taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
EB49A1E08582554484349144B85A6521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1361116	https://doi.org/10.3897/zse.101.153548.figure2	Figure 2. Holotype of Phoxinus ayukensis sp. nov. (SL 54.3 mm, IBIW _ FS _ 501, female). A. Live appearance; B. General appearance of the preserved specimen; C. Radiograph.	Figure 2. Holotype of Phoxinus ayukensis sp. nov. (SL 54.3 mm, IBIW _ FS _ 501, female). A. Live appearance; B. General appearance of the preserved specimen; C. Radiograph.	2025-06-25	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.		Zenodo	biologists	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.			
EB49A1E08582554484349144B85A6521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1361117	https://doi.org/10.3897/zse.101.153548.figure3	Figure 3. Live coloration of female and male of Phoxinus ayukensis sp. nov. (paratypes, IBIW _ FS _ 502) from type locality (Ayuk Stream) and P. adagumicus, another congeneric species inhabiting the Kuban River basin (female with eggs; 44.4960 ° N, 38.4330 ° E; 8 May 2022; IBIW _ FS _ 338) from Zybza River. Side and ventral views are of the same specimens. Photo by Oleg Artaev.	Figure 3. Live coloration of female and male of Phoxinus ayukensis sp. nov. (paratypes, IBIW _ FS _ 502) from type locality (Ayuk Stream) and P. adagumicus, another congeneric species inhabiting the Kuban River basin (female with eggs; 44.4960 ° N, 38.4330 ° E; 8 May 2022; IBIW _ FS _ 338) from Zybza River. Side and ventral views are of the same specimens. Photo by Oleg Artaev.	2025-06-25	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.		Zenodo	biologists	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.			
EB49A1E08582554484349144B85A6521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1361122	https://doi.org/10.3897/zse.101.153548.figure8	Figure 8. Some morphological features of Phoxinus ayukensis sp. nov. A. Example of squamation of middle part of body of an alizarin-stained specimen (colors changed): a diagonal line marks the direction of scale counting between first unbranched rays of dorsal and anal fins (bda), above (all) and below (bll) lateral line (bda = 36, all = 21, and bll = 14 scales); B. Ventral view of pelvic fins, genital and anal openings in females and males; C. Dentition of pharyngeal bones: double-row formula 1.5–4.1 (observed in 10 % of specimens); D. Ventral view of an alizarin-stained specimen showing 3 rd-type scalation on the breast and belly and reduction of ray numbers to seven in both pelvic fins. Scale bar: 0.5 mm (C).	Figure 8. Some morphological features of Phoxinus ayukensis sp. nov. A. Example of squamation of middle part of body of an alizarin-stained specimen (colors changed): a diagonal line marks the direction of scale counting between first unbranched rays of dorsal and anal fins (bda), above (all) and below (bll) lateral line (bda = 36, all = 21, and bll = 14 scales); B. Ventral view of pelvic fins, genital and anal openings in females and males; C. Dentition of pharyngeal bones: double-row formula 1.5–4.1 (observed in 10 % of specimens); D. Ventral view of an alizarin-stained specimen showing 3 rd-type scalation on the breast and belly and reduction of ray numbers to seven in both pelvic fins. Scale bar: 0.5 mm (C).	2025-06-25	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.		Zenodo	biologists	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.			
EB49A1E08582554484349144B85A6521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1361118	https://doi.org/10.3897/zse.101.153548.figure4	Figure 4. Habitat of Phoxinus ayukensis sp. nov.: A. Puddle in the bed of Ayuk Stream, sampling place of the type material (44.4756 ° N, 39.0194 ° E, 19 July 2024, photograph by Oleg Artaev); B. One of a series of waterfalls that restrict the distribution of P. ayukensis sp. nov. (44.4749 ° N, 39.0209 ° E, 9 March 2025, photograph by Andrey Ostapenko).	Figure 4. Habitat of Phoxinus ayukensis sp. nov.: A. Puddle in the bed of Ayuk Stream, sampling place of the type material (44.4756 ° N, 39.0194 ° E, 19 July 2024, photograph by Oleg Artaev); B. One of a series of waterfalls that restrict the distribution of P. ayukensis sp. nov. (44.4749 ° N, 39.0209 ° E, 9 March 2025, photograph by Andrey Ostapenko).	2025-06-25	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.		Zenodo	biologists	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.			
EB49A1E08582554484349144B85A6521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1361119	https://doi.org/10.3897/zse.101.153548.figure5	Figure 5. Map of Phoxinus species distribution in the Northwestern Caucasus (based on own data).	Figure 5. Map of Phoxinus species distribution in the Northwestern Caucasus (based on own data).	2025-06-25	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.		Zenodo	biologists	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.			
EB49A1E08582554484349144B85A6521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1361120	https://doi.org/10.3897/zse.101.153548.figure6	Figure 6. Dendrograms (UPGMA): A. Average values of 41 morphometric characters for each species with bootstrap support on each node (Euclidean distance); B. Eight external meristic characters based on squared Mahalanobis distances defined by discriminant functions in DFA. In both cases, the difference was statistically significant between all species compared (p < 0.005). A schematic map of species ranges in the center, with colors corresponding to the species names.	Figure 6. Dendrograms (UPGMA): A. Average values of 41 morphometric characters for each species with bootstrap support on each node (Euclidean distance); B. Eight external meristic characters based on squared Mahalanobis distances defined by discriminant functions in DFA. In both cases, the difference was statistically significant between all species compared (p < 0.005). A schematic map of species ranges in the center, with colors corresponding to the species names.	2025-06-25	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.		Zenodo	biologists	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.			
EB49A1E08582554484349144B85A6521.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1361121	https://doi.org/10.3897/zse.101.153548.figure7	Figure 7. PCA of Phoxinus spp. based on eight meristic characters and loading plot showing how strongly each character influences principal components.	Figure 7. PCA of Phoxinus spp. based on eight meristic characters and loading plot showing how strongly each character influences principal components.	2025-06-25	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.		Zenodo	biologists	Artaev, Oleg N.;Turbanov, Ilya S.;Bolotovskiy, Aleksey A.;Gandlin, Alexander A.;Levin, Boris A.			
