.9), and P2 chela 5.0-6.1 times as long as wide (vs. 3.7–5.1), but its shorter movable finger 1.3–1.4 times the length of the palm (vs. 1.7–1.9) and P5 propodus 15.5–17.4 times as long as wide (vs. 18.7–24.0) and 4.1 times as long as the dactylus (vs. 4.5–5.5).</p><p>This new species differs from C. celebensis by its shorter rostrum, being 0.5–0.6 times length of cl (vs. 0.6–0.9), and by its P5 propodus 4.1 times as longas the dactylus (vs. 3.4–3.9).</p></div>	https://treatment.plazi.org/id/D477D357D23DA52A7E93C6F0FD9D011B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D23BA52B7E93C220FCEB063F.text	D477D357D23BA52B7E93C220FCEB063F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina rubella Fujino & Shokita 1975	<div><p>Caridina rubella Fujino &amp; Shokita, 1975</p><p>Figure 3F.</p><p>- Caridina rubella Fujino and Shokita, 1975: 102, Figure 6 [type locality: Izaga Cave, Miyako Island, Ryukyu, Japan].—Shokita and Nishijima, 1976 [52]: 71.—Shokita, 1979 [53]: 203; 2003 [54]: 250, Figures 18F and 19I; 2006 [55]: 61.—Maciolek, 1983 [25]: 607.—Cai and Anker, 2004 [56]: 243, Figures 6 and 7.—Fujita and Shokita, 2005 [57]: 203.—Cai and Shokita, 2006 [51]: 2145, Figures 6F–J, 7 and 8.—Fujita, 2007 [58]: 92; 2018: 16.—Cai and Ng, 2009 [59]: 1094.—Weese et al., 2012 [60]: 109.—De Grave et al., 2022 [61]: 711, Figures 1 and 2.</p><p>- Caridina sp. 1 —Marquet et al., 2003 [62]: 74.</p><p>Material examined</p><p>GUAM. ZMB 34612: 1 ♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=144.80388&amp;materialsCitation.latitude=13.528334" title="Search Plazi for locations around (long 144.80388/lat 13.528334)">Fai Fai Cave</a>, 13 ◦ 31.700 ′ N, 144 ◦ 48.233 ′ E, coll. D.C. Rogers .</p><p>Distribution</p><p>This species has a wide distribution in anchialine habitats in the Pacific: Japan (Ryukyu), Philippines (Bohol, Palawan, Panglao), Indonesia (Sulawesi), New Caledonia (Loyalty), Cook Islands, French Polynesia (Tahiti), Niue, Tonga (Tongatapu), Western Samoa (Upolu), Solomon Islands (Nggela Pile), and Guam.</p><p>Remarks</p><p>The single female specimen collected from Fai Fai Cave in Guam agrees well with the published descriptions of the species [51,61,63], having a rostral formula of 10 + 28/12 and a rostrum 1.19 times as long as the carapace. Interestingly, our sequence clusters with a 16S sequence of the short-rostrum morph of the species from Miyako Island which is thought to represent the “real” C. rubella, whereas the long-rostrum morph from the same area potentially represents an undescribed species [60]. Consequently, it appears that this species is morphologically variable, as recently illustrated by De Grave et al. [61], but probably hides cryptic species. Amolecular study of this species from its whole range would be needed to clarify its status.</p></div>	https://treatment.plazi.org/id/D477D357D23BA52B7E93C220FCEB063F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D23AA5287E93C4E5FB21063C.text	D477D357D23AA5287E93C4E5FB21063C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina appendiculata Jalihal & Shenoy 1998	<div><p>Caridina appendiculata Jalihal &amp; Shenoy, 1998</p><p>Figures 3I and 10A.</p><p>Caridina appendiculata Jalihal and Shenoy, 1998: 128 [type locality: River Bari (approx. 8 ◦ 21.070 ′ S, 120 ◦ 10.928 ′ E), Flores, Indonesia].—Cai and Ng, 2007 [64]: 1590.—Klotz et al., 2007 [65]: 7, Figure 3 (part), 4.—de Mazancourt et al., 2018 [66]: 1433, Figure 1, 4.; 2019 [47]: 166; 2020 [43]: 3, 78 (key), Figures 2H and 25C.—Short et al., 2019 [67]: 4, 15, Figure 7.— Hernawati et al., 2020 [68]: Figure 3.</p><p>- Caridina gracilirostris —De Man, 1892 [69]: 399 (part).—Holthuis, 1978 [70]: 35 (part);</p><p>- Caridina sp. E —Page et al., 2007 [21]: 648, Figures 2 and 3.—Cook et al., 2011 [71]: 278, Figure 3.</p><p>Material examined</p><p>PALAU: Babeldaob Island. MNHN-IU-2018-127 (DNA: CA1021): 1 ♀ ovig., cl 3.9 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.63498&amp;materialsCitation.latitude=7.498483" title="Search Plazi for locations around (long 134.63498/lat 7.498483)">small stream East Coast</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.63498&amp;materialsCitation.latitude=7.498483" title="Search Plazi for locations around (long 134.63498/lat 7.498483)">near the estuary</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.63498&amp;materialsCitation.latitude=7.498483" title="Search Plazi for locations around (long 134.63498/lat 7.498483)">Station 5</a>), 07 ◦ 29.909 ′ N, 134 ◦ 38.099 ′ E, 2 March 2011, 0 ma.s.l., coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois.</p><p>FEDERATED STATES OF MICRONESIA: Pohnpei Island . MNHN–IU–2018–128, 1 ♀ ovig. cl 3.2 mm and MNHN–IU–2018–129, 1 ♀ ovig. cl 3.6 mm (DNA: CA1333); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22583&amp;materialsCitation.latitude=6.9434834" title="Search Plazi for locations around (long 158.22583/lat 6.9434834)">Nanpil river estuary</a>, 5 m a.s.l., 06 ◦ 56.609 ′ N, 158 ◦ 13.550 ′ E, coll. P. Keith, P. Gerbeaux, G. Marquet, L. Taillebois, and M. Castelin, 13 March 2012; MNHN–IU–2018–130, 1 ♀ ovig. cl 3.0 mm (DNA: CA1330), MNHN–IU–2018–131, 1 ♀ ovig. cl 3.4 mm and MNHN–IU–2018–132, 1 ♀ cl 3.1 mm; unnamed river, estuary, 5 m a.s.l., 06 ◦ 48.579 ′ N, 158 ◦ 12.639 ′ E, coll. P. Keith, P. Gerbeaux, G. Marquet, L. Taillebois, and M. Castelin, 15 March 2012. MNHN-IU-2024-464 (DNA: CA1183), 2 ♀ ovig., cl 3.2–3.6 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22583&amp;materialsCitation.latitude=6.9434834" title="Search Plazi for locations around (long 158.22583/lat 6.9434834)">Nanpil river estuary</a> (<a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.22583&amp;materialsCitation.latitude=6.9434834" title="Search Plazi for locations around (long 158.22583/lat 6.9434834)">Station 9</a>), 06 ◦ 56.609 ′ N, 158 ◦ 13.550 ′ E; 13 March 2012; 5 ma.s.l., coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. MNHN-IU-2018-130, 1 ♀ ovig., cl 3.0 mm, 1 ♀, cl 3.1 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=158.21065&amp;materialsCitation.latitude=6.80965" title="Search Plazi for locations around (long 158.21065/lat 6.80965)">River 2</a>, 15 March 2012, 5 ma.s.l., 06 ◦ 48.579 ′ N, 158 ◦ 12.639 ′ E, coll. M. Castelin, P. Keith, P. Gerbeaux, G. Marquet, and L. Taillebois. USNM 104882: 1 juv. cl 2.8 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.56998&amp;materialsCitation.latitude=7.5259333" title="Search Plazi for locations around (long 134.56998/lat 7.5259333)">south fork of Arakitaoch river</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.56998&amp;materialsCitation.latitude=7.5259333" title="Search Plazi for locations around (long 134.56998/lat 7.5259333)">station 170-A</a>, 7 ◦ 31.556 ′ N, 134 ◦ 34.199 ′ E, 15 November 1956, coll. H.A. Fehlmann. USNM 172592: 4 ♀, cl 2.9–3.6 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.56998&amp;materialsCitation.latitude=7.5259333" title="Search Plazi for locations around (long 134.56998/lat 7.5259333)">Metengalakumer river</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.56998&amp;materialsCitation.latitude=7.5259333" title="Search Plazi for locations around (long 134.56998/lat 7.5259333)">station 1MR 4-08</a>, 17 November 1977, coll. G. Bright. MNHN–IU–2018–127, 1 ♀ ovig. cl 3.9 mm (DNA: CA1021), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.63498&amp;materialsCitation.latitude=7.498483" title="Search Plazi for locations around (long 134.63498/lat 7.498483)">small stream East Coast</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.63498&amp;materialsCitation.latitude=7.498483" title="Search Plazi for locations around (long 134.63498/lat 7.498483)">near the estuary</a>, 0 m a.s.l., 07 ◦ 29.909 ′ N, 134 ◦ 38.099 ′ E, 2 March 2011, coll. P. Keith, P. Gerbeaux, G. Marquet, L. Taillebois, and M. Castelin.</p><p>Distribution</p><p>C. appendiculata Jalihal and Shenoy, 1998 is known from Australia, Indonesia (Flores, Obira, and Sulawesi), Solomon Islands (Kolombangara, Isabel), Micronesia (Pohnpei), Palau, and Vanuatu (Aneityum, Efate, Epi and Santo) [66].</p><p>Habitat</p><p>All specimens were found in a typical brackish water environment (brackish water pool or colonizing the lower part of rivers, near the estuary).</p><p>Remarks de Mazancourt et al. [66] redescribed this species in detail.</p></div>	https://treatment.plazi.org/id/D477D357D23AA5287E93C4E5FB21063C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D239A5287E93C505FD9F01A5.text	D477D357D239A5287E93C505FD9F01A5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina variabilis de Mazancourt, Marquet, Rogers & Keith 2018	<div><p>Caridina variabilis de Mazancourt, Marquet, Rogers &amp; Keith, 2018</p><p>Figures 3K and 10C,D.</p><p>- Caridina variabilis de Mazancourt, Marquet, Rogers, and Keith, 2018b: 43, Figure 4 [type locality: Palau, Babeldaob Island, TabechedingRiver, 07 ◦ 27.169 ′ N, 134 ◦ 31.748 ′ E].—de Mazancourt et al., 2018 [9]: 6, Figure 2; 2019 [47]: 167, Figures 2 –5; 2020 [43]: 20, 31;</p><p>- Caridina brachydactyla uncata de Mazancourt, Marquet, Rogers, and Keith, 2018: 40 (nomen nudum);</p><p>- Caridina nilotica var. brachydactyla — Bouvier, 1925 [7]: 155 (part, specimens from Marianas);</p><p>- Caridina weberi — Leberer and Nelson, 2001 [19]: 389;</p><p>- Caridina brachydactyla — Bright, 1989 [23]: 35.—Leberer and Cai, 2003 [18]: 354;</p><p>- Caridina mertoni — Leberer and Cai, 2003 [18]: 354;</p><p>- Caridina longirostris — Leberer and Nelson, 2001 [19]: 389;</p><p>Material examined</p><p>See [9].</p><p>Distribution</p><p>This species occurs only in Guam and Palau [9].</p><p>Remarks</p><p>Holthuis (year unknown) named specimens from Palau, based on material collected by H. A. Fehlmann on the 30 October 1956 from Arakitaoch stream and deposited in USNM as C. brachydactyla uncata (“holotype” USNM 105430, “paratypes” UNSNM 105434) but never described or published this taxon. This species was previously confused with C. brachydactyla De Man, 1908 or C. mertoni Roux, 1911, depending on its variable morphology (see [9]).</p></div>	https://treatment.plazi.org/id/D477D357D239A5287E93C505FD9F01A5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D239A5297E93C2BEFD270583.text	D477D357D239A5297E93C2BEFD270583.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina variabilirostris de Mazancourt, Marquet & Keith 2018	<div><p>Caridina variabilirostris de Mazancourt, Marquet &amp; Keith, 2018</p><p>Figures 3J and 10B.</p><p>- Caridina variabilirostris de Mazancourt, Marquet, and Keith, 2018: 7, Figures 3 and 4 [type locality: Federated States of Micronesia, Pohnpei Island, Nanpilriver, 06 ◦ 54.252 ′ N, 158 ◦ 11.491 ′ E, 180 m a.s.l.].—de Mazancourt et al., 2019 [47]: 167, Figures 2–5; 2020 [43]: 20, 30;</p><p>- Caridina brachydactyla —Maciolek and Ford, 1987 [13]: 628;</p><p>- Caridina vitiensis —Maciolek and Ford, 1987 [13]: 628;</p><p>- Caridina longirostris —Maciolek and Ford, 1987 [13]: 628;</p><p>- Caridina sp. —de Mazancourt et al., 2017 [40]: 249, Figures 2 and 5;</p><p>- Caridina weberi —Maciolek and Ford, 1987 [13]: 628.—Nelson et al., 1996 [14]: 11.— Buden et al., 2001 [15]: 257.—Page et al., 2007 [21]: 649, 650, Figures 2 and 3.— von Rintelen et al., 2008 [72]: 2248, Figure 4.—Benstead et al., 2009 [16]: 459.—de Mazancourt et al., 2017 [40]: 226, Figure 4;</p><p>- Caridina nilotica var.? wycki —Roux, 1925 [12]: 149.</p><p>Material examined</p><p>See [8].</p><p>Distribution</p><p>This species is known from Pohnpei only thus far [8].</p><p>Remarks</p><p>This species used to be confused with C. brachydactyla De Man, 1908 or C. mertoni Roux, 1911 (see [8]).</p></div>	https://treatment.plazi.org/id/D477D357D239A5297E93C2BEFD270583	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D237A5267E93C7A7FB210708.text	D477D357D237A5267E93C7A7FB210708.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Caridina gracilirostris De Man 1892	<div><p>Caridina gracilirostris De Man, 1892</p><p>- Caridina gracilirostris De Man, 1892: 399, pl. 25, Figure 31a–d [type locality: River near Maros (approx. 5 ◦ 0.594 ′ S, 119 ◦ 35.381 ′ E), Sulawesi (Celebes), Indonesia];</p><p>- Caridina gracilirostris —Bright, 1979 [17]; 1989 [23]: 34, Table 1.</p><p>Material examined</p><p>PALAU: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.5701&amp;materialsCitation.latitude=7.5259333" title="Search Plazi for locations around (long 134.5701/lat 7.5259333)">Babeldaob Island</a>. USNM 104882: 1 ♂, cl 4.2 mm, 3 ♀ ovig., cl 4.6–5.7 mm and 1 juv. cl 2.8 mm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.5701&amp;materialsCitation.latitude=7.5259333" title="Search Plazi for locations around (long 134.5701/lat 7.5259333)">south fork of Arakitaoch river</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=134.5701&amp;materialsCitation.latitude=7.5259333" title="Search Plazi for locations around (long 134.5701/lat 7.5259333)">station 170-A</a>, 7 ◦ 31.556 ′ N, 134 ◦ 34.206 ′ E, 15 November 1956, coll. H.A. Fehlmann. USNM 172593: 1 ♂, cl 3.1 mm, Metengalakumer river, Station 1MR 4-03, 17 November 1977, coll. G. Bright. USNM 172594: 1 ♂, cl 3.5 mm, Ngerhekuu river, station 1NBK 4-10, 1 June 1978, coll. G. Bright.</p><p>Distribution</p><p>C. gracilirostris is widely distributed in the tropical and subtropical Indo-Pacific region:</p><p>Indonesia, the Philippines, India, Australia, the Solomon Islands, and Palau [43].</p><p>Habitat</p><p>In the lower course of streams under marine influence, very often in brackish water.</p><p>Remarks de Mazancourt et al. [43] redescribed this species in detail.</p></div>	https://treatment.plazi.org/id/D477D357D237A5267E93C7A7FB210708	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
D477D357D235A5257E93C36AFA55062B.text	D477D357D235A5257E93C36AFA55062B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atyidae De Haan 1849	<div><p>Key for studied species:</p><p>1.1 Pereiopods P1 and P2 carpus identically shaped. ...................................... ...2</p><p>1.2 Pereiopods P1 and P2 carpus differently shaped; P2 carpus is more elongated than that of P1. .........................................................................................3</p><p>2.1 Cephalothorax anterior border bearing a strong spine; straight rostrum rounded at its extremity with 2 to 6 small ventral teeth. ............................. Atyopsis spinipes</p><p>2.2 Cephalothorax anterior border acute bearing no spine; curved rostrum with 0 to 4 small ventral teeth. ............................................... Atyoida chacei sp. nov.</p><p>3.1 Eyes with cornea reduced to a small dark spot.. ... ... .. .... ... ... .. ..... Caridina rubella</p><p>3.2 Eyes with cornea normally developed.. ... ... .. .... ... ... .. .... ... ... ... .. .... ... ... ... ...4</p><p>4.1 Teeth absent on dorsalmarginof rostrum. ............................... Caridinatypus</p><p>4.2 More or less abundant teeth present on dorsal margin of rostrum.. ... .... .... ... ... ....5</p><p>5.1 Long stylocerite reaching or exceeding the first article of the antennal peduncle. .....6</p><p>5.2 Short stylocerite, failing toreach the firstarticle of the antennal peduncle.. ... .. .... ....8</p><p>6.1 Dorsal border of rostrum with 1–3 teeth located on the cephalothorax, posterior to the eye; P1 carpus 1.3–1.6 as long as wide.. ... ... .. .... ... ... .. .... ... ... ... .. .. C. lobocensis</p><p>6.2 Dorsal border of rostrum with 7–10 teeth located on the cephalothorax, posterior to the eye; P1 carpus 3.4–4.9 as longas wide. ....................................................7</p><p>7.1 Rostrum reaching to the base of the third segment of the antennular peduncle with 19–21 dorsalteeth ................................................ C. rintelenorum sp. nov.</p><p>7.2 Rostrum reaching beyond the end of the antennular peduncle with 21–26 dorsal teeth. .................................................................... C. cf. serratirostris</p><p>8.1 Short rostrum, curved downwards, 0.2–0.3 of cl, armed with 4–6 teeth on dorsal margin, 0 of them situated on carapace behind orbital margin. ........... C. ponapensis sp. nov.</p><p>8.2 Relatively long rostrum, 0.6–2.1 of cl, armed with 5–26 teeth on dorsal margin, 0–4 of them situated on carapace behind orbital margin. ......................................9</p><p>9.1 Uropodal diaeresis with 6–10 spiniform setae and a very long and upcurved rostrum with 5–9 dorsal teeth, widely spaced.. ................................... C. gracilirostris</p><p>9.2 Number of spiniform setae on uropodal diaeresis 8–15 and a long or short rostrum with 8–26 dorsalteeth, closelyset ........................................................10</p><p>10.1 Pre-anal carina with a prominent finger-like backward striking tooth. ........ C. appendiculata</p><p>10.2 Pre-analcarinaunarmed. ..................................................................11</p><p>11.1 P5 dactylus with a single terminal spine and a short distal propodus seta; posterior margin of the telson with intermediate setae longer or equal than lateral ones. ...................................................................... C. variabilirostris</p><p>11.2 P5 dactylus with two strong distal spines and a very long distal propodus seta; posterior margin of the telson with intermediate setae shorter or equal than lateral ones. ............................................................................. C. variabilis</p></div>	https://treatment.plazi.org/id/D477D357D235A5257E93C36AFA55062B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Mazancourt, Valentin de;Marquet, Gérard;Keith, Philippe	Mazancourt, Valentin de, Marquet, Gérard, Keith, Philippe (2024): An Integrative Taxonomic Revision of the Freshwater Atyid Shrimps (Crustacea: Decapoda: Caridea) of Micronesia. Diversity 16 (200): 1-31, DOI: 10.3390/d16040200, URL: https://doi.org/10.3390/d16040200
