identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B8D9ABD05E0E50B5AF4B1EAA6DA9AFFB.text	B8D9ABD05E0E50B5AF4B1EAA6DA9AFFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agonum (Europhilus) thoreyi Dejean 1828	<div><p>22. Agonum (Europhilus) thoreyi Dejean, 1828</p><p>Fig. 4 C</p><p>Habitat and general distribution.</p><p>Agonum thoreyi is a Holarctic species (Arndt et al. 2011). The species is stenotopic in reeds (Arndt et al. 2011).</p><p>Material examine.</p><p>• Rethymnon: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.358&amp;materialsCitation.latitude=35.35103" title="Search Plazi for locations around (long 24.358/lat 35.35103)">Petres River mouth</a>, 35.351030°N, 24.358000°E, 2 m elev., 9.VII.2015 – 2.XI.2015, 1 spm, leg. Pavlou Chr (NHMC) .</p><p>Comments.</p><p>Agonum thoreyi is a widespread, cosmopolitan species that has been reported from Greece for a first time by Schmidt and Liebherr (2009), while this is the first record of A. thoreyi in Crete. The species was found in a northern small river mouth near a sand dune system that is regularly crowded for recreational activities.</p></div>	https://treatment.plazi.org/id/B8D9ABD05E0E50B5AF4B1EAA6DA9AFFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
179FED983C7455B9A7DA4592B3327710.text	179FED983C7455B9A7DA4592B3327710.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Agonum (Olisares) viridicupreum (Goeze 1777)	<div><p>23. Agonum (Olisares) viridicupreum (Goeze, 1777)</p><p>Fig. 4 D</p><p>Habitat and general distribution.</p><p>Agonum viridicupreum has a wide distribution in West Palaearctic (Arndt et al. 2011). The species prefers open wet habitats with warm temperatures (meadows, open wetlands, fens and floodplains) (Arndt et al. 2011; Drees et al. 2011). It exhibits some tolerance to saline environments (Arndt et al. 2011) and has fluctuations in its range, driven by climate change (Drees et al. 2011).</p><p>Material examined.</p><p>• Heraklion: Dikti Mountain, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.4508&amp;materialsCitation.latitude=35.072395" title="Search Plazi for locations around (long 25.4508/lat 35.072395)">Omalos Viannou plateau</a>, in a temporary wetland, 35.072394°N, 25.450799°E, 1334 m elev., 16.X.2014, handpicking, 1 spm, leg. Pavlou Chr. (NHMC) ; • same data, but 16.X.2014 – 8.12.2014, 974 spms; • same data, but 9.V.2015 – 23.VII.2015, 87 spms .</p><p>Comments.</p><p>Agonum viridicupreum is reported in Greece from the Greek mainland (excluding Peloponnese) and Ionian islands (Arndt et al. 2011), but no further records exist of A. viridicupreum in Crete. The species was found in a temporary pond with grass and wetland vegetation ( Juncus spp. etc.), on a mountain plateau (Omalos Viannou, Dikti Mountain). Perimetrical to the pond, the plateau forms a mixed dehesa-like forest with Quercus coccifera, Crataegus and Acer . This combination of a wet meadow in a warm environment fits the description of the species’ habitat provided by Drees et al. (2011) who also stress the species’ tendency to expand its range towards the northern areas of Palaearctic. Based on the attention given to the southern limits of A. viridicupreum ’ s distribution by Drees et al. (2011), our records could be useful for a better understanding of the species’ range expansion-contraction dynamics.</p></div>	https://treatment.plazi.org/id/179FED983C7455B9A7DA4592B3327710	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
A5320399ACF25BC597497800854D07E8.text	A5320399ACF25BC597497800854D07E8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anaulacus (Aephnidius) ruficornis (Chaudoir 1850)	<div><p>11. Anaulacus (Aephnidius) ruficornis (Chaudoir, 1850)</p><p>Figs 3 B, 5 D</p><p>Habitat and general distribution.</p><p>A. ruficornis occurs in Near and Middle East (Bousquet 2017 a), barely reaching Europe, only from its north-eastern region bordering with Asia (e. g. south Russia) (Bousquet 2017 a). This rare (Austin et al. 2008) species dwells in wetlands and coastal habitats with fresh or saline water (Assmann et al. 2015).</p><p>Material examined.</p><p>• Chania: Frangokastello wetland, 35.184786°N, 24.227804°, 4 m elev., 23.IV.2017 – 8.VI.2017, pitfall traps, 4 spms, leg. Amyntas A. (NHMC) .</p><p>Comments.</p><p>This is the first record of the genus Anaulacus W. S. Macleay, 1825 from Greece, establishing its presence in the southern parts of Europe. Notably, A. ruficornis was found in the south-western part of Crete, while a more eastern locality would be more expected given the species’ distribution. We should highlight that Egypt is excluded from the species’ distribution in Bousquet (2017 a), while it is included in Assmann et al. (2015) who cite an old reference (Alfieri 1976). More research is recommended in order to clarify whether the species also occurs in the Greek mainland and the Aegean, as well as to clarify its distribution in Crete.</p></div>	https://treatment.plazi.org/id/A5320399ACF25BC597497800854D07E8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
6FD716851FAF5DA2961E95C8E85BE89C.text	6FD716851FAF5DA2961E95C8E85BE89C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Anisodactylus (Pseudodichirius) intermedius Dejean 1829	<div><p>16. Anisodactylus (Pseudodichirius) intermedius Dejean, 1829</p><p>Fig. 3 G</p><p>Habitat and general distribution.</p><p>Anisodactylus intermedius is a Turano-Mediterranean species (Arndt et al. 2011). The species is found in wetlands (Austin et al. 2008).</p><p>Material examined.</p><p>• Chania: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.575&amp;materialsCitation.latitude=35.4797" title="Search Plazi for locations around (long 23.575/lat 35.4797)">Falassarna wetland</a>, 35.479700°N, 23.575000°E, 2 m elev., 9.VII.2015 – 2.XI.2015, pitfall traps, 1 spm, leg. Pavlou Chr. (NHMC) .</p><p>Comments.</p><p>Anisodactylus intermedius was reported for the first time in Greece by v. Oertzen (1886) from Corfu and the Greek mainland (Aetoloakarnania, Veluchi). Arndt et al. (2011) also report Peloponnese in the species’ distribution. To our knowledge, this is the first time A. intermedius has been reported from Crete. The species was recorded from Falassarna, in the westernmost coast of the island. Despite the extensive sampling throughout Crete, the species remains known only from the Falassarna area. Hence, we assume that this may be a recent dispersal event from Peloponnese.</p></div>	https://treatment.plazi.org/id/6FD716851FAF5DA2961E95C8E85BE89C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
0F10FDB687CB5EF0A324BE46B8FB7428.text	0F10FDB687CB5EF0A324BE46B8FB7428.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachinus plagiatus Reiche 1868	<div><p>5. Brachinus (s. str.) plagiatus Reiche, 1868</p><p>Figs 2 D, 5 A</p><p>Habitat and general distribution.</p><p>Brachinus plagiatus is a Turano-Mediterranean species (Hrdlička 2017). The species is associated with saline fields and marshes (Nyilas 1994), but also freshwater lakes and ponds (Austin et al. 2008).</p><p>Material examined.</p><p>Heraklion: • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.061403&amp;materialsCitation.latitude=35.338505" title="Search Plazi for locations around (long 25.061403/lat 35.338505)">Almyros estuary</a>, salt-marsh, 35.338506°N, 25.061404°E, 2 m elev., 3.V.2012 – 3.VI.2013, pitfall traps, 1 spm, leg. Aspradaki E. (NHMC) ; • same data, but in phrygana .</p><p>Comments.</p><p>Brachinus plagiatus was recorded for the first time from Greece (Corfu, Attiki, Aetoloakarnania and Zakynthos) by von Oertzen (1886) as B. bombarda . Peloponnese (Tayget, Nauplio) was later on added in the Greek distribution of the species (Apfelbeck 1904). To our knowledge, this is the first record of this species in Crete. The species was found in a saline environment in the marsh formed by the delta of Almyros estuary. The species’ distribution in Crete requires further investigation. Whether this is a recent dispersal or an old relictual presence, is unclear.</p></div>	https://treatment.plazi.org/id/0F10FDB687CB5EF0A324BE46B8FB7428	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
40D67E00751956959BEAAF9DBB54C0B1.text	40D67E00751956959BEAAF9DBB54C0B1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Brachinus psophia Audinet-Serville 1821	<div><p>5. Brachinus (s. str.) psophia Audinet-Serville, 1821</p><p>Figs 2 E, 5 B</p><p>Habitat and general distribution.</p><p>Brachinus psophia has a wide distribution over Europe, North Africa to the Far East (Arndt et al. 2011). It is a hygrophilous, wetland species, that is also common in croplands (Drmić et al. 2016; Lemic et al. 2017).</p><p>Material examined.</p><p>• Chania: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.575&amp;materialsCitation.latitude=35.4797" title="Search Plazi for locations around (long 23.575/lat 35.4797)">Falassarna wetland</a>, 35.479700°N, 23.575000°E, 2 m elev., 15.V.2015 – 9.VII.2015, pitfall traps, 20 spms, leg. Pavlou Chr. (NHMC) ; • same location data, but: 9.VII.2015 – 2.XI.2015, 7 spms</p><p>Comments.</p><p>Brachinus psophia was recorded for the first time from Greece (Corfu, Aetoloakarnania, Peloponnese) by Apfelbeck (1904). Arndt et al. (2011) accept this species for Greece (Ionian islands, Greek mainland and Peloponnese). The extant literature suggests that this is the first record of B. psophia in Crete. The species was only found in Falassarna Bay indicating a dispersal event from a western locality, perhaps Peloponnese. Dispersals from western areas confined in the west coastline of Crete are also recovered for other Carabidae species (e. g. Scarites procerus eurytus Fischer von Waldheim, 1828) and other taxa (e. g. Tentyria grossa Besser, 1832, Tenebrionidae (Trichas 1996)) . The species was found in a coastal salt-marsh.</p></div>	https://treatment.plazi.org/id/40D67E00751956959BEAAF9DBB54C0B1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
650DA829E25559C88E396BF9FECCD55E.text	650DA829E25559C88E396BF9FECCD55E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calathus (Neocalathus) cinctus Motschulsky 1850	<div><p>27. Calathus (Neocalathus) cinctus Motschulsky, 1850</p><p>Figs 4 H, 5 J</p><p>Habitat and general distribution.</p><p>Calathus cinctus has a wide distribution in Europe and Near East (Arndt et al. 2011). The species prefers open habitats, with sparse vegetation (Brigić et al. 2016).</p><p>Material examined.</p><p>• Lasithi: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.227371&amp;materialsCitation.latitude=35.236595" title="Search Plazi for locations around (long 26.227371/lat 35.236595)">Moni Toplou</a>, phrygana, 35.236595°N, 26.227372°E, 170 m elev., 11.XII.1993, handpicking, 1 spm, leg. Trichas A. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.141216&amp;materialsCitation.latitude=34.945328" title="Search Plazi for locations around (long 26.141216/lat 34.945328)">Koufonisi isl.</a>, northern beach, 34.945326°N, 26.141217°E, 2 m elev., 1.XI.1997 – 16.I.1998, pitfall traps, 1 spm, leg. Trichas A. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.236603&amp;materialsCitation.latitude=35.049107" title="Search Plazi for locations around (long 26.236603/lat 35.049107)">Xerokampos salt-lake</a>, 35.049107°N, 26.236602°E, 1 m elev., 1.IV.2015 – 31.V.2015, pitfall traps, 2 spms, leg. Pavlou Chr. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.24008&amp;materialsCitation.latitude=35.05061" title="Search Plazi for locations around (long 26.24008/lat 35.05061)">Xerokampos phrygana</a>, 35.050610°N, 26.240080°E, 2 m elev., 5.VIII.2015 – 26.XI.2015, pitfall traps, 4 spms, leg. Pavlou Chr. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.696203&amp;materialsCitation.latitude=35.049095" title="Search Plazi for locations around (long 25.696203/lat 35.049095)">Bramiana Dam</a>, 35.049097°N, 25.696204°E, 88 m elev., 19.X.2017 – 13.XII.2017, pitfall traps, 1 spm, leg. Amyntas A. (NHMC) ; • same data, but 13.XII.2017 – 15.III.2018, 6 spms; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.697588&amp;materialsCitation.latitude=35.048798" title="Search Plazi for locations around (long 25.697588/lat 35.048798)">Bramiana Dam</a>, in phrygana vegetation, 35.048798°N, 25.697588°E; 72 m elev., 13.XII.2017 – 15.III.2018, pitfall traps, 10 spms, leg. Amyntas A. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.271505&amp;materialsCitation.latitude=35.2044" title="Search Plazi for locations around (long 26.271505/lat 35.2044)">Kouremenos wetland</a>, 35.204399°N, 26.271505°E, 1 m elev., 13.IV.2022 – 10.VI.2022, pitfall traps, 2 spms, leg. Bolanakis G. (NHMC) .</p><p>Comments.</p><p>Calathus cinctus is known in Greece from Greek mainland, Peloponnese and the Aegean (Arndt et al. 2011). To our knowledge, this is the first time C. cinctus has been reported from Crete. The species has been found mainly in coastal saline wetlands (Kouremenos, Xerokampos), but also in inland artificial dams (Bramiana) and phrygana vegetation (Moni Toplou). It appears to be sympatric with the closely related species C. mollis . Calathus cinctus is restricted to east Crete, not crossing the isthmus of Ierapetra, which may indicate an eastern origin.</p></div>	https://treatment.plazi.org/id/650DA829E25559C88E396BF9FECCD55E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
0A4F09E6ADB950668D187AB833B1C63B.text	0A4F09E6ADB950668D187AB833B1C63B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calosoma (Campalita) maderae (Fabricius 1775)	<div><p>3. Calosoma (Campalita) maderae (Fabricius, 1775)</p><p>Fig. 2 B</p><p>General distribution.</p><p>Calosoma maderae has a wide distribution in Europe, North Africa and Near and Middle East (Häckel 2017).</p><p>Material examined.</p><p>• Chania: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.138031&amp;materialsCitation.latitude=35.20152" title="Search Plazi for locations around (long 24.138031/lat 35.20152)">Chora Sfakion</a>, 35.201518°N, 24.138031°E, 40 m elev., 16.V.2019, handpicking, 1 spm, leg. Alexandrakis G. (NHMC) ; • Lefka Ori, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.14765&amp;materialsCitation.latitude=35.29416" title="Search Plazi for locations around (long 24.14765/lat 35.29416)">Niato plateau</a>, 35.29416°N, 24.14765°E, 1226 m elev., pitfall traps, 20.VI.2019 – 31.VII.2019, 1 spm, leg. Bolanakis G. (NHMC) .</p><p>Comments.</p><p>The taxonomic status of this species and its close relatives is ambiguous, with a wide discussion of the species / subspecies rank amongst the different taxa (see Roeschke (1900); Breuning (1927); Jeannel (1941)). Herein, we follow Bruschi (2010) who suggests a lumping approach for the different taxa of the C. maderae group, as well as the inclusion of C. auropunctatum as a synonym of C. maderae s. str. (https://www.calosomas.com/Campalita/cal_maderae.html).</p><p>The first record of C. maderae in Crete is that of Schaum (1857), from material collected by Zebe. Zebe also managed to collect C. maderae from the island of Syros “ Syra ” at the same time (Schaum 1857). Apfelbeck (1904) cites these references and adds two more, both from the volcanic island of Milos. Herein, we report new specimens of C. maderae from Crete for the first time in more than 150 years. This is indicative of its rarity, which is both distributional (until now the only known localities of the species in Crete are confined on Lefka Ori mountain massif) and in terms of abundance (only two specimens), in contrast to the other two Calosoma spp. of Crete which exhibit far greater abundances (and a broad distribution in C. sycophanta case). The species was found in a mountain shrubland (Niato plateau) with Berberis cretica and Quercus coccifera, as well as to an urban environment (Chora Sfakion), near the sea level.</p></div>	https://treatment.plazi.org/id/0A4F09E6ADB950668D187AB833B1C63B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
9508A10561085C8E91B05F1F7F22FDB4.text	9508A10561085C8E91B05F1F7F22FDB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Calosoma inquisitor (Linnaeus 1758)	<div><p>2. Calosoma (s. str.) inquisitor (Linnaeus, 1758)</p><p>Fig. 2 A</p><p>Habitat and general distribution.</p><p>Calosoma inquisitor has a wide distribution in Europe, North Africa, Anatolia, Middle and Far East (Häckel 2017). It is a forest dwelling species, frequently spotted to prey on caterpillars in trees and bushes (Trautner and Geigenmüller 1987; Arndt et al. 2011).</p><p>Material examined.</p><p>• Heraklion: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.9222&amp;materialsCitation.latitude=35.169895" title="Search Plazi for locations around (long 24.9222/lat 35.169895)">Rouvas Forest by the monastery</a>, 35.169894°N, 24.922200°E, 1100 m elev., 6.V.1990 – 27.XI.1990, pitfall traps, 3 spms, leg. Trichas A. (NHMC) ; • Psiloritis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.821604&amp;materialsCitation.latitude=35.196594" title="Search Plazi for locations around (long 24.821604/lat 35.196594)">Nida plateau SW slopes</a>, 35.196594°N, 24.821603°E, 1762 m elev., 13.I.1991, 1 spm, handpicking, leg. Trichas A. (NHMC) ; • Rethymnon: Psiloritis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.7938&amp;materialsCitation.latitude=35.198692" title="Search Plazi for locations around (long 24.7938/lat 35.198692)">Akolita plateau in sub-alpine phrygana</a>, 35.198692°N, 24.793800°E, 1900 m elev., 6.V.1990 – 13.I.1991, pitfall traps, 9 spms, leg. Trichas A. (NHMC) ; • Psiloritis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.792&amp;materialsCitation.latitude=35.197292" title="Search Plazi for locations around (long 24.792/lat 35.197292)">Akolita plateau</a>, 35.197291°N, 24.791999°E, 1900 m elev., 14.IV.2000 – 2.VII.2000, 120 spms, pitfall traps, leg. Chatzaki M. (NHMC) ; • same data, but: 31.X.2000 – 12.VI.2001, 70 spms; same data, but: 35.209192°N, 24.792997°E, 2213 m elev., 16 spms .</p><p>Comments.</p><p>Csiki (1914), summarising all previous classic works on Cretan Coleoptera fauna in his “ Fauna Coleopterorum Insulae Cretae ”, reported only C. sycophanta (Linnaeus, 1758) and C. maderae from Crete. Calosoma inquisitor (Linnaeus, 1758) is reported from Crete for the first time by Breuning (1927), from specimens deposited in the collections of the Natural History Museum of Vienna (“ Mus. Vindob. ”). Later on, Jeannel (1941) listed again C. inquisitor in Crete (“ Crète et îles de la mer Égée ”) without any further details, most probably based on Breuning’s references. Trichas (1996), provided fresh C. inquisitor specimens from Ida Mt., including also the above references as valid, in his doctoral dissertation of South Aegean Archipelago ground-dwelling beetles. More recently, Arndt et al. (2011) also listed C. sycophanta and C. maderae on Crete, but excluded C. inquisitor from the island. Finally, C. inquisitor is included in the appendix of Kaltsas et al. (2013), albeit this is an ecological work, so the species references could easily be overlooked when compiling faunal records. Moreover, as no other (specimen based) records for this taxon in Crete have been presented in the international literature between 1927–2013, this is the first detailed cataloguing of C. inquisitor on the island for almost a century. Clearly, C. inquisitor forms robust populations in the mountain of Psiloritis (Ida Mt.), especially on higher altitudes, found both in forests and mountain shrublands. Additionally, although there are dozens of specimens of C. sycophanta in the NHMC collections, from almost all larger Cretan Mountain massifs (Ida, Dikti, Thrypti – except Lefka Ori Mts. where only C. maderae have been observed twice, see below), C. inquisitor seems to be confined only on the central Cretan massif (Ida).</p></div>	https://treatment.plazi.org/id/9508A10561085C8E91B05F1F7F22FDB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
88D817D464AD5EFFA45006EDCD39B41F.text	88D817D464AD5EFFA45006EDCD39B41F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Carabus (Pachystus) graecus subsp. trojanus Dejean 1826	<div><p>4. Carabus (Pachystus) graecus trojanus Dejean, 1826</p><p>Fig. 2 C</p><p>Habitat and general distribution.</p><p>Carabus graecus trojanus is endemic to Greece (Arndt et al. 2011). The subspecies, as the other members of C. graecus Dejean, 1826, live in a broader range of habitats, found in phrygana, maquis, urban areas and forests.</p><p>Material examined.</p><p>• Heraklion: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.153454&amp;materialsCitation.latitude=35.34339" title="Search Plazi for locations around (long 25.153454/lat 35.34339)">Heraklion Port</a>, 35.343391°N, 25.153453°E, 1 m elev., 9.X.1989, handpicking, 1 spm, leg. Trichas A. (NHMC) .</p><p>Comments.</p><p>Turin et al. (2003) repeatedly mentioned that C. g. trojanus occurs in Crete (although in p. 48, they mark Crete with “? ”). Trichas (1996) mentioned that the original record of C. g. trojanus in Crete by Cecconi (1895) from Chania (as C. trojanus) was most likely erroneous. Nevertheless, Trichas (1996) himself found one specimen of C. g. trojanus in Heraklion port, which he reported, that differs clearly from the Karpathos island subspecies, i. e. C. t. oertzeni Ganglbauer, 1888 – now considered also as a separate subspecies of C. graecus (Häckel, 2017) . Thus, C. g. trojanus has been recorded at least twice from Crete in the span of a century. Given the extremely high abundances of the endemic Carabus (Procrustes) banonii Dejean, 1830 throughout the island (thousands of specimens in the NHMC collections) in almost all habitats; and the systematic and continuous samplings with pitfall traps from the late 80 s till today, we believe that the records of C. g. trojanus in Crete derive from random and not stable dispersal incidents (anthropochorous). The absence of a second Carabus lineage in Crete is an interesting biogeographical problem, since Crete is large enough to host more than one Carabus species, whereas other, much smaller islands host two or more species (e. g. Kythira, Karpathos, Rhodes (Trichas 1996)).</p></div>	https://treatment.plazi.org/id/88D817D464AD5EFFA45006EDCD39B41F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
A112146301905DA3B3AE81A5559204A9.text	A112146301905DA3B3AE81A5559204A9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Chlaenius (Epomis) dejeanii (Dejean 1831)	<div><p>10. Chlaenius (Epomis) dejeanii (Dejean, 1831)</p><p>Fig. 3 A</p><p>Habitat and general distribution.</p><p>Chlaenius dejeanii is an East European-Mediterranean species with its distribution reaching Middle East (e. g. Syria, Iraq) (Arndt et al. 2011; Kirschenhofer 2017).</p><p>Material examined.</p><p>• Heraklion: Aposelemis, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.4208&amp;materialsCitation.latitude=35.234203" title="Search Plazi for locations around (long 25.4208/lat 35.234203)">Sfentyli Dam</a>, 35.234204°N, 25.420799°E, 211 m elev., 24.III.2017 – 25.IV.2017, pitfall traps, 1 spm, leg. Amyntas A. (NHMC) ; • same data but 25.IV.2017 – 30.V.2017, 1 spm; • same data, but 30.V.2017 – 19.X.2017, 21 spms; • Tympaki, salt-lake, 35.075001°Ν, 24.745918°Ε, 2 m elev., 15.IV.2022 – 2.VI.2022, pitfall traps, 4 spms, leg. Bolanakis G. (NHMC) ; • same data, but 2.VI.2022 – 4.VII.2022, 2 spms; • same data, but 15.IV.2022, handpicking, 1 spm; • Lasithi: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.7067&amp;materialsCitation.latitude=35.043095" title="Search Plazi for locations around (long 25.7067/lat 35.043095)">Bramiana artificial lake</a>, 35.043096°N, 25.706700°E, 76 m elev., 4.I.1999 – 3.III.1999, pitfall traps, 1 spm, leg. Papadimitrakis M. (NHMC) ; • same data, but 3.III.1999 – 4.V.1999, 13 spms; • same data, but 4.V.1999 – 22.VII.1999, 13 spms .</p><p>Comments.</p><p>Chlaenius dejeanii is a prominent and scarce (Austin et al. 2008) carabid that was first reported from Greece by Schaum (1857) from Peloponnese (“ Morea ”) and Athens. Apfelbeck (1904) expanded the distribution of the species beyond the Greek mainland in the Ionian islands, by the reporting of specimens from Corfu. This is the first time the species is recorded from Crete. Several populations were captured through pitfall trapping, in the central-east part of the island, from the north (e. g. Aposelemis) to the south (e. g. Tympaki). The species was found in natural (Tympaki) and artificial wetlands (Aposelemis, Bramiana), with different gradients of salinity.</p></div>	https://treatment.plazi.org/id/A112146301905DA3B3AE81A5559204A9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
5585875D8498556EA720047344E8689C.text	5585875D8498556EA720047344E8689C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Clivina ypsilon Dejean & Boisduval 1829	<div><p>6. Clivina (s. str.) ypsilon Dejean &amp; Boisduval, 1829</p><p>Fig. 2 F</p><p>Habitat and general distribution.</p><p>Clivina ypsilon has a wide distribution in Europe, North Africa, Near and Middle East (Balkenohl 2017). The species occurs in saline, surface waters (Trautner and Geigenmüller 1987; Arndt et al. 2011)</p><p>Material examined.</p><p>• Chania: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.274637&amp;materialsCitation.latitude=35.33623" title="Search Plazi for locations around (long 24.274637/lat 35.33623)">Kournas Lake, north bank</a>, 35.336232°N, 24.274637°E, 8.III.2018 – 15.V.2018, pitfall traps, 1 spm, leg. Amyntas A. (NHMC) ; • Heraklion: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.061403&amp;materialsCitation.latitude=35.338505" title="Search Plazi for locations around (long 25.061403/lat 35.338505)">Almyros Estuary</a>, salt-marsh, 35.338506°N, 25.061404°E, 2 m a. s. l, 28.XI.2012 – 30.V.2013, pitfall traps, 2 spms, leg. Aspradaki E. (NHMC) .</p><p>Comments.</p><p>Clivina ypsilon was noted for the first time in Greece by von Oertzen (1886). Apfelbeck (1904) reports again the species providing two localities (Phaleron, in Attica and Thessaly Province). Arndt et al. (2011) accept this species for Greece, but only for the Greek mainland. This constitutes the first documentation of C. ypsilon in Crete. The species was found in two quite different habitats, i. e., in a salt-marsh and in the sandy banks of a freshwater lake (Kournas).</p></div>	https://treatment.plazi.org/id/5585875D8498556EA720047344E8689C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
39FFD737CF51522F8F0BF9824D17FE04.text	39FFD737CF51522F8F0BF9824D17FE04.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cymindis (Arrhostus) andreae Menetries 1832	<div><p>13. Cymindis (Arrhostus) andreae Ménétriés, 1832</p><p>Figs 3 D, 5 E</p><p>Habitat and general distribution.</p><p>Cymindis andreae is an eastern species. The larger part of its range extents to Near and Middle East, while it is also known from North Africa (Egypt) and Eastern Europe (Kabak 2017).</p><p>Material examined.</p><p>• Lasithi: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.697966&amp;materialsCitation.latitude=34.878044" title="Search Plazi for locations around (long 25.697966/lat 34.878044)">Chrysi isl., north beach</a> salt-lake, 34.878045°N, 25.697966°E, 1 m elev., 17.V.1994, handpicking, 3 spms, leg. Trichas A. (NHMC) ; • Chrysi isl., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.728071&amp;materialsCitation.latitude=34.874798" title="Search Plazi for locations around (long 25.728071/lat 34.874798)">Vages Beach</a>, salt-lake, 34.874799°N, 25.728071°E, 1 m elev., 23.III.2023 – 13.VII.2023, pitfall traps, 6 spms, leg. Bolanakis G. (NHMC) .</p><p>Comments.</p><p>In Greece, Cymindis andreae is only known from the Aegean (Arndt et al. 2011). In fact, the only record of this species in Greece (Rhodes) dates back to 1935 (Schatzmayr 1935). This is the first record of C. andreae in Crete. The species was found in Chrysi islet in 1994 (northern salt-lake) (leg. Trichas) and recorded again in 2023 (leg. Bolanakis), in the southern salt-lake of Chrysi (Vages Beach). Since the southern / eastern coastal salt-lakes of Crete have been intensively sampled, the species presence in the island of Crete is unlikely, although more research is required. The species’ origin is obscure. Crete is very poor in African taxa; thus, it is more likely that the species is an eastern dispersal, a common pattern for species restricted in the eastern parts of the island (or its eastern satellite islets) (Koch 1948; Trichas 1996). We confirm the presence of the species in Greece and encourage further research for this species in the eastern part of the Aegean.</p></div>	https://treatment.plazi.org/id/39FFD737CF51522F8F0BF9824D17FE04	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
CC8932358BD7553980686831E877BC75.text	CC8932358BD7553980686831E877BC75.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cymindis ornata Fischer von Waldheim 1823	<div><p>14. Cymindis (s. str.) ornata Fischer von Waldheim, 1823</p><p>Fig. 3 E</p><p>Habitat and general distribution.</p><p>Cymindis ornata is a West Ponitc species (Kabak 2017). The species’ habitat preferences vary. We have found it in river mouths and coastal phrygana, while Teofilova et al. (2015) note that the species occurs strictly in arid, open environments.</p><p>Material examined.</p><p>• Heraklion: Aposelemis River mouth, 35.334994°N, 25.330875°, 1 m elev., 30.VII.2015 – 27.XI.2015, pitfall traps, 1 spm, leg. Pavlou Chr. (NHMC) ; • Rethymnon: Moni Preveli River, phrygana at the west slopes above the beach, 35.151797°N, 24.472496°, 22 m elev., 25.VI.1996 – 28.VIII.1996, pitfall traps, 1 spm, leg. Lymberakis P. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.358007&amp;materialsCitation.latitude=35.350986" title="Search Plazi for locations around (long 24.358007/lat 35.350986)">Petres River mouth</a>, 35.350988°N, 24.358007°E, 1 m elev., 15.V.2015 – 9.VII.2015, pitfall trap, 2 spms, leg. Pavlou Chr. (NHMC) .</p><p>Comments.</p><p>Cymindis ornata is reported in Greece from the Greek mainland (excluding Peloponnese) and Aegean (Schatzmayr 1935; Arndt et al. 2011). To our knowledge, this is the first record of this species in Crete. It is common for Balkan species or genera to occur in Crete, although such species are usually restricted in the west part of the island (Trichas 1996). In this case, all the records are in the central Crete (both south and north).</p></div>	https://treatment.plazi.org/id/CC8932358BD7553980686831E877BC75	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
C9257AC7C6CB549F856030701FF0ED43.text	C9257AC7C6CB549F856030701FF0ED43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicheirotrichus (Pelagophilus) punicus Bedel 1899	<div><p>21. Dicheirotrichus (Pelagophilus) punicus Bedel, 1899</p><p>Fig. 4 B</p><p>Habitat and general distribution.</p><p>Dicheirotrichus punicus has an circum-Mediterranean distribution (Jaeger and Kataev 2017). The species occurs in saline coastal habitats (Austin et al. 2008; Arndt et al. 2011).</p><p>Material examined.</p><p>• Lasithi: Chrysi isl., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.728071&amp;materialsCitation.latitude=34.874798" title="Search Plazi for locations around (long 25.728071/lat 34.874798)">Vages Beach</a>, salt-lake, 34.874799°N, 25.728071°E, 1 m elev., 23.III.2023 – 13.VII.2023, pitfall traps, 2 spms, leg. Bolanakis G. (NHMC) .</p><p>Comments.</p><p>Dicheirotrichus punicus was first recorded in Greece by Apfelbeck (1904) from Kyklades (Naxos). To our knowledge, this is the first record of the species in Crete. Moreover, it is the first time the species has been recorded in Greece in more than one century (Arndt et al. 2011). Its geographical distribution in the Aegean Archipelago requires further research. The species was found in a salt-lake at Chrysi islet. In contrast to D. obsoletus, the species has not been recorded in the Cretan island yet, although it may occur in southern / eastern saline wetlands.</p></div>	https://treatment.plazi.org/id/C9257AC7C6CB549F856030701FF0ED43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
5FA015CE59AC55F8AB78D3F845683418.text	5FA015CE59AC55F8AB78D3F845683418.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dicheirotrichus obsoletus (Dejean 1829)	<div><p>20. Dicheirotrichus (s. str.) obsoletus (Dejean, 1829)</p><p>Figs 4 A, 5 G</p><p>Habitat and general distribution.</p><p>D. obsoletus has a Mediterranean distribution (Jaeger and Kataev 2017). The species occurs in saline coastal habitats (Austin et al. 2008; Arndt et al. 2011).</p><p>Material examined.</p><p>• Heraklion: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.061201&amp;materialsCitation.latitude=35.339104" title="Search Plazi for locations around (long 25.061201/lat 35.339104)">Almyros River</a>, salt-marsh, 35.339104°N, 25.061201°E, 1 m elev., 3.V.2012 – 3.VI.2012, pitfall traps 2 spms, leg. Aspradaki E. (NHMC) ; • same data, but 1.VIII.2012 – 28.XI.2012, 3 spms; • same data, but 28.XI.2012 – 30.V.2013, 1 spm; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.33089&amp;materialsCitation.latitude=35.335" title="Search Plazi for locations around (long 25.33089/lat 35.335)">Aposelemis River mouth</a>, 35.335000°N, 25.330891°E, 1 m elev., 6.XI.2014 – 5.12.2014, pitfall traps, 2 spms, leg. Pavlou Chr. (NHMC) ; • same data, but 30.VII.2015 – 27.XI.2015, 4 ind; • Tympaki, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.746101&amp;materialsCitation.latitude=35.0747" title="Search Plazi for locations around (long 24.746101/lat 35.0747)">Kokkinos Pyrgos</a>, salt-marsh, 35.074700°N, 24.746101°E, 1 m elev., 9.XII.2021, handpicking, 1 spm, leg. Bolanakis G. (NHMC) .</p><p>Comments.</p><p>Dicheirotrichus obsoletus is reported from the Greek mainland (including Peloponnese) and the Aegean (Arndt et al. 2011). This is the first time D. obsoletus is reported from Crete. It is important to note that, in addition to D. punicus (see below), this also constitutes the first documentation of the genus Dicheirotrichus on the island of Crete. Dicheirotrichus obsoletus has been collected from central Crete, both in north and south coastal saline wetlands and marshes.</p></div>	https://treatment.plazi.org/id/5FA015CE59AC55F8AB78D3F845683418	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
DD4D33AE1564521599777020E53E9240.text	DD4D33AE1564521599777020E53E9240.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Grammognatha euphratica Latreille & Dejean 1822	<div><p>1. Grammognatha euphratica Latreille &amp; Dejean, 1822</p><p>Fig. 1</p><p>Habitat and general distribution.</p><p>Grammognatha euphratica has a wide distribution extending from Morocco and southern Spain over northern Africa, parts of the eastern Mediterranean, Arabia and Central Asia as far as Pakistan and northern India (Cassola 1981; Franzen 2001). The species is found in salt marsh and salt meadow habitats, mainly active at hours of twilight (Trautner and Geigenmüller 1987; Arndt et al. 2011) or night (Cassola et al. 2014). Within the eastern Mediterranean, more or less recent distribution data exist from Cyprus, Turkey, Israel and Egypt (Franzen 2001; Şekeroğlu and Aydin 2002; Austin et al. 2008), while recently, the beetle has been discovered on the island of Sant’Antioco in Italy (Cassola et al. 2014). In Greece, it is also present on the island of Rhodes ((Horn 1910) without exact locality); however, this occurrence has not been reconfirmed by the latest known surveys of Wiesner (1990, 1994). The only known locality on Crete until the present study was that of Palaiokastron, in east Crete (Cassola 1981).</p><p>Material examined.</p><p>• <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.35863&amp;materialsCitation.latitude=35.280643" title="Search Plazi for locations around (long 25.35863/lat 35.280643)">Heraklion</a>: Aposelemis River, 35.280642°N, 25.358629°E, 131 m elev., 1.VI.1989, handpicking, 1 spm, leg. Trichas A. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.3309&amp;materialsCitation.latitude=35.335003" title="Search Plazi for locations around (long 25.3309/lat 35.335003)">Aposelemis River mouth</a> 35.335003°N, 25.330901°E, 28.V.2015 – 30.VII.2015, pitfall traps, 9 spms, leg. Pavlou Chr. (NHMC) ; • same data, but 30.VII.2015 – 27.XI.2015, 2 spms; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.27149&amp;materialsCitation.latitude=35.204384" title="Search Plazi for locations around (long 26.27149/lat 35.204384)">Kouremenos Beach wetland</a>, 35.204385°N, 26.271490°E, 13.IV.2022 – 10.VI.2022, pitfall traps, 4 spms, leg. Bolanakis G. (NHMC) ; • same data, but 13.IV.2022, handpicking, 1 spm, leg. Pavlou Chr. (NHMC); • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.728071&amp;materialsCitation.latitude=34.874798" title="Search Plazi for locations around (long 25.728071/lat 34.874798)">Chrysi isl.</a>, Vages Beach, salt-lake, 34.874799°N, 25.728071°E, 23.III.2023 – 13.VII.2023, pitfall traps, 2 spms, leg. Bolanakis G. (NHMC) .</p><p>Comments.</p><p>Cassola (1981) was the first author to report Grammognatha euphratica from Crete (“ Palaiokastron ”, east Crete or Palaikastro / Palekastro – not to be confused with the tautonym in central Crete). Both this record and the ones from Rhodes (Cassola 1973) needed confirmation according to Franzen (2001) and Arndt et al. (2011). For example, the record from Rhodes was never recovered during the surveys of Wiesner in 1990 and 1994 (Franzen 2001). In essence, the records presented here are the first of this rare and fragmentarily distributed beetle from Crete in four decades. We confirm the species’ presence in Crete and also expand its distribution in a southern satellite islet of Crete, Chrysi. All our records are from salt-lakes and are confined in the central-east part of the island. Moreover, the Aposelemis specimens represent the westernmost findings of this species on Crete, pointing to an eastern origin of the species’ dispersal.</p></div>	https://treatment.plazi.org/id/DD4D33AE1564521599777020E53E9240	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
E4193D02709F5C05A384493D52FC0C17.text	E4193D02709F5C05A384493D52FC0C17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laemostenus venustus (Dejean 1828)	<div><p>28. Laemostenus (s. str.) venustus (Dejean, 1828)</p><p>Fig. 4 I</p><p>Habitat and general distribution.</p><p>Laemostenus venustus is a Euro-Mediterranean species that dwells in forests (especially of Q. coccifera and P. orientalis) (Austin et al. 2008).</p><p>Material examined.</p><p>• Rethymno: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.9222&amp;materialsCitation.latitude=35.169895" title="Search Plazi for locations around (long 24.9222/lat 35.169895)">Rouvas Forest</a>, 35.169897°N, 24.922200°E, 1100 m elev., 20.II.1990 – 6.V.1990, pitfall traps, 1 spm, leg. Trichas A. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.7778&amp;materialsCitation.latitude=35.1764" title="Search Plazi for locations around (long 24.7778/lat 35.1764)">Lochria</a>, in Quercus coccifera dehesa-like forest, 35.17640°N 24.77780°E, 980 m elev., 24.V.2019 – 2.VII.2019, pitfall traps, 1 spm leg. G. Bolanakis (NHMC) .</p><p>Comments.</p><p>Laemostenus venustus is known in Greece from the Greek mainland and Peloponnese (von Oertzen 1886; Apfelbeck 1904; Arndt et al. 2011). This is the first record of L. venustus reported from Crete. The species has been recorded only from two localities. Specifically, it has been recorded in Rouvas Forest (a shadowed riparian forest with large, veteran plane trees and kermes oaks). The original record dates back to 1990 (Trichas 1996). After several re-samples with pitfall traps, we managed to find one more specimen in 2018, in Lochria, a dehesa-like shrubland with old kermes oaks. Therefore, L. venustus appears to be a rare species in Crete, with a restricted distribution around Ida mountain.</p></div>	https://treatment.plazi.org/id/E4193D02709F5C05A384493D52FC0C17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
DD9894CC82635F19B7EC028577EA59AE.text	DD9894CC82635F19B7EC028577EA59AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Masoreus aegyptiacus Dejean 1828	<div><p>12. Masoreus (s. str.) aegyptiacus Dejean, 1828</p><p>Fig. 3 C</p><p>Habitat and general distribution.</p><p>Masoreus aegyptiacus is an East-Mediterranean species (Bousquet 2017 a). The species occurs in sand-dune habitats (Assmann et al. 2015).</p><p>Material examined.</p><p>• Lasithi: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.697966&amp;materialsCitation.latitude=34.878044" title="Search Plazi for locations around (long 25.697966/lat 34.878044)">Chrysi isl.</a>, north beach salt-lake, 34.878045°N, 25.697966°E, 1 m elev., 15.VI.1992, handpicking, 2 spms, leg. Lymberakis P. (NHMC) ; • Chrysi isl., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.728071&amp;materialsCitation.latitude=34.874798" title="Search Plazi for locations around (long 25.728071/lat 34.874798)">Vages Beach</a>, salt-lake, 34.874799°N, 25.728071°E, 1 m elev., 23.III.2023 – 13.VII.2023, pitfall traps, 39 spms, leg. Bolanakis G. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.14031&amp;materialsCitation.latitude=34.935085" title="Search Plazi for locations around (long 26.14031/lat 34.935085)">Koufonisi isl.</a>, near salt-marsh, 34.935084°N, 26.140310°E, 60 m elev., 16.XI.1997, handpicking, 2 spms, leg. Trichas, A. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.239803&amp;materialsCitation.latitude=35.049892" title="Search Plazi for locations around (long 26.239803/lat 35.049892)">Xerokampos sand dunes</a>, 35.049891°N, 26.239803°E, 1 m elev., 1.IV.2015 – 31.V.2015, pitfall traps, 4 spms, leg. Kyriakouli Chr. (NHMC) ; • same data, but 31. V. 205– 5.VIII.2015 .</p><p>Comments.</p><p>The genus Masoreus Dejean, 1821 has been known from Greece since the 19 th century (von Oertzen 1886) by the species M. wetterhallii (Gyllenhal, 1813) . Masoreus aegyptiacus has only recently been reported from Greece (Wrase 2009) by some specimens from Peloponnese. Arndt et al. (2011) include only Peloponnese in the species’ distribution for Greece; thus, this is the first time the species is recorded from Crete. Till now, the species has been spotted in three localities, one in the eastern-most part of the island (Xerokampos) and two in a couple of south-eastern satellite islands of Crete (Chrysi and Koufonisi). This may indicate an eastern origin, which is to be expected given the species east-Mediterranean distribution (Bousquet 2017 a). Assmann et al. (2015) report that this species prefers sand-dune habitats. In Crete, we observed a clear preference for coastal salt lakes / marshes, like those of Xerokampos and Chrysi.</p></div>	https://treatment.plazi.org/id/DD9894CC82635F19B7EC028577EA59AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
4E0EF02C67B85A9EB599C32773E7E406.text	4E0EF02C67B85A9EB599C32773E7E406.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ophonus (Metophonus) puncticeps Stephens 1828	<div><p>17. Ophonus (Metophonus) puncticeps Stephens, 1828</p><p>Figs 3 H, 5 F</p><p>Habitat and general distribution.</p><p>Ophonus puncticeps has a Turano-Mediterranean distribution, extending into Europe, Near and Middle East, as well as North Africa (Morocco) (Kataev and Wrase 2017). The species has also been introduced in North America (Kataev and Wrase 2017). Ophonus puncticeps is an euryecous species, occurring in open fields and in anthropogenic areas (Lindroth 1968; Nováková and Šťastná 2014).</p><p>Material examined.</p><p>• Heraklion: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.0488&amp;materialsCitation.latitude=35.334" title="Search Plazi for locations around (long 25.0488/lat 35.334)">Almyros River near phrygana</a>, 35.334000°N, 25.048799°E, 3 m elev., 3.VI.2012 – 1.VIII.2012, pitfall traps, 1 spm, leg. Aspradaki E. (NHMC) ; • <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.048098&amp;materialsCitation.latitude=35.333508" title="Search Plazi for locations around (long 25.048098/lat 35.333508)">Almyros River</a>, in phrygana and cultivations, 35.333509°N, 25.048097°E, 2 m elev., 1.VIII.2012 – 28.XI.2012, pitfall traps, 2 spms, leg. Aspradaki E. (NHMC) ; • Aposelemis River, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.421797&amp;materialsCitation.latitude=35.237072" title="Search Plazi for locations around (long 25.421797/lat 35.237072)">Sfentyli Dam</a>, in a Tamarix grove near road, 35.237073°N, 25.421797°E, 218 m elev., 30.V.2017 – 19.X.2017, pitfall traps, 3 spms, leg. Amyntas A. (NHMC) .</p><p>Comments.</p><p>Ophonus puncticeps is reported in Greece from the Greek mainland (excluding Peloponnese), Ionian islands and the Aegean (Arndt et al. 2011). To our knowledge, this is the first time O. puncticeps has been reported from Crete. Ophonus puncticeps is a good disperser (e. g. it has been introduced and expanding in the USA (Larsen 2013)), thus it is not surprising that it also occurs in Crete, given its current distribution in Greece. Both localities, where the species was found, are located in the north part of central Crete, in rivers that form small deltas and have artificial dams.</p></div>	https://treatment.plazi.org/id/4E0EF02C67B85A9EB599C32773E7E406	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
2A1779048D8151F3B09557C90A5F033B.text	2A1779048D8151F3B09557C90A5F033B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paranchus albipes (Fabricius 1796)	<div><p>24. Paranchus albipes (Fabricius, 1796)</p><p>Fig. 4 E</p><p>Habitat and general distribution.</p><p>Paranchus albipes is a Euro-Mediterranean species, also occurring in Turkey (Schmidt 2017). It is a montane species (Arndt et al. 2011) that dwells in different types of shadow banks (Serrano 1988).</p><p>Material examined.</p><p>• Heraklion: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.165031&amp;materialsCitation.latitude=35.27955" title="Search Plazi for locations around (long 25.165031/lat 35.27955)">Agia Irini</a> temporary stream with Platanus orientalis veteran trees, 35.279550°N, 25.165032°E, 140 m elev., 26.V.2016 – 8.V.2016, pitfall traps, 13 spms, leg. Vlachopoulos N. (NHMC) .</p><p>Comments.</p><p>To our knowledge, this is the first detailed record of P. albipes in Crete and Greece. Schmidt (2017) does not include Greece in the species’ distribution, while Arndt et al. (2011) include it in their key for the Greek ground beetles, but omit it from the catalogue provided. The species was found in a small, thick riparian forest. This is also the first record of the genus Paranchus in Greece and Crete.</p></div>	https://treatment.plazi.org/id/2A1779048D8151F3B09557C90A5F033B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
EC0313283F24595C802468354FAD21BD.text	EC0313283F24595C802468354FAD21BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parophonus (Ophonomimus) hirsutulus (Dejean 1829)	<div><p>18. Parophonus (Ophonomimus) hirsutulus (Dejean, 1829)</p><p>Figs 3 I, 5 H</p><p>Habitat and general distribution.</p><p>Parophonus hirsutulus has a Turano-Mediterranean distribution (Kataev and Wrase 2017).</p><p>Material examined.</p><p>• Rethymno: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.358&amp;materialsCitation.latitude=35.35103" title="Search Plazi for locations around (long 24.358/lat 35.35103)">Petres River mouth</a>, 35.351030°N, 24.358000°E, 2 m elev., 15.V.2015 – 9.VII.2015, pitfall traps, 9 spms, leg. Pavlou Chr. (NHMC) ; • same data, but 9.VII.2015 – 2.XI.2015, 30 spms; • Amari, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.581507&amp;materialsCitation.latitude=35.27571" title="Search Plazi for locations around (long 24.581507/lat 35.27571)">Potamon Dam</a>, 35.275710°N, 24.581507°E, 206 m elev., 4.IV.2017, handpicking, 1 spm, leg. Amyntas A. (NHMC) ; • same data, but 8.VI.2017, 2 spms, leg. Pavlou Chr. (NHMC); • same data, but 8.VI.2017 – 17.X.2017, pitfall traps, 1 spm, leg. Amyntas A. (NHMC) .</p><p>Comments.</p><p>Parophonus hirsutulus has a Turano-Mediterranean distribution (Kataev and Wrase 2017). In Greece, the species has been reported from the Greek mainland and Peloponnese (Apfelbeck 1904; Arndt et al. 2011). This is the first record of P. hirsutulus from Crete. The species appears to be restricted in central Crete.</p></div>	https://treatment.plazi.org/id/EC0313283F24595C802468354FAD21BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
F5D7F4DB9A91543EB30072A30DDF87EF.text	F5D7F4DB9A91543EB30072A30DDF87EF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Polistichus connexus (Geoffroy 1785)	<div><p>29. Polistichus connexus (Geoffroy, 1785)</p><p>Fig. 4 J</p><p>Habitat and general distribution.</p><p>Polistichus connexus has a wide distribution in Europe, North Africa, Near and Middle East (Huber and Marggi 2017). The species occurs in open habitats, often in artificial areas (Holec et al. 2019).</p><p>Material examined.</p><p>• Heraklion: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.061403&amp;materialsCitation.latitude=35.338505" title="Search Plazi for locations around (long 25.061403/lat 35.338505)">Almyros Estuary</a>, salt-marsh, 35.338503°N, 25.061403°E, 1 m elev., 3.V.2012 – 3.VI.2012, pitfall traps, 4 spms, leg. Aspradaki E (NHMC) ; • same data, but 3.VI.2012 – 1.VIII.2012, 1 spm; • Lasithi: Lasithi plateau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.4623&amp;materialsCitation.latitude=35.182903" title="Search Plazi for locations around (long 25.4623/lat 35.182903)">moni Vidiani</a>, 35.182902°N, 25.462299°E, 816 m elev., 13.XII.1997, handpicking, 1 spm, leg. Mylonas M. (NHMC) .</p><p>Comments.</p><p>Polistichus connexus is known from the Greek mainland and Peloponnese (Arndt et al. 2011) and this is the first time the species has been reported from Crete. It has been found in two different habitats (a coastal salt-marsh and a humid mountain plateau with shrublands, dehesa-like forests and cultivations).</p></div>	https://treatment.plazi.org/id/F5D7F4DB9A91543EB30072A30DDF87EF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
24BC178F9D395441A6185A0FB2E3045B.text	24BC178F9D395441A6185A0FB2E3045B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pterostichus (Argutor) cursor (Dejean 1828)	<div><p>25. Pterostichus (Argutor) cursor (Dejean, 1828)</p><p>Fig. 4 F</p><p>Habitat and general distribution.</p><p>Pterostichus cursor is distributed in central-southern Europe, Siberia, Near and Middle East (Arndt et al. 2011). It occurs in freshwater reed-beds (Austin et al. 2008).</p><p>Material examined.</p><p>• Chania: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.25169&amp;materialsCitation.latitude=35.360703" title="Search Plazi for locations around (long 24.25169/lat 35.360703)">Agya Lake</a>, 35.360704°N, 24.251689°E, 5 m elev., pitfall traps, 5 spms, leg. Amyntas A. (NHMC) ; • Rethymno: Amari, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.581507&amp;materialsCitation.latitude=35.27571" title="Search Plazi for locations around (long 24.581507/lat 35.27571)">Potamon Dam</a>, 35.275710°N, 24.581507°E, 206 m elev., 4.IV.2017 – 8.v.2017, pitfall traps, 3 spms., leg. Amyntas A. (NHMC) .</p><p>Comments.</p><p>Pterostichus cursor is reported in Greece from the Greek mainland and Peloponnese (Arndt et al. 2011). This is the first record of P. cursor in Crete. The species was spotted in artificial wetlands, in western and central-west Crete. In the sampling sites, there were plenty of reeds, fitting the species’ habitat preferences.</p></div>	https://treatment.plazi.org/id/24BC178F9D395441A6185A0FB2E3045B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
93C08E87B6E1574DBE1DAF8C53CA28BC.text	93C08E87B6E1574DBE1DAF8C53CA28BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pterostichus (Pseudomaseus) anthracinus (Illiger 1798)	<div><p>26. Pterostichus (Pseudomaseus) anthracinus (Illiger, 1798)</p><p>Figs 4 G, 5 I</p><p>Habitat and general distribution.</p><p>Pterostichus anthracinus is a West Palaearctic species (Arndt et al. 2011). It prefers habitats that combine warm and wet conditions (Kolesnikov and Karyaman 2019) like floodplains and forest stream banks (Lindroth 1992).</p><p>Material examined.</p><p>• <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.25169&amp;materialsCitation.latitude=35.360703" title="Search Plazi for locations around (long 24.25169/lat 35.360703)">Chania Agya Lake</a>, 35.360704°N, 24.251689°E, 5 m elev., 10.IV.2017 – 8.V.2017, pitfall traps, 13 spms, leg. Amyntas A. (NHMC) ; • same data, but 8.V.2017, handpicking, 8 spms, leg. Pavlou Chr. (NHMC); • same data, but 8.V.2017 – 8.VI.2017, pitfall traps, 7 spms, leg. Amyntas A.</p><p>Comments.</p><p>Pterostichus anthracinus is known from the Greek mainland, Peloponnese and the Aegean (von Oertzen 1886; Apfelbeck 1904; Arndt et al. 2011). To our knowledge, this is the first time P. anthracinus has been reported from Crete. It was only recorded from one locality (Agya Lake). Unfortunately, we observed intense human intervention (construction of various artificial structures mainly for recreational activities) near the sampling site. Given that Agya Lake is an artificial lake, it is uncertain whether P. anthracinus is a relatively novel species of the Cretan Carabid fauna. More research regarding its distribution in the island is needed.</p></div>	https://treatment.plazi.org/id/93C08E87B6E1574DBE1DAF8C53CA28BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
D1ABA93062525EC2BFED123C8796DBA3.text	D1ABA93062525EC2BFED123C8796DBA3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scarites (Scallophorites) buparius (Forster 1771)	<div><p>8. Scarites (Scallophorites) buparius (Forster, 1771)</p><p>Fig. 2 I</p><p>Habitat and general distribution.</p><p>Scarites buparius has a Mediterranean distribution, occurring in south Europe and North Africa (Balkenohl 2017). The species is found in coastal sand habitats (Arndt et a. 2017).</p><p>Materials examined.</p><p>• Chania: Platanias, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=24.274637&amp;materialsCitation.latitude=35.33623" title="Search Plazi for locations around (long 24.274637/lat 35.33623)">Platanias River mouth</a> 35.336232°N, 24.274637°E, 1 m elev., 28.V.2021 – 4.VIII.2021, pitfall traps, 1 spm, leg Bolanakis G. (NHMC) .</p><p>Comments.</p><p>Scarites buparius was first mentioned from Crete in 1857 as S. pyracmon (Schaum 1857) from specimens collected by Zebe during his expedition to the island in 1853. This record seems to be the source for all the following references (von Oertzen 1886; Apfelbeck 1904; Csiki 1914; Arndt et al. 2011). In all these references, there is no locality given for the species in Crete. Schaum (1857) described the habitat where Zebe found the specimens, but no precise locality name was given. In this study, we report the first detailed locality for the species in Crete, a coastal wetland that fits perfectly with the habitat described by Schaum (1857) and Arndt et al. (2011). Moreover, we provide concrete data for the species’ existence in Crete for the first time after one and a half centuries. Despite S. buparius being a typical Mediterranean species and the exhaustive samplings that we carried out in the coastal wetlands of Crete, we only managed to obtain one specimen, indicative of the species rarity in the island. Further research is required in order to clarify the species’ distribution in Crete. Unfortunately, when we revisited the small delta where the species was found, the sandy coast had been pressed and cleared to be exploited for recreational activities. Wood and plant debris, as well as stones have been dumped in the small dune where the species was found and we did not manage to find any new specimens. Therefore, further sampling effort is required in order to reconfirm the presence of the S. buparius in Platanias River mouth.</p></div>	https://treatment.plazi.org/id/D1ABA93062525EC2BFED123C8796DBA3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
D8E83208E07B5F4483850CB8D3027970.text	D8E83208E07B5F4483850CB8D3027970.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scarites procerus subsp. eurytus Fischer von Waldheim 1828	<div><p>7. Scarites (s. str.) procerus eurytus Fischer von Waldheim, 1828</p><p>Fig. 2 H</p><p>Habitat and general distribution.</p><p>Scarites p. eurytus has a wide Turano-Mediterranean distribution (Balkenohl 2017). The species occurs in swamp habitats (Arndt et al. 2011).</p><p>Material examined.</p><p>• Chania: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=23.575&amp;materialsCitation.latitude=35.4797" title="Search Plazi for locations around (long 23.575/lat 35.4797)">Falassarna wetland</a>, 35.479700°N, 23.575000°E, 2 m elev., 15.V.2015 – 9.VII.2015, pitfall traps, 18 spms, leg. Pavlou Chr. (NHMC) .</p><p>Comments.</p><p>The species is known from the Greek Mainland and the Aegean (Arndt et al. 2011). More specifically the species was reported for the first time in Greece by von Oertzen (1886) from Andros and Naxos, while the records from the Greek mainland are more recent (Arndt et al. 2011). To our knowledge, this is the first record from Crete. A larger population was spotted in Falassarna Bay, west of Chania. The habitat consists of a small coastal salt-marsh inside a sand-dune system and fits the description given by Arndt et al. (2011) (swampy habitats) for the species’ suitable habitat. It is one of the few areas in Crete combining a marsh habitat with a sand dune system. This could explain the absence of this species in other parts of the island (albeit the extensive research in marshes and coastal deltas). Another explanation could be that the observed population is a result of a dispersal from a western region, something that has been also observed for other beetles found in the west coastline of Crete (Trichas 1996). We consider this scenario less plausible since S. p. eurytus has a wide distribution in all the areas surrounding Crete, although, a stochastic dispersal event from the west cannot be excluded. The species is not yet recorded from Peloponnese, but our record encourages more research regarding the species’ distribution in the southern parts of Greek Mainland. The aggressive touristic development in the wider area of Falassarna Bay and especially around the coastline could pose a serious threat for the species’ population in Crete, even leading to extirpation events.</p></div>	https://treatment.plazi.org/id/D8E83208E07B5F4483850CB8D3027970	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
5884022134FF593386D2CCBF3E183576.text	5884022134FF593386D2CCBF3E183576.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sirdenus (Syrdenopsis) grayii (Wollaston 1862)	<div><p>9. Sirdenus (Syrdenopsis) grayii (Wollaston, 1862)</p><p>Figs 2 G, 5 C</p><p>Habitat and general distribution.</p><p>S. grayii is an Afrotropical-Mediterranean species, mainly found in Cyprus, North Africa and Near East, but also southern Europe (Austin et al. 2008; Bousquet 2017 b). The species dwells in saline wetlands (Austin et al. 2008), burrowing in endogean galleries (Saulleda 1985; Machado 1992).</p><p>Material examined.</p><p>• Lasithi: <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=26.240301&amp;materialsCitation.latitude=35.05098" title="Search Plazi for locations around (long 26.240301/lat 35.05098)">Xerokampos salt-lake (phrygana)</a> 35.050979°N, 26.240301°E, 2 m elev., 5.VIII.2015 – 26.XI.2015, pitfall traps, 1 spm, leg. Pavlou Chr. (NHMC) ; • Chrysi isl., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.728071&amp;materialsCitation.latitude=34.874798" title="Search Plazi for locations around (long 25.728071/lat 34.874798)">Vages Beach salt-lake</a>, 34.874799°N, 25.728071°E, 1 m elev., 23.III.2023 – 13.VII.2023, pitfall traps, 62 spms, leg. Bolanakis G. (NHMC) .</p><p>Comments.</p><p>The genus Sirdenus was mentioned for a first time in Greece in Arndt et al. (2011) by a specimen of S. filliformis from “ Griechenland, Attica ”, deposited in the collection of Museum für Naturkunde, Berlin. Arndt et al. (2011) were critical of the validity of this specimen. Thus, our record is the first concrete record of S. grayii in Crete and consequently Greece, confirming the presence of the genus in Greece. Naturally, the species was first discovered in a salt-lake (the largest of Crete) at the easternmost coastline of the island (Xerokampos), by a single specimen in 2015. However, we re-sampled this locality many times and never managed to re-collect this species. According to Matalin and Makarov (2008), S. grayii is a rare species, difficult to encounter. In 2023, we found a large population of S. grayii (62 specimens) in Chrysi, a southern satellite islet of Crete. The species was found in a small, temporary salt-lake, near a southern beach of the islet. The species has not been spotted in other salt lakes and marshes in the west, central and north parts of Crete (e. g. Falassarna, Almyros, Tympaki). The current known distribution of the species in Crete indicates an eastern origin, echoing the biogeographical remarks of Koch (1948) for a dispersal path as such.</p></div>	https://treatment.plazi.org/id/5884022134FF593386D2CCBF3E183576	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
F631A687C16D51B5967A650695A55B37.text	F631A687C16D51B5967A650695A55B37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichis maculata Klug 1832	<div><p>15. Trichis maculata Klug 1832</p><p>Fig. 3 F</p><p>Habitat and general distribution.</p><p>Trichis maculata is a Turano-Mediterranean species (Kabak 2017). The species occurs in coastal saline habitats (Arndt et al. 2011).</p><p>Material examined.</p><p>• Chrysi isl., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=25.728071&amp;materialsCitation.latitude=34.874798" title="Search Plazi for locations around (long 25.728071/lat 34.874798)">Vages Beach</a>, salt-lake, 34.874799°N, 25.728071°E, 1 m elev., 23.III.2023 – 13.VII.2023, pitfall traps, 6 spms, leg. Bolanakis G. (NHMC) .</p><p>Comments.</p><p>Trichis maculata has been reported from Greece by von Oertzen (1886) from Attiki. In his 1904 report, Apfelbeck also cites T. maculata from Attiki, attributing this information to Krüper. In addition to the record of Apfelbeck (1904), no further documentation pertaining to this species occurs for Greece. This is the first documentation of the genus in Greece for a period of 150 years and, moreover, this signifies the inaugural record of the species in Crete. Trichis maculata was found in a small salt-lake in the southern coast of Chrysi islet. The species occurs in North Africa and near East (Kabak 2017) and has not been found yet anywhere else in Crete. Further research is required in order to confirm whether the species is also present in other parts of Crete or Greece.</p></div>	https://treatment.plazi.org/id/F631A687C16D51B5967A650695A55B37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pavlou, Christoforos;Bolanakis, Giannis;Kardaki, Ljubitsa;Trichas, Apostolos	Pavlou, Christoforos, Bolanakis, Giannis, Kardaki, Ljubitsa, Trichas, Apostolos (2025): Forty years of ground-beetle sampling in Crete. A major contribution to the Carabidae (Coleoptera, Adephaga) fauna of Crete (Greece). Contributions to Entomology 75 (2): 269-288, DOI: 10.3897/contrib.entomol.75.e158430
