taxonID	type	description	language	source
93CBC23C983C5630B5FBC8269D78242E.taxon	description	Figs 9, 18	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
93CBC23C983C5630B5FBC8269D78242E.taxon	diagnosis	Diagnosis. This species is shown for comparison with Euryobeidia xuei sp. nov. Diagnostic characters are provided under E. xuei sp. nov. A female paratype in excellent condition deposited in the ZFMK collection is figured here for the first time (Fig. 18). Moreover, an additional female of larger size and rather untypical pattern is shown (see Fig. 9), discovered under ZFMK material during the present study (identified by genitalia dissection, gen. prep. no. 2459 - DS).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
93CBC23C983C5630B5FBC8269D78242E.taxon	distribution	Distribution. China (Gansu, Sichuan, Shaanxi).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
06859937768E5FF6AE5E640871AC9B9E.taxon	description	Generic description. General appearance. Medium-sized ennomine moths, forewing length 17 – 27 mm. Included species are separated into the following two groups based on ground color: 1 st group, including the type-species E. languidata, with ground color white to grayish-white; 2 nd group orange, often fading to yellow over time. Wings with a large number of black, dark gray or dark brown spots, arranged in a similar pattern, with the exception of Euryobeidia yakushimensis stat. nov. Head. Antennae filiform in both sexes, flagellomeres of a short proximal part cylindrical, more distally they are laterally flattened, ventrally elongated, wedge-shaped, homogeneously covered with very short setae and with a pair of long, straight, spine-like setae, arising mid-laterally on either side near distal margin of each segment. Antennae of females similar, but thinner. Frons narrow, covered with slightly elongated or almost hair-like scales, the latter basally arranged around a tiny, central protrusion of the head-capsule; a concentric arrangement of scales on top of frons, between the bases of antennae, has been observed in some specimens of different species. Vertex covered with slightly longer and broader, distally dentate, obliquely erect scales. Labial palpi slender, roundly curved upwards, just reaching or protruding slightly beyond the frons, third joint small, but clearly visible. Proboscis rather short. Chaetosemata small, near eye margin. Thorax. Dorsum orange, yellow or grayish-yellow, typically with black dots: one on each patagium, two on each tegula, and two on mesothorax; slight variation occurs among species or individuals. Patagia and tegulae with lamellar, partly elongated scales, tegulae in addition with long hair-scales. Legs slender, pale yellow or orange, with a few dark gray or black dots. Index of spurs 0 - 2 - 4, hind tibia not dilated and without scent-brush (hair-pencil) in males. Forewing not or slightly elongate, arched at basal part of costa, apex angled, termen smoothly curved. Fovea absent. Hindwing with a large white area at the basal ⅔ – ¾ (except for E. tigratoides tigratoides and E. yakushimensis), and a broad, rarely narrow, yellow, submarginal band with multiple black streaks or dots. Marginal line with black dots at vein-ends, absent in forewing of 1 st group, absent or strongly reduced in species of 2 nd group. Apex of hindwing rounded, termen minutely concave between vein-ends, posterior margin slightly truncated from the end of vein 3 A to tornus and also on termen from vein-end of CuA 2 to tornus. Venation (Fig. 1). Forewing: costal area very broad at basal ⅔; Sc evenly curved, but rather abruptly bent upwards near the distal 1 / 5 (most clearly noticeable in E. tigratoides, see Fig. 1); R 1 arising from upper vein of cell rather close to the common stem of R 2 - 5; R 1 also often exhibits a distinct curvature opposite to the basal ends of R 2 and R 5 and a second curvature more distally and therefore reaches the costa closer to the apex; R 2, as a typical character for the tribe Baptini, arises from the common stem of R 3 - 5; stem R 2 - 5 arising on a rather large distance from anterior angle of cell (origin of M 1); M 2 from the middle of the discocellulars; CuA 1 widely separated from posterior angle of cell; CuP represented by a rather distinct fold, but with a very short and weak sclerotized vein near the base. Hindwing: Sc + R 1 running closely parallel but not anastomosing with upper vein of cell at base (a common character of most ennomine moths); Rs arising at a rather small distance from anterior angle of cell; M 2 absent; CuA 1 arising at a longer distance from posterior angle of cell; 3 A present. Pregenital abdomen. Abdomen densely covered with fine, whitish, orange, or pale yellow scales. Dorsal side usually with dark spots or transverse bands on each segment from T 1 to T 8, the spots vary in coloration and size between species and also individually, and rarely may even be absent in some or all segments (e. g. Euryobeidia tigratoides tigratoides). Laterally, a row of dark spots present, reduced or absent. Ventral side often with a small number of dark spots and patches of various size and shape, especially in the species-group with white or grayish-white ground color, fewer or even absent in the group of orange / yellow specimens. Integument with most tergites and sternites not conspicuously modified. T 1 narrower than T 2, appendages of intersegmental tergal phragma T 1 / T 2 very long, strap-like. The 8 th segment in males slightly elongate, broader than the 7 th segment, posteriorly slightly sclerotized. In females, the 7 th segment distinctly enlarged, 8 th segment very small, tergite T 8 rather membranous, with a cup-shaped, round or oval invagination of unknown function (See the black arrow on Fig. 50; visible on dorsal side, behind the right arm of lamella postvaginalis; so far only observed in E. languidata). Tympanal organs of moderate size, shallow, without lacinia. Setal comb on third sternite and sterno-tympanal process both absent, which is consistent with the absence of dilated hind-tibiae with scent-brushes. Coremata not developed. Male genitalia. Very heterogeneous, especially the uncus with various peculiar shapes among species. It may exhibit the shape of a bird’s head, densely covered with setae, with a small or larger beak-shaped apex, pointed in lateral view, and a narrow neck of differing length. In addition, large, plate-like appendages from “ head ” may be present. The base of the uncus mostly is a transverse straight or triangular bar, with lateral socii, covered with fine setae. Only one species (E. largeteaui) exhibits a rather unmodified uncus. Common characters are rare, e. g. the weak gnathos, reduced to fine, sclerotized lateral arms, not fused at middle, or the large tegumen, consisting of two narrow, elongate arms, which are rather common also in other genera of Baptini. In Euryobeidia, the tegumen sometimes is swollen distally and curved ventrad, in other species it is not swollen, but stronger distally and narrower caudally. Transtilla sinuous, broad, band-shaped, slightly to heavily sclerotized. Vinculum short and strong, fused laterally with tegumen. Saccus short, rounded. Juxta groove-like. Valva long and slender, with a well-developed cucullus, reaching from apex back to center of valva, ending with a small, setose knob. Valva distally strongly curved dorsad, apex with a smaller or larger, sometimes lobe-like protrusion, resulting in a more ventral notch. Costa smooth or slightly protruding or with a large, broad process. Base of sacculus often with a lamellar, oval or triangular projection, dorsally more or less dentate and variable in size among species. Aedeagus elongate, apically narrowed, and sclerotized ventrally, often with a pointed process of variable length (extremely long in E. largeteaui). Cornuti spine-like or replaced by sclerotized folds and patches of vesica of various shapes. Bulbus ejaculatorius shorter than the aedeagus shaft; proximal part (near aedeagus) tube-like and short; central part U-shaped, open ventrally; distal part cap-shaped, large, with a long rectangular extension distally (Figs 40, 45, 46, 49). Female genitalia. Ovipositor short, papillae anales slightly elongated, densely and shortly setose, tip rounded. Posterior apophyses long and narrow, anterior apophyses shorter, strong, ~ ⅓ – ½ the length of posterior apophyses, their bases dilated or narrow. A triangular sclerite present between the bases of posterior apophyses, its size and shape vary considerably between species and subspecies. Sterigma well developed, sclerotized; lamella antevaginalis spined on posterior margin and more strongly so laterally; lamella postvaginalis a sclerotized plate of different shape, consisting of two layers of plates, connected proximally. Introitus bursae strongly sclerotized, more or less twisted, usually placed asymmetrically on left side in ventral view (situated in the center in E. largeteaui only). Ductus seminalis arising close to the end of the short, narrow, strongly sclerotized posterior part of bursa copulatrix (ductus bursae of authors). Anterior part of bursa large, oval or pyriform, distal ½ – ¾ abundantly spined inside, remaining proximal portion membranous, without spines.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
06859937768E5FF6AE5E640871AC9B9E.taxon	distribution	Distribution. China, Korea, Japan, India, Nepal, Vietnam (new record).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
06859937768E5FF6AE5E640871AC9B9E.taxon	diagnosis	Diagnosis. Euryobeidia species are very similar in appearance to certain species and subspecies of the Obeidia - complex (Inoue, 2003) of genera (mainly of the genus Epobeidia Wehrli, 1939) and the genus Abraxas Leach, [1815], which probably serve as models in a mimicry relationship. However, they are easily distinguished from the previously mentioned genera by their less elongate, basally arched forewings and markedly different genitalic structures. Within the tribe Baptini, Euryobeidia species can be easily identified by the pattern of prominent dark spots on the wings and the distinctive genitalic structures, particularly the peculiar uncus of the male genitalia.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
4423D5E68DAA5785B2D61D86ED21263B.taxon	description	Figs 2 – 8, 38, 39, 50, 56	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
4423D5E68DAA5785B2D61D86ED21263B.taxon	diagnosis	Diagnosis. Images of adults and genitalia of the nominate subspecies are shown here for comparison with E. languidata incrassata stat. nov. and E. yakushimensis stat. nov. The diagnoses are given under each of the two taxa.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
4423D5E68DAA5785B2D61D86ED21263B.taxon	distribution	Distribution. Nepal (type-locality), India (Assam-Meghalaya), N. Vietnam, China (Sichuan, Yunnan, Jiangxi, Zhejiang, Fujian, Taiwan, Guangdong, Guangxi), Korea, Japan.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
0899FC5C5BAF53D697B673F7883E65C9.taxon	description	Figs 10 – 13, 40, 41, 53, 57	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
0899FC5C5BAF53D697B673F7883E65C9.taxon	diagnosis	Diagnosis. Euryobeidia languidata incrassata Xiang & Han, 2017 differs significantly from the nominate subspecies in the wing pattern being paler gray and a larger size, but the male genitalia of both subspecies are almost identical. The female genitalia of E. languidata incrassata, represented by a single specimen only, exhibit slight differences from those of the nominate subspecies, such as the size and shape of the lamella postvaginalis. The results of the molecular studies show that the genetic distance between them is 0.5 – 0.9 % (see Table 2).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
0899FC5C5BAF53D697B673F7883E65C9.taxon	distribution	Distribution. China (Hainan).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
3067910A681D551796EBDC636FA95F71.taxon	description	Figs 20 – 24, 59	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
3067910A681D551796EBDC636FA95F71.taxon	diagnosis	Diagnosis. This widespread and abundant species is presented here for comparison with E. tigratoides leopardiformis subsp. nov., due to their nearly identical appearance. Diagnostic characters are given under the description of the latter.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
3067910A681D551796EBDC636FA95F71.taxon	distribution	Distribution. China (Gansu, Zhejiang, Hubei, Jiangxi, Hunan, Fujian, Taiwan, Guangdong, Hong Kong, Guangxi, Sichuan, Chongqing, Guizhou), Vietnam.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
02884497086B5E3C85311167F1542E62.taxon	description	Figs 25, 26, 60, 61	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
02884497086B5E3C85311167F1542E62.taxon	diagnosis	Diagnosis. This species is also shown for comparison with E. tigratoides sp. nov. and E. tigratoides leopardiformis subsp. nov., particularly the latter. Diagnostic characters are provided under E. tigratoides leopardiformis subsp. nov.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
02884497086B5E3C85311167F1542E62.taxon	distribution	Distribution. China (Anhui, Zhejiang, Hubei, Hunan, Jiangxi, Fujian, Guangdong, Hong Kong, Guangxi, Sichuan).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
73973B9EF01E5F5080A5BE7A3C2CE449.taxon	description	Figs 14, 15, 44	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
73973B9EF01E5F5080A5BE7A3C2CE449.taxon	diagnosis	Diagnosis. This new species is nearly identical in habitus, pattern, and coloration to the sympatric Euryobeidia languidata incrassata Xiang & Han, but it is considerably smaller in size. Moreover, there are some extremely subtle differences. For example, the section near the inner margin of the forewing has almost no black streaks; the hindwing discal spot is very small, appearing as a tiny dot. However, with only one specimen, we cannot be certain if these are stable characters. In addition, the male genitalia of this species, featuring an elongated uncus and an extremely broad costal protrusion, are distinctly different from those of other congeneric species.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
73973B9EF01E5F5080A5BE7A3C2CE449.taxon	description	Description. The length of the forewing in the single male is ~ 20 mm. The wing pattern is almost identical to that of ssp. incrassata (see the redescription of E. languidata incrassata), except for the uncertain subtle differences already mentioned in the diagnosis. Male genitalia. Uncus long, subapical part (or stem) conspicuously elongated, slightly dilated dorsally, densely covered with setae, beak-like part long and stout. Socii absent or barely visible (the single type specimen). Gnathos weak, with a pair of short, fine sclerotized lateral arms, widely open or membranous in the middle. Tegumen moderately short (compared to most of other members of the genus), distally strongly curved ventrad. Transtilla long, broad, ribbon-like, slightly sclerotized. Valva broad, apex less curved dorsad (compared to most of congeneric species), ventral margin smooth, without notch. Costa with a markedly broad projection close to apex, dorsal edge faintly serrated in the middle. Cucullus well developed, from middle to apex of valva. Basal saccular process broad, lamellar, dorsal edge with minute denticles. Juxta strongly sclerotized, groove-like. Saccus short, rounded. Aedeagus slender, apically slightly tapering. Cornuti consisting of multiple irregularly joined sclerotized folds and patches. Female. Unknown.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
73973B9EF01E5F5080A5BE7A3C2CE449.taxon	distribution	Distribution. China (Hainan).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
73973B9EF01E5F5080A5BE7A3C2CE449.taxon	etymology	Etymology. The specific name is derived from the extremely broad costa of the male genitalia.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
33976CE5A1715DD9B8A88655DB644BD7.taxon	description	Figs 32 – 37, 46, 47, 54, 63, 64	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
33976CE5A1715DD9B8A88655DB644BD7.taxon	diagnosis	Diagnosis. Euryobeidia tigratoides is the only entirely orange / yellow species without a white pattern or area on the hindwing. This external character, together with its large body size, makes it very easy to distinguish from all other congeners. In addition to its distinct appearance, this species can also be readily differentiated from all other congeneric species (except for the next new subspecies to be described: E. tigratoides leopardiformis subsp. nov.) by the following two features of the male genitalia: 1. There is a protruding, rounded lobe bare of setae at the apex of valva, with a neighboring evident notch in E. tigratoides, while in other species, it is absent or inconspicuous. 2. The basal process of the sacculus is broader than in any other congeneric species.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
33976CE5A1715DD9B8A88655DB644BD7.taxon	description	Description. Forewing length 21.5 – 24.4 mm in males, 25.2 – 26.1 mm in females. Ground color orange or yellow after fading, densely covered with numerous dark spots. More detailed general features see the previous generic description. Head. Antennae filiform in both sexes, ventro-laterally with a pair of long, spine-like setae on each segment. Frons narrow, covered with smooth, narrow, light orange scales, with a small, rounded, central protrusion near the base. Labial palpus slightly extending beyond frons. Vertex covered with erect, lamellar, light orange scales. Chaetosemata small, near eye margin. Proboscis short. Thorax. Dorsum bright orange, two large separate black dots present on mesothorax. Patagia and tegulae bright orange, patagium with a small black spot in most individuals, tegula with a small black spot at base and a large black spot at middle. Legs orange, a few small black spots mainly on the base, middle and end of the femur and tibia segments. Index of spurs 0 - 2 - 4. Hind tibia not dilated, without scent-brush in males. Wings entirely orange, without white pattern. Forewing not elongated, arched at basal part of costa, apex angled, termen smoothly curved, fovea absent. Forewing scattered with numerous small streaks or spots on basal, costal and terminal areas; antemedial line represented by three large dark spots, the large streaked patch on the middle of the costal aera, extending from the costa down to near the cell-fold (weak or barely visible in few individuals); six separate dark spots present outside the discal spot, getting larger from upper to lower; discal spot appearing as a large dark dot; fringes matching the ground color, interspersed with black. Hindwing scattered with numerous tiny spots or streaks on basal and terminal areas; the first two spots of postmedial line always fused, the second and third pairs are typically separate, the two spots within the second pair or within the third pair, very close or even connected only in few individuals; discal spot large, rounded. Underside of wings almost identical to upperside, but slightly darker. Area of wing-coupling pale. Pregenital abdomen. Abdomen covered with fine, orange scales; dorsal dark spots fewer than those of other congeners, and inconspicuous or absent in some individuals. Tergites and sternites not conspicuously modified. The 8 th segment in males slightly elongate, broader than the 7 th segment, posteriorly slightly sclerotized, posterior edge slightly concave in the center. Tympanal organs of moderate size, without lacinia. Sterno-tympanal process, setal comb and coremata absent. Male genitalia. Uncus short, “ bird-headed ”, dorsal apical part markedly dilated, densely covered with setae, “ beak ” part pointed in lateral view, stem short, strongly curved dorsad. Socii small, with fine setae. Gnathos weak, with a pair of short, fine sclerotized lateral arms only. Transtilla long, broad, sclerotized, band-shaped. Tegumen large, with long, stout lateral arms, distally significantly swollen, curved ventrad. Valva slender, apex strongly curved dorsad at a right angle, with a large, round, non-setose extension of costa, forming a conspicuous notch ventrally. Costa narrow, smooth. Cucullus well developed, from apex reaching back to center of valva. Basal process of sacculus quite broad, lamellar, dorsal edge with minute denticles. Juxta broad, heavily sclerotized, groove-like. Saccus small, rounded. Aedeagus slender, apically slightly tapering and ridge-like. Cornutus small, with a central groove. Bulbus ejaculatorius shorter than the aedeagus shaft, with a rather large cap. Female genitalia. Ovipositor very short, papillae anales small, densely setose. Apophyses anteriores slightly shorter than apophyses posteriores, basal ¼ slightly broadened. A narrow, triangular sclerite present between the bases of posterior apophyses. Lamella antevaginalis well developed, with large irregular serrations. The ventral central plate of lamella postvaginalis large, broad at top, narrow at bottom, apical center slightly concave; dorsal layer with a pair of large, sclerotized, irregularly shaped, lateral projections. Introitus bursae slightly displaced to left side, strongly sclerotized. Posterior part of bursa fine, strongly sclerotized and twisted, connected to the ductus seminalis at middle, proximally swollen close to the bursa copulatrix. Anterior part of bursa pyriform, the distal ½ with many spines inside, the proximal ½ membranous, without spines.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
33976CE5A1715DD9B8A88655DB644BD7.taxon	distribution	Distribution. China (Hainan).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
33976CE5A1715DD9B8A88655DB644BD7.taxon	etymology	Etymology. The specific name is derived from its potential mimicry model, the nominotypical subspecies of Epobeidia tigrata (Guenée).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
AB238D9FAEBB5084B2BF5ABD3240BA44.taxon	description	Figs 27 – 31, 48, 49, 55	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
AB238D9FAEBB5084B2BF5ABD3240BA44.taxon	diagnosis	Diagnosis. This new subspecies is significantly smaller than the nominate subspecies and exhibits a strikingly distinct appearance, characterized by the basal ⅔ of the hindwing being white. However, the highly similar male and female genitalia and a minimal genetic divergence of 0.94 % (see Table 2) strongly indicate a close taxonomic relationship, thereby supporting the treatment of leopardiformis as a new subspecies of E. tigratoides. In addition to the markedly different wing patterns, the former can also be clearly distinguished from the latter by the following characters of male and female genitalia: the uncus is clearly longer with its stem straight or only slightly curved when observed from a lateral aspect, the spines on the posterior part of bursa are more concentrated compared to the nominate subspecies, and the lateral process of lamella postvaginalis is less sclerotized than in the nominate subspecies. Furthermore, Euryobeidia tigratoides leopardiformis closely resembles the sympatric E. largeteaui (Oberthür) in size, habitus, coloration, and wing pattern, rendering them nearly indistinguishable by appearance alone. Another sympatric species, E. quadrata Xiang & Han, also similar in appearance to the two species mentioned above, can be distinguished by a combination of characters: a white patch or area on the forewing (this single feature is also present in some individuals of E. largeteaui) and the nearly fused second pair of dots of the postmedial line on the hindwing. The new subspecies can be readily differentiated from E. largeteaui and E. quadrata by the following genitalic characters: 1. Uncus is small, bird-headed, while in E. largeteaui it is large, semicircular; in E. quadrata, it is also larger, with an elongated apex and an extremely dilated, somewhat square, flake-like protrusion on the dorsal side. 2. Basal process of sacculus is quite broad with minute denticles along the dorsal edge, whereas in both E. largeteaui and E. quadrata, it is narrow and lacks denticles. 3. The valval apex bears a conspicuous lobe and a neighboring notch, which is absent or inconspicuous in the other two species. 4. Apex of aedeagus is moderately long, slightly tapering, and ridged; in contrast, it is rather long and rod-like in E. largeteaui, and short and broad in E. quadrata. 5. Lamella antevaginalis is well-developed with large, irregular serrations, whereas in E. largeteaui, it is large and triangular with an upright sclerite in the center, and in E. quadrata, consists of two semicircular sclerites. 6. Lamella postvaginalis is rather large and M-shaped, with a pair of large, slightly sclerosed lateral processes, whereas it is much smaller in E. quadrata and quite small, barely visible, in E. largeteaui.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
AB238D9FAEBB5084B2BF5ABD3240BA44.taxon	description	Description. Forewing length 18 – 20 mm in males, 21 mm in the single female. Adults of E. tigratoides leopardiformis are almost identical to E. largeteaui in habitus, coloration, pattern and size, there are no consistent features to distinguish them. Typically, the six spots of postmedial line are fused in pairs, with the second and third pairs sometimes also slightly fused in certain specimens. However, the wing pattern observed in all four known type specimens of E. tigratoides leopardiformis is also found in specimens with nearly identical patterns within the extensive E. largeteaui collection. Currently, we believe that the two taxa cannot be accurately distinguished based on adult external morphological characters other than genitalia. Male and female genitalia. The male and female genitalia are strikingly similar to those of E. tigratoides tigratoides (see the previous description of E. tigratoides), with the following stable differences: the shaft of the uncus is longer, its stem is less curved, and its dorsal apical part is less dilated; The spines on the posterior part of bursa are sparser; the lateral process of lamella postvaginalis is less sclerotized.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
AB238D9FAEBB5084B2BF5ABD3240BA44.taxon	distribution	Distribution. China (Sichuan, Hubei).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
AB238D9FAEBB5084B2BF5ABD3240BA44.taxon	etymology	Etymology. The subspecific name is derived from its potential mimicry model, the subspecies leopardaria (Oberthür) of Epobeidia tigrata (Guenée).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
CE6E62A30DA452428B067634DCE4542A.taxon	description	Figs 19, 45, 58	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
CE6E62A30DA452428B067634DCE4542A.taxon	diagnosis	Diagnosis. Euryobeidia xuei is generally similar to E. languidata (Walker, 1862) in habitus, pattern, coloration and size, but can be distinguished from the latter on the basis of the shorter, less elongate wings, with apex and tornus more rounded and the compact, partly fused, very dark basal spotting, and a large, nearly fully fused, black discal spot forming a triangular shape in forewing which renders it more similar to E. ellipsoidea Xiang & Han, 2017. It differs from the latter and also from languidata by the complete or almost complete absence of a large spot or a dense group of smaller spots in the middle of the hind margin of the forewing. This feature that rarely occurs in other species and the extreme reduction of spots in the hindwings are distinctive for E. xuei. The structure of the male genitalia is quite similar to that of E. ellipsoidea, especially the shape of valva and the presence of a large, plate-like structure of the uncus, which indicates a rather close relationship of the two taxa. Distinguishing characters are the round, apical lobes of the valve costa which is markedly larger in xuei and the stalk connecting the central, beak-shaped part of uncus and the large dorsal plate is longer and narrower, the plate itself has a different shape. Moreover, the 3 % genetic distance based on molecular data separates this taxon from E. ellipsoidea, justifying its recognition as a distinct, but closely related species.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
CE6E62A30DA452428B067634DCE4542A.taxon	description	Description. Forewing length 19 – 21 mm in males. Similar to E. languidata, but wing-shape different: wings shorter, broader, apex and tornus of forewing and hindwing more rounded. Head. Antennae filiform, agreeing with generic description, shaft dorsally covered with silvery-gray scales. Labial palpus dark gray, with a few yellow scales at base of first segment, the latter with moderately long, obliquely upright scales. Second joint rather smooth, third joint very small, tapering. Frons narrow, covered with dark brownish-gray scales, those on dorsal ½ smooth, slightly elongated, basal ½ with a brush of more strongly elongated scales, covering a small, rounded protrusion of the integument. Vertex small, consisting of large, dark gray, obliquely upright scales, forming a triangle. Posterior of it and around the head runs a conspicuous, yellow collar. Chaetosemata present, near eye-margin, consisting of a few sensillae only. Thorax. Dorsum as described in the generic description, anteriorly yellow, posteriorly grayish yellow, with two very large, almost black spots behind each other. Patagia yellow with large, dark gray spots, tegulae with a yellow transverse band, a basal dark gray spot and a second spot and lighter gray hair-scales posteriorly. Legs yellow and yellowish gray, with some black dots. Tibia of hind-leg not dilated, without scent pencil. Wings with ground color clear white. Pattern of very dark, rather large, round spots, compact and partly fused near base of forewing. The more internal dots are suffused with brown scales, while the large apical patch and narrow bands along costa and anterior ⅔ of termen are dark silvery gray. A large spot in the middle of the hind margin absent, resulting in a broad, white band stretching obliquely through the forewing from near apex to hind margin. This seems to be the most obvious distinguishing element and is present in all known specimens. There are small yellow streaks near tornus. Fringe almost entirely black. Spots on hindwing reduced in size or absent, rendering the white area the largest of all species. Discal dot round, rather large. Base of hindwing with a group of very small dots. Yellow band along hindwing termen narrow, only sparsely dotted, fringe yellow with black, almost round dots. Underside with pattern very similar, but much paler. Venation agreeing with generic description, except that CuP is represented by a very faint fold, without a weakly sclerotized basal portion. Pregenital abdomen. Ground color of proximal ¾ of dorsal side pale yellowish gray, distal ¼ deeply yellow. All tergites with spots or patches of different sizes, shapes, and colors: T 1 pale gray, with a deep incision proximally in the middle. Spots from T 2 to T 8 almost black. T 2, T 3 smaller, almost semicircular, T 4 - T 6 rectangular, transverse bands (T 6 smaller), T 7 a very small spot, T 8 a pair of rather large, quadrate patches (see Figs 19, 58). Laterally a row of black, irregular spots, decreasing in size towards tip of abdomen. Lateral and ventral sides deep yellow, with moderately elongated scales and irregularly distributed black spots of different size and shape. Shape and number of these spots is due to ample variation. Integument without distinct variations, tergite 8 narrowly rectangular, sternite 8 much larger, laterally and distally rounded. Tympanal organs of moderate size, shallow, without lacinia. Setal comb on sternite 3 and sterno-tympanal process absent. Tergal phragma T 1 / T 2 long, strap-like. Male genitalia. Uncus with a central, beak-shaped part, carried by a long and narrow “ neck ” which combines it with the forked base and the semicircular connection with the tegumen. From the central part, a long and narrow, spined stalk arises which supports a large plate of unknown function. It is rounded posteriorly and tapering to a slightly narrower end anteriorly (Fig. 45). The gnathos consists of tiny, sclerotized lateral arms, which are widely separated from each other. Valves long and narrow, with the costa smooth, slightly protruding on both sides. Apex of valva strongly curved back, with a large, round, transparent, sclerotized plate (but hardly visible). Sacculus with a large, oval, basal plate, its margin irregularly indented and covered with minute spines. Aedeagus moderately long, the sclerotized distal ½ flat and terminating with a short spine at the rather broad apex. Vesica with two cornuti of different shape which are fused at one end. Female. Unknown.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
CE6E62A30DA452428B067634DCE4542A.taxon	distribution	Distribution. Vietnam, China (Yunnan).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
CE6E62A30DA452428B067634DCE4542A.taxon	etymology	Etymology. The specific epithet is honoring the renowned Chinese taxonomist Prof. Dayong Xue for his outstanding contributions to the study of Geometridae.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
E77EDDEA299E5505939A48BB72826139.taxon	description	Figs 16, 17, 42, 43, 51, 52	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
E77EDDEA299E5505939A48BB72826139.taxon	diagnosis	Diagnosis. This species is the only one with an almost unspotted wing pattern in the genus Euryobeidia, making it very easy to distinguish from other congeneric species. In terms of genitalia, this species is quite similar to E. languidata, indicating that they are rather closely related. However, there are still many stable differences between them, and they can be easily distinguished by the following characters: 1. Apex of the uncus is hardly dilated in E. yakushimensis, whereas it is distinctly inflated in E. languidata; 2. Costa of the male genitalia, with a small process in the middle in E. yakushimensis, while it is smooth, without process in E. languidata; 3. The triangular sclerite between the bases of the apophyses posteriores is distinctly narrower in E. yakushimensis than in E. languidata; 4. Lamella postvaginalis is smaller in E. yakushimensis than in E. languidata; 5. The spines on the posterior part of the bursa are sparser in E. yakushimensis than in E. languidata.	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
E77EDDEA299E5505939A48BB72826139.taxon	distribution	Distribution. Japan (Yakushima).	en	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
