identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
93CBC23C983C5630B5FBC8269D78242E.text	93CBC23C983C5630B5FBC8269D78242E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia ellipsoidea Xiang & Han 2017	<div><p>Euryobeidia ellipsoidea Xiang &amp; Han, 2017</p><p>Figs 9, 18</p><p>Euryobeidia ellipsoidea Xiang &amp; Han, 2017, Zootaxa 4317 (2): 372.</p><p>Type material examined.</p><p>For figures of the male holotype and a female paratype, their genitalia and detailed type material data from IZCAS and ZFMK, see Xiang et al. (2017).</p><p>Additional material examined.</p><p>China – Sichuan Province • 1 ♀; “ Chasseurs indigènes de Tâ-tsien-lou, Récolte de 1910 ”; Kangding; 1910; local collectors; coll. Charles Oberthür, ex coll. Wehrli, ZFMK .</p><p>Diagnosis.</p><p>This species is shown for comparison with Euryobeidia xuei sp. nov. Diagnostic characters are provided under E. xuei sp. nov. A female paratype in excellent condition deposited in the ZFMK collection is figured here for the first time (Fig. 18). Moreover, an additional female of larger size and rather untypical pattern is shown (see Fig. 9), discovered under ZFMK material during the present study (identified by genitalia dissection, gen. prep. no. 2459 - DS).</p><p>Distribution.</p><p>China (Gansu, Sichuan, Shaanxi).</p></div>	https://treatment.plazi.org/id/93CBC23C983C5630B5FBC8269D78242E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
06859937768E5FF6AE5E640871AC9B9E.text	06859937768E5FF6AE5E640871AC9B9E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia Fletcher 1979	<div><p>Euryobeidia Fletcher, 1979</p><p>Euryobeidia Wehrli, 1939, in Seitz, Gross-Schmett. Erde 4 (Suppl.): 269. Unavailable, type species not designated.</p><p>Euryobeidia Fletcher, 1979, The Generic Names of Moths of the World, 3: 84. Type species: Abraxas languidata Walker, 1862 .</p><p>Generic description.</p><p>General appearance. Medium-sized ennomine moths, forewing length 17–27 mm. Included species are separated into the following two groups based on ground color: 1 st group, including the type-species E. languidata, with ground color white to grayish-white; 2 nd group orange, often fading to yellow over time. Wings with a large number of black, dark gray or dark brown spots, arranged in a similar pattern, with the exception of Euryobeidia yakushimensis stat. nov. Head. Antennae filiform in both sexes, flagellomeres of a short proximal part cylindrical, more distally they are laterally flattened, ventrally elongated, wedge-shaped, homogeneously covered with very short setae and with a pair of long, straight, spine-like setae, arising mid-laterally on either side near distal margin of each segment. Antennae of females similar, but thinner. Frons narrow, covered with slightly elongated or almost hair-like scales, the latter basally arranged around a tiny, central protrusion of the head-capsule; a concentric arrangement of scales on top of frons, between the bases of antennae, has been observed in some specimens of different species. Vertex covered with slightly longer and broader, distally dentate, obliquely erect scales. Labial palpi slender, roundly curved upwards, just reaching or protruding slightly beyond the frons, third joint small, but clearly visible. Proboscis rather short. Chaetosemata small, near eye margin. Thorax. Dorsum orange, yellow or grayish-yellow, typically with black dots: one on each patagium, two on each tegula, and two on mesothorax; slight variation occurs among species or individuals. Patagia and tegulae with lamellar, partly elongated scales, tegulae in addition with long hair-scales. Legs slender, pale yellow or orange, with a few dark gray or black dots. Index of spurs 0-2 - 4, hind tibia not dilated and without scent-brush (hair-pencil) in males. Forewing not or slightly elongate, arched at basal part of costa, apex angled, termen smoothly curved. Fovea absent. Hindwing with a large white area at the basal ⅔ – ¾ (except for E. tigratoides tigratoides and E. yakushimensis), and a broad, rarely narrow, yellow, submarginal band with multiple black streaks or dots. Marginal line with black dots at vein-ends, absent in forewing of 1 st group, absent or strongly reduced in species of 2 nd group. Apex of hindwing rounded, termen minutely concave between vein-ends, posterior margin slightly truncated from the end of vein 3 A to tornus and also on termen from vein-end of CuA 2 to tornus. Venation (Fig. 1). Forewing: costal area very broad at basal ⅔; Sc evenly curved, but rather abruptly bent upwards near the distal 1 / 5 (most clearly noticeable in E. tigratoides, see Fig. 1); R 1 arising from upper vein of cell rather close to the common stem of R 2-5; R 1 also often exhibits a distinct curvature opposite to the basal ends of R 2 and R 5 and a second curvature more distally and therefore reaches the costa closer to the apex; R 2, as a typical character for the tribe Baptini, arises from the common stem of R 3-5; stem R 2-5 arising on a rather large distance from anterior angle of cell (origin of M 1); M 2 from the middle of the discocellulars; CuA 1 widely separated from posterior angle of cell; CuP represented by a rather distinct fold, but with a very short and weak sclerotized vein near the base. Hindwing: Sc + R 1 running closely parallel but not anastomosing with upper vein of cell at base (a common character of most ennomine moths); Rs arising at a rather small distance from anterior angle of cell; M 2 absent; CuA 1 arising at a longer distance from posterior angle of cell; 3 A present. Pregenital abdomen. Abdomen densely covered with fine, whitish, orange, or pale yellow scales. Dorsal side usually with dark spots or transverse bands on each segment from T 1 to T 8, the spots vary in coloration and size between species and also individually, and rarely may even be absent in some or all segments (e. g. Euryobeidia tigratoides tigratoides). Laterally, a row of dark spots present, reduced or absent. Ventral side often with a small number of dark spots and patches of various size and shape, especially in the species-group with white or grayish-white ground color, fewer or even absent in the group of orange / yellow specimens. Integument with most tergites and sternites not conspicuously modified. T 1 narrower than T 2, appendages of intersegmental tergal phragma T 1 / T 2 very long, strap-like. The 8 th segment in males slightly elongate, broader than the 7 th segment, posteriorly slightly sclerotized. In females, the 7 th segment distinctly enlarged, 8 th segment very small, tergite T 8 rather membranous, with a cup-shaped, round or oval invagination of unknown function (See the black arrow on Fig. 50; visible on dorsal side, behind the right arm of lamella postvaginalis; so far only observed in E. languidata). Tympanal organs of moderate size, shallow, without lacinia. Setal comb on third sternite and sterno-tympanal process both absent, which is consistent with the absence of dilated hind-tibiae with scent-brushes. Coremata not developed.</p><p>Male genitalia. Very heterogeneous, especially the uncus with various peculiar shapes among species. It may exhibit the shape of a bird’s head, densely covered with setae, with a small or larger beak-shaped apex, pointed in lateral view, and a narrow neck of differing length. In addition, large, plate-like appendages from “ head ” may be present. The base of the uncus mostly is a transverse straight or triangular bar, with lateral socii, covered with fine setae. Only one species ( E. largeteaui) exhibits a rather unmodified uncus. Common characters are rare, e. g. the weak gnathos, reduced to fine, sclerotized lateral arms, not fused at middle, or the large tegumen, consisting of two narrow, elongate arms, which are rather common also in other genera of Baptini . In Euryobeidia, the tegumen sometimes is swollen distally and curved ventrad, in other species it is not swollen, but stronger distally and narrower caudally. Transtilla sinuous, broad, band-shaped, slightly to heavily sclerotized. Vinculum short and strong, fused laterally with tegumen. Saccus short, rounded. Juxta groove-like. Valva long and slender, with a well-developed cucullus, reaching from apex back to center of valva, ending with a small, setose knob. Valva distally strongly curved dorsad, apex with a smaller or larger, sometimes lobe-like protrusion, resulting in a more ventral notch. Costa smooth or slightly protruding or with a large, broad process. Base of sacculus often with a lamellar, oval or triangular projection, dorsally more or less dentate and variable in size among species. Aedeagus elongate, apically narrowed, and sclerotized ventrally, often with a pointed process of variable length (extremely long in E. largeteaui). Cornuti spine-like or replaced by sclerotized folds and patches of vesica of various shapes. Bulbus ejaculatorius shorter than the aedeagus shaft; proximal part (near aedeagus) tube-like and short; central part U-shaped, open ventrally; distal part cap-shaped, large, with a long rectangular extension distally (Figs 40, 45, 46, 49).</p><p>Female genitalia. Ovipositor short, papillae anales slightly elongated, densely and shortly setose, tip rounded. Posterior apophyses long and narrow, anterior apophyses shorter, strong, ~ ⅓ – ½ the length of posterior apophyses, their bases dilated or narrow. A triangular sclerite present between the bases of posterior apophyses, its size and shape vary considerably between species and subspecies. Sterigma well developed, sclerotized; lamella antevaginalis spined on posterior margin and more strongly so laterally; lamella postvaginalis a sclerotized plate of different shape, consisting of two layers of plates, connected proximally. Introitus bursae strongly sclerotized, more or less twisted, usually placed asymmetrically on left side in ventral view (situated in the center in E. largeteaui only). Ductus seminalis arising close to the end of the short, narrow, strongly sclerotized posterior part of bursa copulatrix (ductus bursae of authors). Anterior part of bursa large, oval or pyriform, distal ½ – ¾ abundantly spined inside, remaining proximal portion membranous, without spines.</p><p>Distribution.</p><p>China, Korea, Japan, India, Nepal, Vietnam (new record).</p><p>Diagnosis.</p><p>Euryobeidia species are very similar in appearance to certain species and subspecies of the Obeidia - complex (Inoue, 2003) of genera (mainly of the genus Epobeidia Wehrli, 1939) and the genus Abraxas Leach, [1815], which probably serve as models in a mimicry relationship. However, they are easily distinguished from the previously mentioned genera by their less elongate, basally arched forewings and markedly different genitalic structures. Within the tribe Baptini, Euryobeidia species can be easily identified by the pattern of prominent dark spots on the wings and the distinctive genitalic structures, particularly the peculiar uncus of the male genitalia.</p></div>	https://treatment.plazi.org/id/06859937768E5FF6AE5E640871AC9B9E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
4423D5E68DAA5785B2D61D86ED21263B.text	4423D5E68DAA5785B2D61D86ED21263B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia languidata (Walker 1862)	<div><p>Euryobeidia languidata (Walker, 1862)</p><p>Figs 2–8, 38, 39, 50, 56</p><p>Abraxas languidata Walker, 1862, List Specimens lepid. Insects Colln Br. Mus. 24: 1122. Holotype, Nepal. (NHMUK)</p><p>Euryobeidia languidata: Wehrli 1939, in Seitz, Gross-Schmett. Erde 4 (Suppl.): 269. Unavailable.</p><p>Euryobeidia languidata: Fletcher 1979, The Generic Names of Moths of the World 3: 84; Stüning, D. 2000, Moths of Nepal, part 6: 110; Xiang et al. 2017, Zootaxa 4317 (2): 371.</p><p>Type material examined.</p><p>Holotype. Nepal • ♀; ‘ Nepal’; Hardwicke Bequest; NHMUK. (Abdomen lost; see Fig. 2)</p><p>Additional material examined.</p><p>Nepal • 1 ♀; Kathmandu Valley, Godavari; 1600–1800 m; 5 Jun. 1967; leg. Dierl-Schacht; ZSM . India • 1 ♂; Khasis; May 1896; Nat. Coll.; coll. Wehrli, ZFMK . China – Fujian Province • 1 ♂; Quanzhou City, Dehua County, Jiuxianshan; 1200 m; 12 May 2024; C. L. Huang &amp; Z. Peng leg.; GenBank no.: PQ 083540; gen. prep. no. CRICATAS 00283; CRICATAS – Taiwan • 1 ♂; Nantou, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=120.916664&amp;materialsCitation.latitude=23.966667" title="Search Plazi for locations around (long 120.916664/lat 23.966667)">Sang-Gan nr. Pu-li</a>; 23°58'N, 120°55'E; Apr. – Jul. 2008; local coll.; ZFMK . Vietnam • 1 ♂; N. Vietnam, Vinhu, Tam Dao; 900 m; 29 May 1997; B. Tanaka leg.; CKY . Japan – • 1 ♂, 1 ♀; ‘ Japon, Wileman, 1898 ’; male gen. prep. Wehrli no. 5525, female gen. prep. no. 2460 - DS; ZFMK – Tokyo • 1 ♂; Tokio; 17 Jun. 1910; gen. prep. Wehrli. no. 5517; ZFMK • 1 ♂; Abiko, Chiba Pref. nr. Tokyo; May 1914; ZFMK • 1 ♂; Chiyoda, Kokyo; 21 May 1998; Y. Kishida leg.; gen. prep. no. RS 8877; CRS / NIAES (shown by Sato 2011, pl. 1-015 - 9) • 1 ♀; Chiyoda, Kokyo; 11 Jun. 1998; Y. Kishida leg.; gen. prep. no. RS 8878; CRS / NIAES (shown by Sato 2011, pl. 1-015 - 10) – Niigata Prefecture • 1 ♀; Niigata City, Nishikan-ku, Iwamuro, Parking of Yahiko; 23 Jul. 2007; R. Sato leg.; CRS / NIAES (shown by Sato 2011, pl. 1-015 - 11) .</p><p>Remarks.</p><p>Euryobeidia languidata obviously is extremely rare at its type locality, Nepal. The female holotype at NHMUK is the only specimen among a collection of ~ 50 specimens, half of them originating from NE India, Assam-Meghalaya region, the second half from Japan. The volumes 1–5 of the “ Moths of Nepal ” - series do not mention it and several thousand specimens from many localities in Nepal at ZFMK definitely do not contain it. The doubt about the correctness of the type locality has arisen, but was answered by the fact that a second specimen was found at the ZSM collection, Munich, a female collected near Kathmandu, the specimen mentioned above (Stüning 2000: 110, Moths of Nepal, vol. 6). Unfortunately, this and a male from Khasi Hills, Meghalaya could be studied by images only (Figs 2–5). However, in our experience they will not affect our taxonomic conclusions when investigated more closely in future.</p><p>Diagnosis.</p><p>Images of adults and genitalia of the nominate subspecies are shown here for comparison with E. languidata incrassata stat. nov. and E. yakushimensis stat. nov. The diagnoses are given under each of the two taxa.</p><p>Distribution.</p><p>Nepal (type-locality), India (Assam-Meghalaya), N. Vietnam, China (Sichuan, Yunnan, Jiangxi, Zhejiang, Fujian, Taiwan, Guangdong, Guangxi), Korea, Japan.</p><p>Remarks.</p><p>Euryobeidia languidata is the type-species of Euryobeidia, designated by Fletcher (1979), and a member of the white or grayish-white group of species (1 st Group). It is the species with the widest area of distribution, from east Nepal in the west to north Japan in the east, with southernmost occurrence on the island of Hainan ( ssp. incrassata). Within China, it appears to be primarily distributed along the southeastern coast. More to the north, there appears to be a gap between northern Zhejiang and South Korea, but this region is mostly low-lying plains with high population density, extensive agriculture and hardly any forests, obviously unsuitable for the existence of this species.</p></div>	https://treatment.plazi.org/id/4423D5E68DAA5785B2D61D86ED21263B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
0899FC5C5BAF53D697B673F7883E65C9.text	0899FC5C5BAF53D697B673F7883E65C9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia languidata subsp. incrassata Xiang & Han 2017	<div><p>Euryobeidia languidata incrassata Xiang &amp; Han stat. nov.</p><p>Figs 10–13, 40, 41, 53, 57</p><p>Euryobeidia incrassata Xiang &amp; Han, 2017, Zootaxa 4317 (2): 372 (part).</p><p>Type material examined.</p><p>Holotype. China – Hainan Province • ♂; Lingshui, Diaoluoshan; 8 May 1984; coll. Gu Maobin; IOZ-CAS slide no. Geom- 7235; IZCAS . Paratypes. China – Hainan Province • 1 ♂; same collection data as for holotype; IZCAS • 1 ♂; Baisha, Nankai, Nanmaola; 1261 m; 12–14 May 2009; coll. Chen Fuqiang; IZCAS .</p><p>Additional material examined.</p><p>China – Hainan Province • 3 ♂♂, 1 ♀; Lingshui, Diaoluoshan; 922 m; 10 May 2023; Bo Liu leg.; GenBank no.: PQ 083537; gen. prep. nos. CRICATAS 00220, CRICATAS 00223; CRICATAS / ZFMK • 1 ♂; same locality as for preceding; 07–12 May 2024; Bo Liu &amp; Wei Yan leg.; GenBank no.: PQ 083539; gen. prep. no. CRICATAS 00222; CRICATAS • 1 ♂; Lingshui, Diaoluoshan; 997 m; 21 Apr. 2025; Bo Liu &amp; Wei Yan leg.; CRICATAS .</p><p>Diagnosis.</p><p>Euryobeidia languidata incrassata Xiang &amp; Han, 2017 differs significantly from the nominate subspecies in the wing pattern being paler gray and a larger size, but the male genitalia of both subspecies are almost identical. The female genitalia of E. languidata incrassata, represented by a single specimen only, exhibit slight differences from those of the nominate subspecies, such as the size and shape of the lamella postvaginalis. The results of the molecular studies show that the genetic distance between them is 0.5–0.9 % (see Table 2).</p><p>Redescription.</p><p>Forewing length 24.9–28.0 mm in males, 26.5 mm in the single female (19–23 mm in E. languidata languidata). Ground color grayish-white, wings on upper- and underside covered with numerous pale gray spots and streaks (more numerous, larger, dark gray spots in E. l. languidata). More detailed general characters see the generic description. Head. Antennae filiform in both sexes, laterally with a pair of long, spine-like setae on each segment. Frons pale yellow, with a small, rounded protrusion in the middle near the base. Labial palpus pale yellow, slightly extending beyond frons. Vertex covered with erect, pale yellow scales. Chaetosemata weak. Proboscis short. Thorax. Dorsum pale yellow, two large separate black dots on mesothorax. Patagia and tegulae pale yellow, a small black dot at center of each patagium, a small black dot at base and a moderately large black dot at middle of each tegula. Legs pale yellow with a few dark spots. Index of spurs 0-2 - 4. Hind tibia not dilated, without scent-brush in males. Forewing not elongate, rather broadly triangular and arched at basal part of costa, apex angled, termen smoothly curved. Fovea absent. Forewing grayish-white, covered with lots of dark gray spots or streaks, especially on wing base, costa, inner margin and outer margin; antemedial line appearing as three dark gray spots on the upper and lower veins of cell and 2 A respectively; six dark gray spots present outside the discal spot, positioned on veins R, M 1, M 3, CuA 1, CuA 2, and 2 A, the first two spots commonly fused, the second two spots on M 3 and CuA 1, located further outwards, almost fused with the distal band; discal spot large, dark gray, elongate-oval, weakly curved inwards; terminal area with a band of tiny dark gray streaks, getting broader and more dense towards apex; fringes dark gray. Hindwing grayish-white, tinged with dark gray and pale yellow terminally, yellow part mixed with short gray transverse streaks; postmedial line similar to that of forewing, but spots smaller; discal spot present as a small dark gray dot; terminal line appearing as a row of black spots at the end of veins; fringes black corresponding to those spots, pale yellow between veins. Underside almost identical to upperside, but paler. Pregenital abdomen. Abdomen yellow, dorsal side lightly grayish-yellow, with dark gray spots at middle of each segment (a pair of two spots on first tergite which are fused in the middle on second tergite and get gradually more fused and reduced in size until tergite eight); lateral side with a row of black spots, and ventral side scattered with irregular dark gray spots. Tergites and sternites not conspicuously modified. The 8 th segment in males slightly elongate, broader than the 7 th segment, posteriorly slightly sclerotized, posterior edge straight. Tympanal organs of moderate size, without lacinia. Sterno-tympanal process, setal comb, and coremata absent.</p><p>Male genitalia. Uncus moderately long, “ bird-headed ”, dorsal apical portion distinctly expanded semicircularly, densely covered with setae, “ beak ” portion small, pointed in lateral view, stem portion slightly curved, basal portion weak. Socii small, with fine setae. Gnathos weak, with short, narrow, sclerotized lateral arms which are separated by a large (membranous) opening at middle. Tegumen large, with long, stout lateral arms, distally swollen, curved ventrad. Transtilla long, partly broadened, ribbon-like, slightly sclerotized. Valva slender, apex strongly curved dorsad, with a small, faintly visible notch more ventrally. Costa narrow, smooth, apex slightly protruding. Cucullus well developed, from apex reaching back to center of valva. Basal saccular process broad, lamellar, dorsal edge serrate. Juxta broad, heavily sclerotized, groove-like. Saccus rounded. Aedeagus slender, curved, apically tapering, with an arrow-shaped tip in lateral view. Cornuti consisting of multiple irregularly joined sclerotized folds and patches.</p><p>Female genitalia. Ovipositor rather short, papillae anales small, densely setose. Apophyses anteriores short, about ½ length of the very thin apophyses posteriores, basal two thirds of the former strongly broadened. A broad, triangular sclerite with a small tip present between the bases of apophyses posteriores. Lamella antevaginalis well developed, with many large irregular serrations. The ventral central plate of lamella postvaginalis strongly sclerotized, large, I-shaped; dorsal layer less sclerotized. Introitus bursae placed asymmetrically on left side, strongly sclerotized. Posterior part of bursa strongly sclerotized, barely twisted, with a few spines on left side, proximally swollen close to the bursa copulatrix, distally tapering, with a small colliculum-like structure adjacent to ductus seminalis. Anterior part of bursa almost oval, the distal ¾ with many spines inside and the proximal ¼ membranous, without spines.</p><p>Distribution.</p><p>China (Hainan).</p><p>Remarks.</p><p>This subspecies was originally described as Euryobeidia incrassata by Xiang et al. (2017). In their paper, the adult holotype, and the male genitalia of a paratype were illustrated, with the treatment of the new species primarily based on the distinctive genitalia of the latter, which set it apart from all other species in the genus. However, a detailed examination of the male genitalia of the holotype, two paratypes and non-types revealed that they were almost identical with those of E. languidata languidata (Walker, 1862) and incrassata therefore should be treated as a distinct subspecies of the latter. The only paratype of E. incrassata with distinctive male genitalia is redescribed in the present study as a new species, E. supercostata sp. nov. In addition, the authors examined all remaining paratypes and all freshly collected non-type males of E. languidata incrassata stat. nov. by dissecting the genitalia or brushing off scales to reveal the uncus and the characteristic part of the valva, ensuring that they were not confused with the almost identical in appearance and sympatric E. supercostata sp. nov.</p></div>	https://treatment.plazi.org/id/0899FC5C5BAF53D697B673F7883E65C9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
3067910A681D551796EBDC636FA95F71.text	3067910A681D551796EBDC636FA95F71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia largeteaui (Oberthur 1884)	<div><p>Euryobeidia largeteaui (Oberthür, 1884)</p><p>Figs 20–24, 59</p><p>Rhyparia largeteaui Oberthür, 1884, Études ent. 10: 32, pl. 1, fig. 5. holotype ♂, China: Kouy-Tchéou, in coll. ZFMK .</p><p>Euryobeidia largeteaui: Wehrli 1939, in Seitz, Gross-Schmett. Erde 4 (Suppl.): 269. Unavailable.</p><p>Euryobeidia largeteaui: Fletcher 1979, The Generic Names of Moths of the World 3: 84; Xiang et al. 2017, Zootaxa 4317 (2): 373.</p><p>Type material examined.</p><p>Holotype. China – Guizhou Province • ♂; Kouy-Tchéou (Guizhou); Abbé Largeteau leg.; gen. slide Wehrli no. 5578; ex coll. Oberthür, ex coll. Wehrli, ZFMK. (left forewing absent; see Fig. 20)</p><p>Additional material examined.</p><p>China – Hunan Province • 20 ♂♂, 12 ♀♀; Huaihua City, Xupu County, Taojinping Township, Shannaoao Village, Mountain woodland; 800 m; Jul. 2022; Chao Dai leg.; GenBank nos: PQ 083536, PQ 083541; CRICATAS, CRICATAS 00251 to CRICATAS 00282 . – Hubei Province • 4 ♂♂, 1 ♀; W. Hubei Province, Wufeng, Yizhuxiang Mt.; 1560 m; Jun. 1998; Wang &amp; Li leg.; ZFMK . Further 26 ♂♂ ♀♀ in ZFMK from Zhejiang, Hunan, Fujian, Guangdong, mainly ex coll. Wehrli, Oberthür, Höne. Vietnam – • 1 ♂; N. Vietnam, Lao Cai, Sa Pa; 1500 m; 25–28 May 1997; B. Tanak leg.; CKY • 6 ♂♂; N. Vietnam, Cha-pa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.76667&amp;materialsCitation.latitude=22.25" title="Search Plazi for locations around (long 103.76667/lat 22.25)">Mt. Fan-Si-Pan</a>; 22°15'N, 103°46'E; 1600–1800 m; leg. Sinjaev &amp; Simonov; ZFMK .</p><p>Diagnosis.</p><p>This widespread and abundant species is presented here for comparison with E. tigratoides leopardiformis subsp. nov., due to their nearly identical appearance. Diagnostic characters are given under the description of the latter.</p><p>Distribution.</p><p>China (Gansu, Zhejiang, Hubei, Jiangxi, Hunan, Fujian, Taiwan, Guangdong, Hong Kong, Guangxi, Sichuan, Chongqing, Guizhou), Vietnam.</p><p>Remarks.</p><p>A specimen from Baoxing, Sichuan, China, identified by Xiang et al. (2017) as E. largeteaui, is described in the present study as a new subspecies of E. tigratoides sp. nov., and is designated as the holotype.</p></div>	https://treatment.plazi.org/id/3067910A681D551796EBDC636FA95F71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
02884497086B5E3C85311167F1542E62.text	02884497086B5E3C85311167F1542E62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia quadrata Xiang & Han 2017	<div><p>Euryobeidia quadrata Xiang &amp; Han, 2017</p><p>Figs 25, 26, 60, 61</p><p>Euryobeidia quadrata Xiang &amp; Han, 2017, Zootaxa 4317 (2): 374.</p><p>Type material examined.</p><p>The description was based on a large series of male and female specimens, the holotype designated from Zhejiang. Detailed type material data from IZCAS and ZFMK, see Xiang et al. (2017).</p><p>Additional material examined.</p><p>China – Hunan Province • 1 ♂; Huaihua City, Xupu County, Taojinping Township, Shannaoao Village, Mountain woodland; 800 m; Jul. 2022; Chao Dai leg.; GenBank no.: PQ 083535; CRICATAS, CRICATAS 00250 .</p><p>Diagnosis.</p><p>This species is also shown for comparison with E. tigratoides sp. nov. and E. tigratoides leopardiformis subsp. nov., particularly the latter. Diagnostic characters are provided under E. tigratoides leopardiformis subsp. nov.</p><p>Distribution.</p><p>China (Anhui, Zhejiang, Hubei, Hunan, Jiangxi, Fujian, Guangdong, Hong Kong, Guangxi, Sichuan).</p></div>	https://treatment.plazi.org/id/02884497086B5E3C85311167F1542E62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
73973B9EF01E5F5080A5BE7A3C2CE449.text	73973B9EF01E5F5080A5BE7A3C2CE449.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia supercostata Liu, Stuning & Han 2025	<div><p>Euryobeidia supercostata Liu, Stüning &amp; Han sp. nov.</p><p>Figs 14, 15, 44</p><p>Euryobeidia incrassata Xiang &amp; Han, 2017, Zootaxa 4317 (2): 372 (part).</p><p>Type material.</p><p>Holotype. China – Hainan Province • ♂; Lingshui, Diaoluoshan; 8 May 1984; coll. Gu Maobin; IOZ-CAS slide no. Geom- 4140; IZCAS .</p><p>Diagnosis.</p><p>This new species is nearly identical in habitus, pattern, and coloration to the sympatric Euryobeidia languidata incrassata Xiang &amp; Han, but it is considerably smaller in size. Moreover, there are some extremely subtle differences. For example, the section near the inner margin of the forewing has almost no black streaks; the hindwing discal spot is very small, appearing as a tiny dot. However, with only one specimen, we cannot be certain if these are stable characters. In addition, the male genitalia of this species, featuring an elongated uncus and an extremely broad costal protrusion, are distinctly different from those of other congeneric species.</p><p>Description.</p><p>The length of the forewing in the single male is ~ 20 mm. The wing pattern is almost identical to that of ssp. incrassata (see the redescription of E. languidata incrassata), except for the uncertain subtle differences already mentioned in the diagnosis.</p><p>Male genitalia. Uncus long, subapical part (or stem) conspicuously elongated, slightly dilated dorsally, densely covered with setae, beak-like part long and stout. Socii absent or barely visible (the single type specimen). Gnathos weak, with a pair of short, fine sclerotized lateral arms, widely open or membranous in the middle. Tegumen moderately short (compared to most of other members of the genus), distally strongly curved ventrad. Transtilla long, broad, ribbon-like, slightly sclerotized. Valva broad, apex less curved dorsad (compared to most of congeneric species), ventral margin smooth, without notch. Costa with a markedly broad projection close to apex, dorsal edge faintly serrated in the middle. Cucullus well developed, from middle to apex of valva. Basal saccular process broad, lamellar, dorsal edge with minute denticles. Juxta strongly sclerotized, groove-like. Saccus short, rounded. Aedeagus slender, apically slightly tapering. Cornuti consisting of multiple irregularly joined sclerotized folds and patches.</p><p>Female. Unknown.</p><p>Distribution.</p><p>China (Hainan).</p><p>Etymology.</p><p>The specific name is derived from the extremely broad costa of the male genitalia.</p><p>Remarks.</p><p>The holotype of Euryobeidia supercostata sp. nov. was originally designated as a paratype of E. incrassata Xiang &amp; Han, 2017 . However, our examination of the holotype revealed that E. incrassata should be treated as a distinct subspecies of E. languidata (see E. languidata incrassata stat. nov.).</p></div>	https://treatment.plazi.org/id/73973B9EF01E5F5080A5BE7A3C2CE449	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
33976CE5A1715DD9B8A88655DB644BD7.text	33976CE5A1715DD9B8A88655DB644BD7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia tigratoides Liu, Stuning & Han 2025	<div><p>Euryobeidia tigratoides Liu, Stüning &amp; Han sp. nov.</p><p>Figs 32–37, 46, 47, 54, 63, 64</p><p>Type material.</p><p>Holotype. China – Hainan Province • ♂; Lingshui, Diaoluoshan; 922 m; 01–03 Apr. 2024; Bo Liu, Wei Lin &amp; Miaofeng Xu leg.; CRICATAS / IZCAS, CRICATAS 00230 . Paratypes (19 ♂♂, 6 ♀♀). China – Hainan Province • 2 ♂♂, 3 ♀♀; same locality as for holotype; 20 Apr. 2023; Bo Liu leg.; gen. prep. nos. CRICATAS 00243, CRICATAS 00244; CRICATAS / IZCAS / ZFMK • 2 ♂♂; same locality as for holotype; 10 May 2023; Bo Liu leg.; GenBank no. PQ 083534; gen. prep. no. CRICATAS 00247; CRICATAS / IZCAS • 7 ♂♂, 1 ♀; same collection data as for holotype • 7 ♂♂; same locality as for holotype; 07–12 May 2024; Bo Liu &amp; Wei Yan leg.; GenBank no.: PQ 083538; CRICATAS, IZCAS / ZFMK • 1 ♂; Lingshui, Diaoluoshan; 997 m; 21 Apr. 2025; Bo Liu &amp; Wei Yan leg.; CRICATAS • 2 ♀; Lingshui, Diaoluoshan; 974 m; 10 Jun. 2025; Bo Liu &amp; Wei Yan leg.; CRICATAS .</p><p>Diagnosis.</p><p>Euryobeidia tigratoides is the only entirely orange / yellow species without a white pattern or area on the hindwing. This external character, together with its large body size, makes it very easy to distinguish from all other congeners. In addition to its distinct appearance, this species can also be readily differentiated from all other congeneric species (except for the next new subspecies to be described: E. tigratoides leopardiformis subsp. nov.) by the following two features of the male genitalia: 1. There is a protruding, rounded lobe bare of setae at the apex of valva, with a neighboring evident notch in E. tigratoides, while in other species, it is absent or inconspicuous. 2. The basal process of the sacculus is broader than in any other congeneric species.</p><p>Description.</p><p>Forewing length 21.5–24.4 mm in males, 25.2–26.1 mm in females. Ground color orange or yellow after fading, densely covered with numerous dark spots. More detailed general features see the previous generic description. Head. Antennae filiform in both sexes, ventro-laterally with a pair of long, spine-like setae on each segment. Frons narrow, covered with smooth, narrow, light orange scales, with a small, rounded, central protrusion near the base. Labial palpus slightly extending beyond frons. Vertex covered with erect, lamellar, light orange scales. Chaetosemata small, near eye margin. Proboscis short. Thorax. Dorsum bright orange, two large separate black dots present on mesothorax. Patagia and tegulae bright orange, patagium with a small black spot in most individuals, tegula with a small black spot at base and a large black spot at middle. Legs orange, a few small black spots mainly on the base, middle and end of the femur and tibia segments. Index of spurs 0-2 - 4. Hind tibia not dilated, without scent-brush in males. Wings entirely orange, without white pattern. Forewing not elongated, arched at basal part of costa, apex angled, termen smoothly curved, fovea absent. Forewing scattered with numerous small streaks or spots on basal, costal and terminal areas; antemedial line represented by three large dark spots, the large streaked patch on the middle of the costal aera, extending from the costa down to near the cell-fold (weak or barely visible in few individuals); six separate dark spots present outside the discal spot, getting larger from upper to lower; discal spot appearing as a large dark dot; fringes matching the ground color, interspersed with black. Hindwing scattered with numerous tiny spots or streaks on basal and terminal areas; the first two spots of postmedial line always fused, the second and third pairs are typically separate, the two spots within the second pair or within the third pair, very close or even connected only in few individuals; discal spot large, rounded. Underside of wings almost identical to upperside, but slightly darker. Area of wing-coupling pale. Pregenital abdomen. Abdomen covered with fine, orange scales; dorsal dark spots fewer than those of other congeners, and inconspicuous or absent in some individuals. Tergites and sternites not conspicuously modified. The 8 th segment in males slightly elongate, broader than the 7 th segment, posteriorly slightly sclerotized, posterior edge slightly concave in the center. Tympanal organs of moderate size, without lacinia. Sterno-tympanal process, setal comb and coremata absent.</p><p>Male genitalia. Uncus short, “ bird-headed ”, dorsal apical part markedly dilated, densely covered with setae, “ beak ” part pointed in lateral view, stem short, strongly curved dorsad. Socii small, with fine setae. Gnathos weak, with a pair of short, fine sclerotized lateral arms only. Transtilla long, broad, sclerotized, band-shaped. Tegumen large, with long, stout lateral arms, distally significantly swollen, curved ventrad. Valva slender, apex strongly curved dorsad at a right angle, with a large, round, non-setose extension of costa, forming a conspicuous notch ventrally. Costa narrow, smooth. Cucullus well developed, from apex reaching back to center of valva. Basal process of sacculus quite broad, lamellar, dorsal edge with minute denticles. Juxta broad, heavily sclerotized, groove-like. Saccus small, rounded. Aedeagus slender, apically slightly tapering and ridge-like. Cornutus small, with a central groove. Bulbus ejaculatorius shorter than the aedeagus shaft, with a rather large cap.</p><p>Female genitalia. Ovipositor very short, papillae anales small, densely setose. Apophyses anteriores slightly shorter than apophyses posteriores, basal ¼ slightly broadened. A narrow, triangular sclerite present between the bases of posterior apophyses. Lamella antevaginalis well developed, with large irregular serrations. The ventral central plate of lamella postvaginalis large, broad at top, narrow at bottom, apical center slightly concave; dorsal layer with a pair of large, sclerotized, irregularly shaped, lateral projections. Introitus bursae slightly displaced to left side, strongly sclerotized. Posterior part of bursa fine, strongly sclerotized and twisted, connected to the ductus seminalis at middle, proximally swollen close to the bursa copulatrix. Anterior part of bursa pyriform, the distal ½ with many spines inside, the proximal ½ membranous, without spines.</p><p>Distribution.</p><p>China (Hainan).</p><p>Etymology.</p><p>The specific name is derived from its potential mimicry model, the nominotypical subspecies of Epobeidia tigrata (Guenée) .</p></div>	https://treatment.plazi.org/id/33976CE5A1715DD9B8A88655DB644BD7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
AB238D9FAEBB5084B2BF5ABD3240BA44.text	AB238D9FAEBB5084B2BF5ABD3240BA44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia tigratoides subsp. leopardiformis Liu, Stuning & Han 2025	<div><p>Euryobeidia tigratoides leopardiformis Liu, Stüning &amp; Han subsp. nov.</p><p>Figs 27–31, 48, 49, 55</p><p>Euryobeidia largeteaui: Xiang et al. 2017, Zootaxa 4317 (2): 374 (part).</p><p>Type material.</p><p>Holotype. China – Sichuan Province • ♂; Baoxing County, Dashuigou; 1591 m; 1–5 Aug. 2016; Le Cui leg.; gen. prep. no. Geom- 04555; IZCAS, IOZ LEP M 22920 . Paratypes. China – Hubei Province • 2 ♂♂, 1 ♀; W. Hubei Province, Wufeng, Yizhuxiang Mt.; 1560 m; Jun. 1998, Wang &amp; Li leg.; gen. prep. nos. 2449 - DS, 2450 - DS; ZFMK .</p><p>Diagnosis.</p><p>This new subspecies is significantly smaller than the nominate subspecies and exhibits a strikingly distinct appearance, characterized by the basal ⅔ of the hindwing being white. However, the highly similar male and female genitalia and a minimal genetic divergence of 0.94 % (see Table 2) strongly indicate a close taxonomic relationship, thereby supporting the treatment of leopardiformis as a new subspecies of E. tigratoides . In addition to the markedly different wing patterns, the former can also be clearly distinguished from the latter by the following characters of male and female genitalia: the uncus is clearly longer with its stem straight or only slightly curved when observed from a lateral aspect, the spines on the posterior part of bursa are more concentrated compared to the nominate subspecies, and the lateral process of lamella postvaginalis is less sclerotized than in the nominate subspecies.</p><p>Furthermore, Euryobeidia tigratoides leopardiformis closely resembles the sympatric E. largeteaui (Oberthür) in size, habitus, coloration, and wing pattern, rendering them nearly indistinguishable by appearance alone. Another sympatric species, E. quadrata Xiang &amp; Han, also similar in appearance to the two species mentioned above, can be distinguished by a combination of characters: a white patch or area on the forewing (this single feature is also present in some individuals of E. largeteaui) and the nearly fused second pair of dots of the postmedial line on the hindwing. The new subspecies can be readily differentiated from E. largeteaui and E. quadrata by the following genitalic characters: 1. Uncus is small, bird-headed, while in E. largeteaui it is large, semicircular; in E. quadrata, it is also larger, with an elongated apex and an extremely dilated, somewhat square, flake-like protrusion on the dorsal side. 2. Basal process of sacculus is quite broad with minute denticles along the dorsal edge, whereas in both E. largeteaui and E. quadrata, it is narrow and lacks denticles. 3. The valval apex bears a conspicuous lobe and a neighboring notch, which is absent or inconspicuous in the other two species. 4. Apex of aedeagus is moderately long, slightly tapering, and ridged; in contrast, it is rather long and rod-like in E. largeteaui, and short and broad in E. quadrata . 5. Lamella antevaginalis is well-developed with large, irregular serrations, whereas in E. largeteaui, it is large and triangular with an upright sclerite in the center, and in E. quadrata, consists of two semicircular sclerites. 6. Lamella postvaginalis is rather large and M-shaped, with a pair of large, slightly sclerosed lateral processes, whereas it is much smaller in E. quadrata and quite small, barely visible, in E. largeteaui .</p><p>Description.</p><p>Forewing length 18–20 mm in males, 21 mm in the single female. Adults of E. tigratoides leopardiformis are almost identical to E. largeteaui in habitus, coloration, pattern and size, there are no consistent features to distinguish them. Typically, the six spots of postmedial line are fused in pairs, with the second and third pairs sometimes also slightly fused in certain specimens. However, the wing pattern observed in all four known type specimens of E. tigratoides leopardiformis is also found in specimens with nearly identical patterns within the extensive E. largeteaui collection. Currently, we believe that the two taxa cannot be accurately distinguished based on adult external morphological characters other than genitalia.</p><p>Male and female genitalia. The male and female genitalia are strikingly similar to those of E. tigratoides tigratoides (see the previous description of E. tigratoides), with the following stable differences: the shaft of the uncus is longer, its stem is less curved, and its dorsal apical part is less dilated; The spines on the posterior part of bursa are sparser; the lateral process of lamella postvaginalis is less sclerotized.</p><p>Distribution.</p><p>China (Sichuan, Hubei).</p><p>Etymology.</p><p>The subspecific name is derived from its potential mimicry model, the subspecies leopardaria (Oberthür) of Epobeidia tigrata (Guenée) .</p></div>	https://treatment.plazi.org/id/AB238D9FAEBB5084B2BF5ABD3240BA44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
CE6E62A30DA452428B067634DCE4542A.text	CE6E62A30DA452428B067634DCE4542A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia xuei Liu, Stuning & Han 2025	<div><p>Euryobeidia xuei Liu, Stüning &amp; Han sp. nov.</p><p>Figs 19, 45, 58</p><p>Type material.</p><p>Holotype. Vietnam – • ♂; North Vietnam, Cha-pa, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=103.76667&amp;materialsCitation.latitude=22.25" title="Search Plazi for locations around (long 103.76667/lat 22.25)">Mt. Fan-si-pan</a>; 22°15'N, 103°46'E; 1500–1800 m; 10 Jun. – 6 Jul. 1994; leg. V. Sinjaev &amp; einh. Sammler; lux (light trap); gen. prep. no. 2458 - DS; ZFMK . Paratype. Vietnam – • 1 ♂; same collection data as for holotype; ZFMK .</p><p>Additional material examined.</p><p>China – Yunnan Province • 1 ♂; Yuxi, Zhenyuan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.264&amp;materialsCitation.latitude=24.277" title="Search Plazi for locations around (long 101.264/lat 24.277)">Qianjiazhai, Ailaoshan</a>; 24.277°N, 101.264°E; 2200 m; 5 Aug. 2011; Kitching &amp; Ashton leg.; KIZCAS, ARB 00027908 • 1 ♂ or ♀ (image only); Diqing, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.96534&amp;materialsCitation.latitude=27.358805" title="Search Plazi for locations around (long 98.96534/lat 27.358805)">Weixi</a>; 27.358805°N, 98.965343°E; 2720 m; 01 Aug. 2024; Photographed by Fan Gao .</p><p>Diagnosis.</p><p>Euryobeidia xuei is generally similar to E. languidata (Walker, 1862) in habitus, pattern, coloration and size, but can be distinguished from the latter on the basis of the shorter, less elongate wings, with apex and tornus more rounded and the compact, partly fused, very dark basal spotting, and a large, nearly fully fused, black discal spot forming a triangular shape in forewing which renders it more similar to E. ellipsoidea Xiang &amp; Han, 2017 . It differs from the latter and also from languidata by the complete or almost complete absence of a large spot or a dense group of smaller spots in the middle of the hind margin of the forewing. This feature that rarely occurs in other species and the extreme reduction of spots in the hindwings are distinctive for E. xuei .</p><p>The structure of the male genitalia is quite similar to that of E. ellipsoidea, especially the shape of valva and the presence of a large, plate-like structure of the uncus, which indicates a rather close relationship of the two taxa. Distinguishing characters are the round, apical lobes of the valve costa which is markedly larger in xuei and the stalk connecting the central, beak-shaped part of uncus and the large dorsal plate is longer and narrower, the plate itself has a different shape. Moreover, the 3 % genetic distance based on molecular data separates this taxon from E. ellipsoidea, justifying its recognition as a distinct, but closely related species.</p><p>Description.</p><p>Forewing length 19–21 mm in males. Similar to E. languidata, but wing-shape different: wings shorter, broader, apex and tornus of forewing and hindwing more rounded.</p><p>Head. Antennae filiform, agreeing with generic description, shaft dorsally covered with silvery-gray scales. Labial palpus dark gray, with a few yellow scales at base of first segment, the latter with moderately long, obliquely upright scales. Second joint rather smooth, third joint very small, tapering. Frons narrow, covered with dark brownish-gray scales, those on dorsal ½ smooth, slightly elongated, basal ½ with a brush of more strongly elongated scales, covering a small, rounded protrusion of the integument. Vertex small, consisting of large, dark gray, obliquely upright scales, forming a triangle. Posterior of it and around the head runs a conspicuous, yellow collar. Chaetosemata present, near eye-margin, consisting of a few sensillae only. Thorax. Dorsum as described in the generic description, anteriorly yellow, posteriorly grayish yellow, with two very large, almost black spots behind each other. Patagia yellow with large, dark gray spots, tegulae with a yellow transverse band, a basal dark gray spot and a second spot and lighter gray hair-scales posteriorly. Legs yellow and yellowish gray, with some black dots. Tibia of hind-leg not dilated, without scent pencil. Wings with ground color clear white. Pattern of very dark, rather large, round spots, compact and partly fused near base of forewing. The more internal dots are suffused with brown scales, while the large apical patch and narrow bands along costa and anterior ⅔ of termen are dark silvery gray. A large spot in the middle of the hind margin absent, resulting in a broad, white band stretching obliquely through the forewing from near apex to hind margin. This seems to be the most obvious distinguishing element and is present in all known specimens. There are small yellow streaks near tornus. Fringe almost entirely black. Spots on hindwing reduced in size or absent, rendering the white area the largest of all species. Discal dot round, rather large. Base of hindwing with a group of very small dots. Yellow band along hindwing termen narrow, only sparsely dotted, fringe yellow with black, almost round dots. Underside with pattern very similar, but much paler. Venation agreeing with generic description, except that CuP is represented by a very faint fold, without a weakly sclerotized basal portion. Pregenital abdomen. Ground color of proximal ¾ of dorsal side pale yellowish gray, distal ¼ deeply yellow. All tergites with spots or patches of different sizes, shapes, and colors: T 1 pale gray, with a deep incision proximally in the middle. Spots from T 2 to T 8 almost black. T 2, T 3 smaller, almost semicircular, T 4 - T 6 rectangular, transverse bands (T 6 smaller), T 7 a very small spot, T 8 a pair of rather large, quadrate patches (see Figs 19, 58). Laterally a row of black, irregular spots, decreasing in size towards tip of abdomen. Lateral and ventral sides deep yellow, with moderately elongated scales and irregularly distributed black spots of different size and shape. Shape and number of these spots is due to ample variation. Integument without distinct variations, tergite 8 narrowly rectangular, sternite 8 much larger, laterally and distally rounded. Tympanal organs of moderate size, shallow, without lacinia. Setal comb on sternite 3 and sterno-tympanal process absent. Tergal phragma T 1 / T 2 long, strap-like.</p><p>Male genitalia. Uncus with a central, beak-shaped part, carried by a long and narrow “ neck ” which combines it with the forked base and the semicircular connection with the tegumen. From the central part, a long and narrow, spined stalk arises which supports a large plate of unknown function. It is rounded posteriorly and tapering to a slightly narrower end anteriorly (Fig. 45). The gnathos consists of tiny, sclerotized lateral arms, which are widely separated from each other. Valves long and narrow, with the costa smooth, slightly protruding on both sides. Apex of valva strongly curved back, with a large, round, transparent, sclerotized plate (but hardly visible). Sacculus with a large, oval, basal plate, its margin irregularly indented and covered with minute spines. Aedeagus moderately long, the sclerotized distal ½ flat and terminating with a short spine at the rather broad apex. Vesica with two cornuti of different shape which are fused at one end.</p><p>Female. Unknown.</p><p>Distribution.</p><p>Vietnam, China (Yunnan).</p><p>Etymology.</p><p>The specific epithet is honoring the renowned Chinese taxonomist Prof. Dayong Xue for his outstanding contributions to the study of Geometridae .</p></div>	https://treatment.plazi.org/id/CE6E62A30DA452428B067634DCE4542A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
E77EDDEA299E5505939A48BB72826139.text	E77EDDEA299E5505939A48BB72826139.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Euryobeidia yakushimensis Inoue 1976	<div><p>Euryobeidia yakushimensis Inoue, 1976 stat. nov.</p><p>Figs 16, 17, 42, 43, 51, 52</p><p>Euryobeidia languidata yakushimensis Inoue, 1976, Tinea 10 (2): 20. Holotype, Yakushima Isl., S. Japan (NHMUK) [images examined] .</p><p>Material examined.</p><p>Japan – Kagoshima Pref. • 1 ♂; Yakushima Island, Miyanoura, Siratani; alt. 600 m; 18–20 Jun. 1993; M. Owada leg.; gen. prep. no. RS 7496; CRS / NIAES (shown by Sato 2011: pl. 1-015 - 12) • 1 ♀; Yakushima Island, Mt. Toimodake; alt. 1016 m; 26 Jun. 2008; Y. Kubota leg.; gen. prep. no. RS 8879; CRS / NIAES (shown by Sato 2011: pl. 1-015 - 13) • 1 ♂; Yakushima Island, Yakusugi Land; alt. 1000–1300 m; 12 Jul. 2008; Y. Kubota leg.; gen. prep. no. RS 7411; CRS / NIAES • 1 ♂; same locality and collector as for preceding; 16 Aug. 2009; gen. prep. no. RS 7407; CRS / NIAES • 1 ♀; same locality and collector as for preceding; 5 Jul. 2008; gen. prep. no. RS 7412; CRS / NIAES • 1 ♀; same locality and collector as for preceding; 5 Jul. 2009; gen. prep. no. RS 7808; CRS / NIAES .</p><p>Diagnosis.</p><p>This species is the only one with an almost unspotted wing pattern in the genus Euryobeidia, making it very easy to distinguish from other congeneric species. In terms of genitalia, this species is quite similar to E. languidata, indicating that they are rather closely related. However, there are still many stable differences between them, and they can be easily distinguished by the following characters: 1. Apex of the uncus is hardly dilated in E. yakushimensis, whereas it is distinctly inflated in E. languidata; 2. Costa of the male genitalia, with a small process in the middle in E. yakushimensis, while it is smooth, without process in E. languidata; 3. The triangular sclerite between the bases of the apophyses posteriores is distinctly narrower in E. yakushimensis than in E. languidata; 4. Lamella postvaginalis is smaller in E. yakushimensis than in E. languidata; 5. The spines on the posterior part of the bursa are sparser in E. yakushimensis than in E. languidata .</p><p>Redescription.</p><p>Wingspan 35–41 mm in males, 40–42 mm in females. Forewing almost entirely covered with large dark gray patches, the grayish-white ground color remaining only in narrow bands along the position of postmedial line or forming a few small, patches on discal cell or antemedial line. Hindwing similar to forewing, with a yellow marginal band intermingled with several small, dark gray transverse streaks and spots. Underside almost identical to upperside, but slightly darker. Male and female genitalia. The morphology of the genitalia is quite similar to that of Euryobeidia languidata (see description of E. languidata incrassata), with the following different features: Uncus fine, stick-like, apex rounded, bent ventrad, dorsal apical portion barely dilated. Apical lobe-like protrusion and ventral notch of valva are more conspicuous. Costa exhibits a distinctive small protruding structure in the center. The triangular sclerite between the bases of the apophyses posteriores is narrower and the lamella postvaginalis is smaller compared to E. languidata . Basal part of apophyses anteriores weakly broadened. The spines on the posterior part of bursa are sparser than in E. languidata . Anterior part of bursa almost oval, the upper ⅔ with numerous spines inside and the lower ⅓ membranous (the membranous part is smaller in E. languidata, but probably variable in size).</p><p>Distribution.</p><p>Japan (Yakushima).</p><p>Remarks.</p><p>Sato (2011) proposed that E. languidata yakushimensis should be treated as a distinct species based on his comparative study of E. languidata languidata and E. languidata yakushimensis from Japan. In the present study, we additionally include specimens from continental China and Vietnam, as well as the new subspecies E. languidata incrassata from Hainan, in our comparative analysis. The results show that in addition to significant differences in wing pattern, there are also specific differences in male and female genitalia (see the previous diagnosis and description). We agree with Sato’s comments on E. languidata yakushimensis and elevate it to species rank.</p></div>	https://treatment.plazi.org/id/E77EDDEA299E5505939A48BB72826139	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Liu, Bo;Stüning, Dieter;Han, Hongxiang	Liu, Bo, Stüning, Dieter, Han, Hongxiang (2025): Further taxonomic studies of the mimetic genus Euryobeidia Fletcher, 1979 (Lepidoptera, Geometridae, Ennominae, Baptini), with descriptions of four new taxa and two status changes. ZooKeys 1260: 313-343, DOI: 10.3897/zookeys.1260.157773
