identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
B0966550CC455A5A860F36FB59882E5A.text	B0966550CC455A5A860F36FB59882E5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allopeas clavulinum (Potiez & Michaud 1838) BOR	<div><p>Allopeas clavulinum (Potiez &amp; Michaud, 1838)</p><p>Fig. 7 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15416  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15418  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2921&amp;materialsCitation.latitude=5.4341" title="Search Plazi for locations around (long 100.2921/lat 5.4341)">Moon Gate Station 5 trail, Plot. 11</a>; 5.4341 ° N, 100.2921 ° E; 110 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15420  •   Penang Hill, Teluk Bahang-Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2159&amp;materialsCitation.latitude=5.418" title="Search Plazi for locations around (long 100.2159/lat 5.418)">Entrance of a durian farm, near Boulder Valley, Old rubber tree, Plot. 17</a>; 5.418 ° N, 100.2159 ° E; 180 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15415  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2263&amp;materialsCitation.latitude=5.4218" title="Search Plazi for locations around (long 100.2263/lat 5.4218)">Lam durian farm, Plot. 19</a>; 5.4218 ° N, 100.2263 ° E; 280 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15419  •  same data as for preceding; BOR/MOL 15421 •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15417  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15750  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15751  .</p><p>Remark.</p><p>New record for Penang Hill. It is possibly an introduced species and is widespread in Malaysia (Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/B0966550CC455A5A860F36FB59882E5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
E6B13B9CDD00585092B8C952C2145316.text	E6B13B9CDD00585092B8C952C2145316.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Allopeas gracile (T. Hutton 1834) BOR	<div><p>Allopeas gracile (T. Hutton, 1834)</p><p>Fig. 7 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15414  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2259&amp;materialsCitation.latitude=5.4198" title="Search Plazi for locations around (long 100.2259/lat 5.4198)">Lam durian farm, Plot. 13</a>; 5.4198 ° N, 100.2259 ° E; 290 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15546  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2187&amp;materialsCitation.latitude=5.4148" title="Search Plazi for locations around (long 100.2187/lat 5.4148)">Tropical fruit farm, Plot. 14</a>; 5.4148 ° N, 100.2187 ° E; 260 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15547  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.22&amp;materialsCitation.latitude=5.416" title="Search Plazi for locations around (long 100.22/lat 5.416)">Tropical fruit farm, Plot. 15</a>; 5.4160 ° N, 100.2200 ° E; 250 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15548  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2263&amp;materialsCitation.latitude=5.4218" title="Search Plazi for locations around (long 100.2263/lat 5.4218)">Lam durian farm, Plot. 19</a>; 5.4218 ° N, 100.2263 ° E; 280 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15284  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15749  .</p><p>Remarks.</p><p>More common in disturbed habitats on Penang Hill. It is possibly an introduced species and is widespread in Malaysia (Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/E6B13B9CDD00585092B8C952C2145316	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
51C9047E640E5D53B482BCC921C62F21.text	51C9047E640E5D53B482BCC921C62F21.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Alycaeus gibbosulus Stoliczka 1872	<div><p>Alycaeus gibbosulus Stoliczka, 1872</p><p>Remark.</p><p>Old record from the base of Penang Hill by Stoliczka (1872).</p></div>	https://treatment.plazi.org/id/51C9047E640E5D53B482BCC921C62F21	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
E99DB71D225F58369C29A61E47C5C572.text	E99DB71D225F58369C29A61E47C5C572.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidromus atricallosus (A. Gould 1843)	<div><p>Amphidromus atricallosus (A. Gould, 1843)</p><p>Remark.</p><p>This species was previously described by Stoliczka (1873) from Penang Hill but was not found during the sampling of this study.</p></div>	https://treatment.plazi.org/id/E99DB71D225F58369C29A61E47C5C572	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
970FF6340EA75969A44CC6B0957B3792.text	970FF6340EA75969A44CC6B0957B3792.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Amphidromus perversus (Linnaeus 1758)	<div><p>Amphidromus perversus (Linnaeus, 1758)</p><p>Remarks.</p><p>This species was previously described by Stoliczka (1873) from Penang Hill. However, it was not found during the sampling of this study.</p></div>	https://treatment.plazi.org/id/970FF6340EA75969A44CC6B0957B3792	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
26F32015AC1F5D07907D21E35021BEF4.text	26F32015AC1F5D07907D21E35021BEF4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopos punctata Collinge 1902	<div><p>Atopos punctata Collinge, 1902</p><p>Figs 18 B, 22 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, Trail from Viaduct to Claremont, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">after shelter, Plot. 27</a>; 5.4205 ° N, 100.2724 ° E; 500 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15228  .</p><p>Remarks.</p><p>This species was also previously recorded by Stoliczka (1873). It differs from  Atopos tourannensis by having a brown mantle.</p></div>	https://treatment.plazi.org/id/26F32015AC1F5D07907D21E35021BEF4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
DA698A0517A658D987281C376E2F7279.text	DA698A0517A658D987281C376E2F7279.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Atopos tourannensis (Souleyet 1852) BOR	<div><p>Atopos tourannensis (Souleyet, 1852)</p><p>Figs 18 A, 22 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang-Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2159&amp;materialsCitation.latitude=5.418" title="Search Plazi for locations around (long 100.2159/lat 5.418)">Entrance of a durian farm, near Boulder Valley, Old rubber tree, Plot. 17</a>; 5.418 ° N, 100.2159 ° E; 180 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15227  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. This species differs from  Atopos punctata by having a black mantle with white areas at both ends (head and tail) of the mantle.</p></div>	https://treatment.plazi.org/id/DA698A0517A658D987281C376E2F7279	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
DE80B908E07E53F4A8C1CF52DABC6C89.text	DE80B908E07E53F4A8C1CF52DABC6C89.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Bradybaena similaris (A. Ferussac 1821) BOR	<div><p>Bradybaena similaris (A. Férussac, 1821)</p><p>Figs 11 E, 21 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, Along Jalan Tunku Yahya Petra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2662&amp;materialsCitation.latitude=5.4225" title="Search Plazi for locations around (long 100.2662/lat 5.4225)">Random 6</a>.; 5.4225 ° N, 100.2662 ° E; 730 m a. s. l.; 20 Jun. 2023; T. S. Liew leg.; BOR/MOL 15772  .</p><p>Remarks.</p><p>This species was found by Stoliczka (1873) from Penang Hill. It is an introduced species and, as noted by Stoliczka (1873), it was only found in cocoa and palm plantations at the foothills and was not encountered in the forest. Similar observations were also made in this study.</p></div>	https://treatment.plazi.org/id/DE80B908E07E53F4A8C1CF52DABC6C89	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
C2F091FC4E4D59289942181ED191DF08.text	C2F091FC4E4D59289942181ED191DF08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Charopa perlata van Benthem Jutting 1959	<div><p>Charopa perlata van Benthem Jutting, 1959</p><p>Fig. 12 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2921&amp;materialsCitation.latitude=5.4341" title="Search Plazi for locations around (long 100.2921/lat 5.4341)">Moon Gate Station 5 trail, Plot. 11</a>; 5.4341 ° N, 100.2921 ° E; 110 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15482  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15560  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15753  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell characters match the description of the species originally described from Sumatra (van Benthem Jutting 1959).</p></div>	https://treatment.plazi.org/id/C2F091FC4E4D59289942181ED191DF08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
5DFD23C9B6FA571583B7CEC81F748CBF.text	5DFD23C9B6FA571583B7CEC81F748CBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Charopa undetermined-2	<div><p>Charopa Penang sp. 2</p><p>Fig. 12 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15483  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2259&amp;materialsCitation.latitude=5.4198" title="Search Plazi for locations around (long 100.2259/lat 5.4198)">Lam durian farm, Plot. 13</a>; 5.4198 ° N, 100.2259 ° E; 290 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15562  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.22&amp;materialsCitation.latitude=5.416" title="Search Plazi for locations around (long 100.22/lat 5.416)">Tropical fruit farm, Plot. 15</a>; 5.4160 ° N, 100.2200 ° E; 250 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15561  •   Penang Hill, Teluk Bahang-Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2159&amp;materialsCitation.latitude=5.418" title="Search Plazi for locations around (long 100.2159/lat 5.418)">Entrance of a durian farm, near Boulder Valley, Old rubber tree, Plot. 17</a>; 5.418 ° N, 100.2159 ° E; 180 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15484  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15761  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell spire is moderately raised. Teleoconch with densely spirally arranged pits above periphery, transitioning to densely placed spiral grooves on the subsequent whorls, which become more densely placed towards the periphery. Below periphery with densely placed spiral grooves that continue from the upper shell and fade away towards the umbilicus. The shell sculptures of this species are similar to  Microcystina Penang sp. 6, but it differs by having a smaller shell size, only half the size of  Microcystina Penang sp. 6 .</p></div>	https://treatment.plazi.org/id/5DFD23C9B6FA571583B7CEC81F748CBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
BFA47B62A03E59EBBC5EE76BBBB21EC8.text	BFA47B62A03E59EBBC5EE76BBBB21EC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Coptocheilus sectilabris (A. Gould 1844) BOR	<div><p>Coptocheilus sectilabris (A. Gould, 1844)</p><p>Figs 6 E, 19 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2645&amp;materialsCitation.latitude=5.4229" title="Search Plazi for locations around (long 100.2645/lat 5.4229)">Research trail, Random 4</a>.; 5.4229 ° N, 100.2645 ° E; 740 m a. s. l.; 27 Oct. 2022; M. Y. Lee leg.; BOR/MOL 15270  .</p><p>Remarks.</p><p>So far, all the records were found at the top of Penang Hill, specifically around the Habitat, but it is very rare elsewhere on Penang Hill. It is a widespread species in Indochina and Peninsular Malaysia (Maassen 2001; Tumpeesuwan and Panha 2008; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/BFA47B62A03E59EBBC5EE76BBBB21EC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
8C872F2255565ADEBEA0749CC3407493.text	8C872F2255565ADEBEA0749CC3407493.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclophorus malayanus (W. H. Benson 1852) BOR	<div><p>Cyclophorus malayanus (W. H. Benson, 1852)</p><p>Figs 5 A, 19 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2669&amp;materialsCitation.latitude=5.4248" title="Search Plazi for locations around (long 100.2669/lat 5.4248)">Research trail, Plot. 01</a>; 5.4248 ° N, 100.2669 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15163  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15119  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15126  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15140  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2648&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2648/lat 5.4223)">Habitat nature trail, Plot. 08</a>; 5.4223 ° N, 100.2648 ° E; 760 m a. s. l.; 6 Sept. 2022; J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15099  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2921&amp;materialsCitation.latitude=5.4341" title="Search Plazi for locations around (long 100.2921/lat 5.4341)">Moon Gate Station 5 trail, Plot. 11</a>; 5.4341 ° N, 100.2921 ° E; 110 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15100  •  same data as for preceding; BOR/MOL 15101 •  same data as for preceding; BOR/MOL 15152 •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.286&amp;materialsCitation.latitude=5.4396" title="Search Plazi for locations around (long 100.286/lat 5.4396)">Botanical garden, Plot. 12</a>; 5.4396 ° N, 100.2860 ° E; 85 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15091  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15221  •  same data as for preceding; BOR/MOL 15273 •   Penang Hill, Teluk Bahang-Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2159&amp;materialsCitation.latitude=5.418" title="Search Plazi for locations around (long 100.2159/lat 5.418)">Entrance of a durian farm, near Boulder Valley, Old rubber tree, Plot. 17</a>; 5.418 ° N, 100.2159 ° E; 180 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15222  •  same data as for preceding; BOR/MOL 15278 •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2265&amp;materialsCitation.latitude=5.4199" title="Search Plazi for locations around (long 100.2265/lat 5.4199)">Bukit Kerajaan ForestReserve next to Lam durian farm, Plot. 20</a>; 5.4199 ° N, 100.2265 ° E; 330 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15281  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15219  •  same data as for preceding; BOR/MOL 15232 •  same data as for preceding; BOR/MOL 15289 •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2575&amp;materialsCitation.latitude=5.39" title="Search Plazi for locations around (long 100.2575/lat 5.39)">Forest near Dam, Plot. 25</a>; 5.3900 ° N, 100.2575 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15295  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3977" title="Search Plazi for locations around (long 100.2612/lat 5.3977)">Forest near Dam, Plot. 26</a>; 5.3977 ° N, 100.2612 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15240  •  same data as for preceding; BOR/MOL 15296 •   Penang Hill, Trail - Moniot Road East, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15298  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15729  •  same data as for preceding; BOR/MOL 15766 •   Penang Hill, Moniot trail, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2574&amp;materialsCitation.latitude=5.4165" title="Search Plazi for locations around (long 100.2574/lat 5.4165)">Plot. 32</a>; 5.4165 ° N, 100.2574 ° E; 695 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15769  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15730  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4216" title="Search Plazi for locations around (long 100.2674/lat 5.4216)">Path B, Random 1. PathB</a>; 5.4216 ° N, 100.2674 ° E; 730 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15103  •  same data as for preceding; BOR/MOL 15107 •  same data as for preceding; BOR/MOL 15149 .</p><p>Remarks.</p><p>One of the most common large land snails at Penang Hill, with a shell width of up to 45 mm. It is also known from other localities in the northern part of Peninsular Malaysia. This species can be found in Perak as well (Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/8C872F2255565ADEBEA0749CC3407493	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
52C61D009CC853CA8BD37B2D6CE7AE50.text	52C61D009CC853CA8BD37B2D6CE7AE50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclophorus perdix subsp. perdix (Broderip & G. B. Sowerby I 1830)	<div><p>Cyclophorus perdix perdix (Broderip &amp; G. B. Sowerby I, 1830)</p><p>Figs 5 B, 19 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2645&amp;materialsCitation.latitude=5.4226" title="Search Plazi for locations around (long 100.2645/lat 5.4226)">Research trail, Plot. 03</a>; 5.4226 ° N, 100.2645 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15157  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15127  •  same data as for preceding; BOR/MOL 15118 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15135  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sep. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15141  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2218&amp;materialsCitation.latitude=5.442" title="Search Plazi for locations around (long 100.2218/lat 5.442)">Taman Rimba Teluk Bahang, Trail Stesen 3, Plot. 18</a>; 5.4420 ° N, 100.2218 ° E; 150 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15282  •   Penang Hill, Trail - Moniot Road East, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15299  •   Penang Hill, Moniot trail, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2574&amp;materialsCitation.latitude=5.4165" title="Search Plazi for locations around (long 100.2574/lat 5.4165)">Plot. 32</a>; 5.4165 ° N, 100.2574 ° E; 695 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15728  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15726  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4216" title="Search Plazi for locations around (long 100.2674/lat 5.4216)">Path B, Random 1. PathB</a>; 5.4216 ° N, 100.2674 ° E; 730 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15102  .</p><p>Remarks.</p><p>Less common than  Cyclophorus malayanus on Penang Hill, it was found to occur sympatrically with  Cyclophorus malayanus in this study. This species can also be found in Perak (Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/52C61D009CC853CA8BD37B2D6CE7AE50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
DA42C1C9819055FCA2B6A09786B5F73F.text	DA42C1C9819055FCA2B6A09786B5F73F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cyclotus solutus (Stoliczka 1872)	<div><p>Cyclotus solutus (Stoliczka, 1872)</p><p>Remarks.</p><p>This species is common in Perak (Foon et al. 2017) and was recorded by Stoliczka (1872) at the base of Penang Hill. However, it was not recorded in this study.</p></div>	https://treatment.plazi.org/id/DA42C1C9819055FCA2B6A09786B5F73F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
A4C1FB8D692B541CAFB374C0846736D7.text	A4C1FB8D692B541CAFB374C0846736D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina crosseana Godwin-Austen & Nevill 1879	<div><p>Diplommatina crosseana Godwin-Austen &amp; Nevill, 1879</p><p>Fig. 6 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.286&amp;materialsCitation.latitude=5.4396" title="Search Plazi for locations around (long 100.286/lat 5.4396)">Plot. 12</a>; 5.4396 ° N, 100.2860 ° E; 85 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15443  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15543  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2218&amp;materialsCitation.latitude=5.442" title="Search Plazi for locations around (long 100.2218/lat 5.442)">Taman Rimba Teluk Bahang, Trail Stesen 3, Plot. 18</a>; 5.4420 ° N, 100.2218 ° E; 150 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15444  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2637&amp;materialsCitation.latitude=5.3908" title="Search Plazi for locations around (long 100.2637/lat 5.3908)">Forest near farm, Plot. 21</a>; 5.3908 ° N, 100.2637 ° E; 340 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15544  .</p><p>Remark.</p><p>New record for Penang Hill. Previously known only from Perak (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/A4C1FB8D692B541CAFB374C0846736D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
E36BE821E71C546591E17C81610882B8.text	E36BE821E71C546591E17C81610882B8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplommatina undetermined-1	<div><p>Diplommatina Penang sp. 1</p><p>Fig. 6 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2913&amp;materialsCitation.latitude=5.4305" title="Search Plazi for locations around (long 100.2913/lat 5.4305)">Moon Gate Station 5 trail, Plot. 10</a>; 5.4305 ° N, 100.2913 ° E; 250 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15442  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2517&amp;materialsCitation.latitude=5.4203" title="Search Plazi for locations around (long 100.2517/lat 5.4203)">Trail to Western hill, Km 3.6, Plot. 23</a>; 5.4203 ° N, 100.2517 ° E; 750 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15542  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. Based on the only shell fragments collected, it is likely  Diplommatina ventriculus, which was originally described from Perak (von Möllendorff 1891).</p></div>	https://treatment.plazi.org/id/E36BE821E71C546591E17C81610882B8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
4A7B896866545B0295BF4A3CC17C69E9.text	4A7B896866545B0295BF4A3CC17C69E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gastrocopta palmira (Stoliczka 1873)	<div><p>Gastrocopta palmira (Stoliczka, 1873)</p><p>Remark.</p><p>Previously described and recorded by Stoliczka (1873) from Penang Hill, but not found in this study.</p></div>	https://treatment.plazi.org/id/4A7B896866545B0295BF4A3CC17C69E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
8F09C9BAB9B85326AED60FF22BFBDAE1.text	8F09C9BAB9B85326AED60FF22BFBDAE1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Gulella bicolor (T. Hutton 1834)	<div><p>Gulella bicolor (T. Hutton, 1834)</p><p>Remark.</p><p>Previously recorded by Stoliczka (1873) from the base of Penang Hill but not found in this study.</p></div>	https://treatment.plazi.org/id/8F09C9BAB9B85326AED60FF22BFBDAE1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
BA505C825B745084AAF275A2C6081E18.text	BA505C825B745084AAF275A2C6081E18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Helicarion permolle Stoliczka 1873	<div><p>Helicarion permolle Stoliczka, 1873</p><p>Figs 11 D, 21 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15117  •  same data as for preceding; BOR/MOL 15480 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15196  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15144  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.286&amp;materialsCitation.latitude=5.4396" title="Search Plazi for locations around (long 100.286/lat 5.4396)">Plot. 12</a>; 5.4396 ° N, 100.2860 ° E; 85 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15121  •   Penang Hill, Trail from Viaduct to Claremont, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">after shelter, Plot. 27</a>; 5.4205 ° N, 100.2724 ° E; 500 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15235  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15742  •  same data as for preceding; BOR/MOL 15768 •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15741  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4216" title="Search Plazi for locations around (long 100.2674/lat 5.4216)">Path B, Random 1. PathB</a>; 5.4216 ° N, 100.2674 ° E; 730 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15109  .</p><p>Remarks.</p><p>This species was previously described by Stoliczka (1873), and it is one of the most common semi-slugs from Penang Hill. It was listed as taxon inquirendum on MolluscaBase (2024) [https://molluscabase.org/aphia.php?p=taxdetails&amp;id=1337679]. We suggest that this is a valid species as the specimens fit the description in Stoliczka (1873).</p></div>	https://treatment.plazi.org/id/BA505C825B745084AAF275A2C6081E18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
447C34808AE251E4B2E448934E05DD94.text	447C34808AE251E4B2E448934E05DD94.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemiplecta cymatium (L. Pfeiffer 1856) BOR	<div><p>Hemiplecta cymatium (L. Pfeiffer, 1856)</p><p>Figs 8 C, 20 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2669&amp;materialsCitation.latitude=5.4248" title="Search Plazi for locations around (long 100.2669/lat 5.4248)">Research trail, Plot. 01</a>; 5.4248 ° N, 100.2669 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15161  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2649&amp;materialsCitation.latitude=5.4237" title="Search Plazi for locations around (long 100.2649/lat 5.4237)">Research
trail, Plot. 02</a>; 5.4237 ° N, 100.2649 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15158  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat
nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15136  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research
trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15137  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat
nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15087  •  same data as for preceding; BOR/MOL 15105 •  same data as for preceding; BOR/MOL 15142 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2648&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2648/lat 5.4223)">Habitat nature trail, Plot. 08</a>; 5.4223 ° N, 100.2648 ° E; 760 m a. s. l.; 6 Sept. 2022; J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15096  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.286&amp;materialsCitation.latitude=5.4396" title="Search Plazi for locations around (long 100.286/lat 5.4396)">Plot. 12</a>; 5.4396 ° N, 100.2860 ° E; 85 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15092  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15269  •   Penang Hill, Teluk Bahang-Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2159&amp;materialsCitation.latitude=5.418" title="Search Plazi for locations around (long 100.2159/lat 5.418)">Entrance
of a durian farm, near Boulder Valley, Old rubber tree, Plot. 17</a>; 5.418 ° N, 100.2159 ° E; 180 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15277  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest
near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15220  •  same data as for preceding; BOR/MOL 15290 •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2575&amp;materialsCitation.latitude=5.39" title="Search Plazi for locations around (long 100.2575/lat 5.39)">Forest near Dam, Plot. 25</a>; 5.3900 ° N, 100.2575 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15294  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15736  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15735  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4216" title="Search Plazi for locations around (long 100.2674/lat 5.4216)">Path B, Random 1. PathB</a>; 5.4216 ° N, 100.2674 ° E; 730 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15104  •  same data as for preceding; BOR/MOL 15148 .</p><p>Remarks.</p><p>One of the most common large land snails at Penang Hill, with a shell width of up to 40 mm. It is also known from other localities in Peninsular Malaysia. This species can also be found in Perak (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/447C34808AE251E4B2E448934E05DD94	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
C7F7EFF08A3357E49A60CA6C52ADFF76.text	C7F7EFF08A3357E49A60CA6C52ADFF76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kaliella barrakporensis (Reeve 1852) BOR	<div><p>Kaliella barrakporensis (Reeve, 1852)</p><p>Fig. 14 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15739  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. It is a widespread species in Malaysia (Vermeulen et al. 2015; Foon et al. 2017; Foon and Marzuki 2023).</p></div>	https://treatment.plazi.org/id/C7F7EFF08A3357E49A60CA6C52ADFF76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
D613301E214456AEA4632D1038E64B38.text	D613301E214456AEA4632D1038E64B38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kaliella scandens (J. C. Cox 1872) BOR	<div><p>Kaliella scandens (J. C. Cox, 1872)</p><p>Fig. 15 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2187&amp;materialsCitation.latitude=5.4148" title="Search Plazi for locations around (long 100.2187/lat 5.4148)">Tropical fruit farm, Plot. 14</a>; 5.4148 ° N, 100.2187 ° E; 260 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15566  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.22&amp;materialsCitation.latitude=5.416" title="Search Plazi for locations around (long 100.22/lat 5.416)">Tropical
fruit farm, Plot. 15</a>; 5.4160 ° N, 100.2200 ° E; 250 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15565  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2263&amp;materialsCitation.latitude=5.4218" title="Search Plazi for locations around (long 100.2263/lat 5.4218)">Lam
durian farm, Plot. 19</a>; 5.4218 ° N, 100.2263 ° E; 280 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15476  •   Penang Hill, Trail from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">Viaduct</a> to Claremont, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">after shelter, Plot. 27</a>; 5.4205 ° N, 100.2724 ° E; 500 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15475  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15744  •  same data as for preceding; BOR/MOL 15765 .</p><p>Remarks.</p><p>This is a new record for Penang Hill. This species is widespread, found from Sundaland to Australia and the Pacific Islands (Vermeulen et al. 2015).</p></div>	https://treatment.plazi.org/id/D613301E214456AEA4632D1038E64B38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
6DC1DEB97F2554EE9A70997D70D351BC.text	6DC1DEB97F2554EE9A70997D70D351BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kaliella undetermined-1	<div><p>Kaliella Penang sp. 1</p><p>Fig. 14 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15466  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15465  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell periphery is angular with a spiral thread visible above the suture of the penultimate whorl. Above the shell, the apical whorls are unknown, followed by regularly placed shallow radial ribs on the subsequent whorls. Below the shell, there are densely spaced spiral grooves at the periphery, which reduce towards the umbilicus.</p></div>	https://treatment.plazi.org/id/6DC1DEB97F2554EE9A70997D70D351BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
AE3AB79442655F468520EDC0774307DB.text	AE3AB79442655F468520EDC0774307DB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kaliella undetermined-2	<div><p>Kaliella Penang sp. 2</p><p>Fig. 14 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15468  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2648&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2648/lat 5.4223)">Habitat nature trail, Plot. 08</a>; 5.4223 ° N, 100.2648 ° E; 760 m a. s. l.; 6 Sept. 2022; J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15469  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2517&amp;materialsCitation.latitude=5.4203" title="Search Plazi for locations around (long 100.2517/lat 5.4203)">Trail to Western hill, Km 3.6, Plot. 23</a>; 5.4203 ° N, 100.2517 ° E; 750 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15563  •   Penang Hill, Trail - Moniot Road East, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15467  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2574&amp;materialsCitation.latitude=5.4165" title="Search Plazi for locations around (long 100.2574/lat 5.4165)">Moniot trail, Plot. 32</a>; 5.4165 ° N, 100.2574 ° E; 695 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15752  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell is convex discus-shaped. The shell periphery is angular with a spiral thread. Above the shell, the apical whorls have shallow spiral ribs crossed by growth lines, transitioning to regularly placed prominent radial ribs on the subsequent whorls. Below the shell, radial ribs continue from above at the periphery, transitioning to spaced spiral grooves which reduce towards the umbilicus.</p></div>	https://treatment.plazi.org/id/AE3AB79442655F468520EDC0774307DB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
27C73C8902D0594AB24F5F4699BA0D99.text	27C73C8902D0594AB24F5F4699BA0D99.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kaliella undetermined-4	<div><p>Kaliella Penang sp. 4</p><p>Fig. 14 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2645&amp;materialsCitation.latitude=5.4226" title="Search Plazi for locations around (long 100.2645/lat 5.4226)">Research trail, Plot. 03</a>; 5.4226 ° N, 100.2645 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15473  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15474  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15470  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2575&amp;materialsCitation.latitude=5.39" title="Search Plazi for locations around (long 100.2575/lat 5.39)">Forest near Dam, Plot. 25</a>; 5.3900 ° N, 100.2575 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15471  •   Penang Hill, Trail - Moniot Road East, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15472  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell is conical in shape with an angular periphery adorned with a spiral thread. Teleoconch with elevated radial ribs above periphery that transition to densely placed fine spiral ribs on the subsequent whorls, crossed by irregular growth lines. Below periphery with less densely spaced spiral grooves at the periphery, which diminish towards the umbilicus.</p></div>	https://treatment.plazi.org/id/27C73C8902D0594AB24F5F4699BA0D99	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
D9278023E466520C8BC1B1DBAB5AF875.text	D9278023E466520C8BC1B1DBAB5AF875.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kaliella undetermined-5	<div><p>Kaliella Penang sp. 5</p><p>Fig. 15 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.22&amp;materialsCitation.latitude=5.416" title="Search Plazi for locations around (long 100.22/lat 5.416)">Tropical fruit farm, Plot. 15</a>; 5.4160 ° N, 100.2200 ° E; 250 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15564  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell is flat conical with an angular periphery adorned with a visible spiral thread above the suture of the penultimate whorl. Teleoconch with elevated radial ribs above periphery that transition to densely placed radial ribs on the subsequent whorls. Below periphery with dense radial ribs continuing from above and transitioning to spaced spiral grooves that diminish towards the umbilicus.</p></div>	https://treatment.plazi.org/id/D9278023E466520C8BC1B1DBAB5AF875	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
2366391C699B585F880E2F12A77C6FE5.text	2366391C699B585F880E2F12A77C6FE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Laevicaulis alte (A. Ferussac 1822) BOR	<div><p>Laevicaulis alte (A. Férussac, 1822)</p><p>Figs 18 C, 22 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4216" title="Search Plazi for locations around (long 100.2674/lat 5.4216)">Path B, Random 1. PathB</a>; 5.4216 ° N, 100.2674 ° E; 730 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15111  •   Penang Hill, Plaza, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2689&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2689/lat 5.4247)">Random 2. Plaza</a>; 5.4247 ° N, 100.2689 ° E; 740 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15112  •   Penang Hill, Along Jalan Tunku Yahya Petra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2662&amp;materialsCitation.latitude=5.4225" title="Search Plazi for locations around (long 100.2662/lat 5.4225)">Random 6</a>.; 5.4225 ° N, 100.2662 ° E; 730 m a. s. l.; 20 Jun. 2023; T. S. Liew leg.; BOR/MOL 15773  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. So far, this species has only been found in human-altered habitats. The most reliable method for determining species identity involves detailed anatomical examination. However, as this specimen is a juvenile, some identifying features may not be fully developed. Based on the available evidence, we tentatively identify it as  L. alte .</p></div>	https://treatment.plazi.org/id/2366391C699B585F880E2F12A77C6FE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
401E5B6DFF735D3C803A740348E1586E.text	401E5B6DFF735D3C803A740348E1586E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagocheilus striolatus Stoliczka 1872	<div><p>Lagocheilus striolatus Stoliczka, 1872</p><p>Fig. 5 F</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15200  .</p><p>Remark.</p><p>A rare species on Penang Hill.</p></div>	https://treatment.plazi.org/id/401E5B6DFF735D3C803A740348E1586E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
6F1B86BB7CB7542FBA1DF08C795B5BEE.text	6F1B86BB7CB7542FBA1DF08C795B5BEE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagocheilus trochoides (Stoliczka 1872) BOR	<div><p>Lagocheilus trochoides (Stoliczka, 1872)</p><p>Figs 5 E, 19 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2669&amp;materialsCitation.latitude=5.4248" title="Search Plazi for locations around (long 100.2669/lat 5.4248)">Research trail, Plot. 01</a>; 5.4248 ° N, 100.2669 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15081  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15134  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15084  •  same data as for preceding; BOR/MOL 15085 •  same data as for preceding; BOR/MOL 15139 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15143  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2648&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2648/lat 5.4223)">Habitat nature trail, Plot. 08</a>; 5.4223 ° N, 100.2648 ° E; 760 m a. s. l.; 6 Sept. 2022; J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15156  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15540  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2265&amp;materialsCitation.latitude=5.4199" title="Search Plazi for locations around (long 100.2265/lat 5.4199)">Bukit Kerajaan ForestReserve next to Lam durian farm, Plot. 20</a>; 5.4199 ° N, 100.2265 ° E; 330 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15425  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15287  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">Trail from Viaduct to Claremont, after shelter, Plot. 27</a>; 5.4205 ° N, 100.2724 ° E; 500 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15426  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15738  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15737  .</p><p>Remark.</p><p>This species is more common than  L. striolatus at Penang Hill.</p></div>	https://treatment.plazi.org/id/6F1B86BB7CB7542FBA1DF08C795B5BEE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
2CF6B8FB0AA15F90BF5662A0CA2672D0.text	2CF6B8FB0AA15F90BF5662A0CA2672D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lagocheilus undetermined-2	<div><p>Lagocheilus Penang sp. 2</p><p>Fig. 5 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15430  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15428  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2921&amp;materialsCitation.latitude=5.4341" title="Search Plazi for locations around (long 100.2921/lat 5.4341)">Moon Gate Station 5 trail, Plot. 11</a>; 5.4341 ° N, 100.2921 ° E; 110 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15427  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15541  •   Penang Hill, Teluk Bahang-Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2159&amp;materialsCitation.latitude=5.418" title="Search Plazi for locations around (long 100.2159/lat 5.418)">Entrance of a durian farm, near Boulder Valley, Old rubber tree, Plot. 17</a>; 5.418 ° N, 100.2159 ° E; 180 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15429  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. This species differs from  L. trochoides by having more, less prominent spiral ribs, and from  L. striolatus by having more spiral ribs. It also differs from  L. garelli (Souleyet, 1852), which was previously recorded and described from Penang, by having more rapidly expanding whorl size.</p></div>	https://treatment.plazi.org/id/2CF6B8FB0AA15F90BF5662A0CA2672D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
721EBD42328D5A6DB72DE1C1125D8FFB.text	721EBD42328D5A6DB72DE1C1125D8FFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lissachatina fulica (Bowdich 1822) BOR	<div><p>Lissachatina fulica (Bowdich, 1822)</p><p>Figs 7 E, 20 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15486  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15131  •  same data as for preceding; BOR/MOL 15195 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15088  •  same data as for preceding; MOL 15147 •  same data as for preceding; BOR/MOL 15423 •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2259&amp;materialsCitation.latitude=5.4198" title="Search Plazi for locations around (long 100.2259/lat 5.4198)">Lam durian farm, Plot. 13</a>; 5.4198 ° N, 100.2259 ° E; 290 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15271  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.22&amp;materialsCitation.latitude=5.416" title="Search Plazi for locations around (long 100.22/lat 5.416)">Tropical fruit farm, Plot. 15</a>; 5.4160 ° N, 100.2200 ° E; 250 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15272  •   Penang Hill, Teluk Bahang-Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2159&amp;materialsCitation.latitude=5.418" title="Search Plazi for locations around (long 100.2159/lat 5.418)">Entrance of a durian farm, near Boulder Valley, Old rubber tree, Plot. 17</a>; 5.418 ° N, 100.2159 ° E; 180 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15412  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2263&amp;materialsCitation.latitude=5.4218" title="Search Plazi for locations around (long 100.2263/lat 5.4218)">Lam durian farm, Plot. 19</a>; 5.4218 ° N, 100.2263 ° E; 280 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15230  •  same data as for preceding; BOR/MOL 15283 •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15731  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15727  .</p><p>Remarks.</p><p>New record for Penang Hill. This species was likely introduced to Malaysia as earlier as 1911 in Kedah, and 1922 in Penang (South 1926; Berry and Chan 1968). A widespread introduced species in Malaysia.</p></div>	https://treatment.plazi.org/id/721EBD42328D5A6DB72DE1C1125D8FFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
BB4B341367225281B428C8A1BD2CFB05.text	BB4B341367225281B428C8A1BD2CFB05.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Meghimatium pictum (Stoliczka 1873)	<div><p>Meghimatium pictum (Stoliczka, 1873)</p><p>Fig. 22 E</p><p>Remarks.</p><p>This species was previously described by Stoliczka (1873) from the base of the hill in the north, at ca 100 feet above sea level. It was not found during the sampling of this study. However, this species was recently photographed on Penang Hill (Fig. 22 E). It is thought to be a widespread species found from East Asia to Indochina (Tsai et al. 2011 Inkhavilay et al. 2019; Ito et al. 2023).</p></div>	https://treatment.plazi.org/id/BB4B341367225281B428C8A1BD2CFB05	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
D8CCB3B5E7465A6189D1BDA7EA15FC65.text	D8CCB3B5E7465A6189D1BDA7EA15FC65.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microcystina undetermined-1	<div><p>Microcystina Penang sp. 1</p><p>Fig. 8 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15453  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15455  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15448  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2648&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2648/lat 5.4223)">Habitat nature trail, Plot. 08</a>; 5.4223 ° N, 100.2648 ° E; 760 m a. s. l.; 6 Sept. 2022; J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15450  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2857&amp;materialsCitation.latitude=5.4216" title="Search Plazi for locations around (long 100.2857/lat 5.4216)">Moon Gate Station 5 trail, Plot. 09</a>; 5.4216 ° N, 100.2857 ° E; 410 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15445  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2913&amp;materialsCitation.latitude=5.4305" title="Search Plazi for locations around (long 100.2913/lat 5.4305)">Moon Gate Station 5 trail, Plot. 10</a>; 5.4305 ° N, 100.2913 ° E; 250 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15451  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2921&amp;materialsCitation.latitude=5.4341" title="Search Plazi for locations around (long 100.2921/lat 5.4341)">Moon Gate Station 5 trail, Plot. 11</a>; 5.4341 ° N, 100.2921 ° E; 110 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15449  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2259&amp;materialsCitation.latitude=5.4198" title="Search Plazi for locations around (long 100.2259/lat 5.4198)">Lam durian farm, Plot. 13</a>; 5.4198 ° N, 100.2259 ° E; 290 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15558  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15557  •   Penang Hill, Teluk Bahang-Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2159&amp;materialsCitation.latitude=5.418" title="Search Plazi for locations around (long 100.2159/lat 5.418)">Entrance of a durian farm, near Boulder Valley, Old rubber tree, Plot. 17</a>; 5.418 ° N, 100.2159 ° E; 180 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15447  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2218&amp;materialsCitation.latitude=5.442" title="Search Plazi for locations around (long 100.2218/lat 5.442)">Taman Rimba Teluk Bahang, Trail Stesen 3, Plot. 18</a>; 5.4420 ° N, 100.2218 ° E; 150 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15452  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2637&amp;materialsCitation.latitude=5.3908" title="Search Plazi for locations around (long 100.2637/lat 5.3908)">Forest near farm, Plot. 21</a>; 5.3908 ° N, 100.2637 ° E; 340 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15555  •   Penang Hill, Trail to Western hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2494&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2494/lat 5.4223)">Junction to Teluk Bahang from Penang Hill, Plot. 22</a>; 5.4223 ° N, 100.2494 ° E; 760 m a. s. l.; 24 Oct. 2022; T. S. Liew, J, Dulipat, M. Y. Lee leg.; BOR/MOL 15556  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2517&amp;materialsCitation.latitude=5.4203" title="Search Plazi for locations around (long 100.2517/lat 5.4203)">Trail to Western hill, Km 3.6, Plot. 23</a>; 5.4203 ° N, 100.2517 ° E; 750 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15554  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3977" title="Search Plazi for locations around (long 100.2612/lat 5.3977)">Forest near Dam, Plot. 26</a>; 5.3977 ° N, 100.2612 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15454  •   Penang Hill, Trail - Moniot Road East, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15446  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15754  .</p><p>Remarks.</p><p>This is a new record for Penang Hill of a widespread species. The shell has a slightly raised spire with a height less than half of the last whorl height. The upper shell, including the apical and subsequent whorls, shows growth lines, while the lower shell exhibits faint spiral lines.</p></div>	https://treatment.plazi.org/id/D8CCB3B5E7465A6189D1BDA7EA15FC65	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
F7B4952B3B255B0ABCE91EC73901F42C.text	F7B4952B3B255B0ABCE91EC73901F42C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microcystina undetermined-2	<div><p>Microcystina Penang sp. 2</p><p>Fig. 9 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2669&amp;materialsCitation.latitude=5.4248" title="Search Plazi for locations around (long 100.2669/lat 5.4248)">Research trail, Plot. 01</a>; 5.4248 ° N, 100.2669 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15456  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15458  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2575&amp;materialsCitation.latitude=5.39" title="Search Plazi for locations around (long 100.2575/lat 5.39)">Forest near Dam, Plot. 25</a>; 5.3900 ° N, 100.2575 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15459  •   Penang Hill, Trail - Moniot Road East, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15457  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15763  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell spire is highly raised, with a height more than half of the last whorl height. This species has the most convex shell compared to other  Microcystina species from Penang Hill. On the upper shell, the apical whorls have elevated spiral ribs that continue onto subsequent whorls, which are crossed by radial ribs forming granule-like sculptures. Below the shell, there are spaced grooves with spiral lines at the periphery, which reduce towards the umbilicus.</p></div>	https://treatment.plazi.org/id/F7B4952B3B255B0ABCE91EC73901F42C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
45B384260F6C5804BE8F8BB777AB9739.text	45B384260F6C5804BE8F8BB777AB9739.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microcystina undetermined-4	<div><p>Microcystina Penang sp. 4</p><p>Fig. 9 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2645&amp;materialsCitation.latitude=5.4226" title="Search Plazi for locations around (long 100.2645/lat 5.4226)">Research trail, Plot. 03</a>; 5.4226 ° N, 100.2645 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15462  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.286&amp;materialsCitation.latitude=5.4396" title="Search Plazi for locations around (long 100.286/lat 5.4396)">Plot. 12</a>; 5.4396 ° N, 100.2860 ° E; 85 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15463  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2574&amp;materialsCitation.latitude=5.4165" title="Search Plazi for locations around (long 100.2574/lat 5.4165)">Moniot trail, Plot. 32</a>; 5.4165 ° N, 100.2574 ° E; 695 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15755  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell spire is moderately raised, with a height less than half of the last whorl height. On the upper shell, the apical whorls are smooth, and subsequent whorls have regularly spaced deep radial grooves. This species has the most prominent radial grooves compared to other  Microcystina species from Penang Hill. Below the shell, there are regularly spaced, deep, radial grooves that continue from the upper shell.</p></div>	https://treatment.plazi.org/id/45B384260F6C5804BE8F8BB777AB9739	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
58EA9368F87952DA8C2C4FEDA56DD2BC.text	58EA9368F87952DA8C2C4FEDA56DD2BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microcystina undetermined-5	<div><p>Microcystina Penang sp. 5</p><p>Fig. 9 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3977" title="Search Plazi for locations around (long 100.2612/lat 5.3977)">Forest near Dam, Plot. 26</a>; 5.3977 ° N, 100.2612 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15464  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15762  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell spire is hardly raised, with a height less than half of the last whorl height. On the upper shell, there are densely spirally arranged pits on the apical and subsequent whorls. This species is unique among the  Microcystina species from Penang Hill in having densely spirally arranged pits on the whorls near the periphery. Below the shell, there are densely spirally arranged pits that continue from the upper shell and fade away towards the umbilicus.</p></div>	https://treatment.plazi.org/id/58EA9368F87952DA8C2C4FEDA56DD2BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
CA378F8A47985D029FD02BBB84B10A9B.text	CA378F8A47985D029FD02BBB84B10A9B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microcystina undetermined-6	<div><p>Microcystina Penang sp. 6</p><p>Fig. 9 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2259&amp;materialsCitation.latitude=5.4198" title="Search Plazi for locations around (long 100.2259/lat 5.4198)">Lam durian farm, Plot. 13</a>; 5.4198 ° N, 100.2259 ° E; 290 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15559  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15756  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. The shell spire is slightly raised, and the shell whorls are more rapidly expanded compared to other  Microcystina species from Penang Hill. On the upper shell, there are densely spirally arranged pits on the apical whorls, transitioning to densely placed spiral grooves on the subsequent whorls, which become more densely placed towards the periphery. Below the shell, there are densely placed spiral grooves that continue from the upper shell and fade away towards the umbilicus.</p></div>	https://treatment.plazi.org/id/CA378F8A47985D029FD02BBB84B10A9B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
4FBFF62B9E035B4888EC1DD3E7543363.text	4FBFF62B9E035B4888EC1DD3E7543363.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microcystis palmicola Stoliczka 1873	<div><p>Microcystis palmicola Stoliczka, 1873</p><p>Remark.</p><p>This species was previously described by Stoliczka (1873) from the base of Penang Hill but was not found during the sampling of this study.</p></div>	https://treatment.plazi.org/id/4FBFF62B9E035B4888EC1DD3E7543363	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
79CE69893838526A80D902FCE267FF71.text	79CE69893838526A80D902FCE267FF71.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Oospira penangensis (Stoliczka 1873) BOR	<div><p>Oospira penangensis (Stoliczka, 1873)</p><p>Figs 8 A, 20 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15223  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2575&amp;materialsCitation.latitude=5.39" title="Search Plazi for locations around (long 100.2575/lat 5.39)">Forest near Dam, Plot. 25</a>; 5.3900 ° N, 100.2575 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15224  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Trail - Moniot Road East, Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15243  .</p><p>Remark.</p><p>This species was described and recorded by Stoliczka (1873) from Penang Hill, and it can also be found in Perak (Maassen 2001).</p></div>	https://treatment.plazi.org/id/79CE69893838526A80D902FCE267FF71	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
5B4D3629788151AC963448341634BCDB.text	5B4D3629788151AC963448341634BCDB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Opisthoporus penangensis Stoliczka 1872	<div><p>Opisthoporus penangensis Stoliczka, 1872</p><p>Fig. 6 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15275  .</p><p>Remarks.</p><p>Originally described from base of Penang Hill (Stoliczka 1872) and recorded in only one plot of this study. This species has also been recorded in the northern part of Peninsular Malaysia and southern Thailand (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/5B4D3629788151AC963448341634BCDB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
C8675D6B57F55DF29FAA307D4DCE2827.text	C8675D6B57F55DF29FAA307D4DCE2827.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Parmarion martensi Simroth 1893	<div><p>Parmarion martensi Simroth, 1893</p><p>Figs 10 A, 20 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.22&amp;materialsCitation.latitude=5.416" title="Search Plazi for locations around (long 100.22/lat 5.416)">Tropical fruit farm, Plot. 15</a>; 5.4160 ° N, 100.2200 ° E; 250 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15229  •  same data as for preceding; BOR/MOL 15239 .</p><p>Remark.</p><p>This is a new record for Penang Hill. This species is widespread in Malaysia and might have been introduced to this location (Maassen 2001).</p></div>	https://treatment.plazi.org/id/C8675D6B57F55DF29FAA307D4DCE2827	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
4BEE8EC6610F55B2838D66F373798297.text	4BEE8EC6610F55B2838D66F373798297.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Paropeas tchehelense (de Morgan 1885)	<div><p>Paropeas tchehelense (de Morgan, 1885)</p><p>Fig. 8 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2669&amp;materialsCitation.latitude=5.4248" title="Search Plazi for locations around (long 100.2669/lat 5.4248)">Research trail, Plot. 01</a>; 5.4248 ° N, 100.2669 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15162  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2648&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2648/lat 5.4223)">Habitat nature trail, Plot. 08</a>; 5.4223 ° N, 100.2648 ° E; 760 m a. s. l.; 6 Sept. 2022; J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15097  .</p><p>Remark.</p><p>New record from Penang Hill. This species is also recorded from the mainland of Peninsular Malaysia and southern Thailand (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/4BEE8EC6610F55B2838D66F373798297	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
7D7F43FFC8665EC689A113123E32AEE2.text	7D7F43FFC8665EC689A113123E32AEE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Phaedusa filicostata (Stoliczka 1873)	<div><p>Phaedusa filicostata (Stoliczka, 1873)</p><p>Remark.</p><p>Previously described and recorded by Stoliczka (1873) from Penang Hill but not found in this study.</p></div>	https://treatment.plazi.org/id/7D7F43FFC8665EC689A113123E32AEE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
78F76A091DCE5CEDBD00749A2A141E8A.text	78F76A091DCE5CEDBD00749A2A141E8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philalanka carinifera (Stoliczka 1873) BOR	<div><p>Philalanka carinifera (Stoliczka, 1873)</p><p>Fig. 13 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2669&amp;materialsCitation.latitude=5.4248" title="Search Plazi for locations around (long 100.2669/lat 5.4248)">Research trail, Plot. 01</a>; 5.4248 ° N, 100.2669 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15434  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat
nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15435  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat
nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15433  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2648&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2648/lat 5.4223)">Habitat
nature trail, Plot. 08</a>; 5.4223 ° N, 100.2648 ° E; 760 m a. s. l.; 6 Sept. 2022; J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15437  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest
near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15436  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2575&amp;materialsCitation.latitude=5.39" title="Search Plazi for locations around (long 100.2575/lat 5.39)">Forest
near Dam, Plot. 25</a>; 5.3900 ° N, 100.2575 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15440  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3977" title="Search Plazi for locations around (long 100.2612/lat 5.3977)">Forest
near Dam, Plot. 26</a>; 5.3977 ° N, 100.2612 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15438  •   Penang Hill, Trail from <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">Viaduct</a> to Claremont, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">after shelter, Plot. 27</a>; 5.4205 ° N, 100.2724 ° E; 500 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15441  •   Penang Hill, Trail - Moniot Road East, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15439  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15746  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15745  •  same data as for preceding; BOR/MOL 15764 .</p><p>Remarks.</p><p>This species was previously described by Stoliczka (1873) from Penang Hill. It differs from the other two  Philalanka species by having three spiral ribs on the penultimate whorls.</p></div>	https://treatment.plazi.org/id/78F76A091DCE5CEDBD00749A2A141E8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
B954EFA52453580EB8363ADE22C9B077.text	B954EFA52453580EB8363ADE22C9B077.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philalanka kusana (Aldrich 1889) BOR	<div><p>Philalanka kusana (Aldrich, 1889)</p><p>Fig. 13 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15760  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. This species differs from the other two  Philalanka species by having no more than two spiral ribs on the penultimate whorls.</p></div>	https://treatment.plazi.org/id/B954EFA52453580EB8363ADE22C9B077	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
5D8C50C8E41E5FDDA6B03A760DABE37F.text	5D8C50C8E41E5FDDA6B03A760DABE37F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Philalanka undetermined-2	<div><p>Philalanka Penang sp. 2</p><p>Fig. 13 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15539  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. This species differs from the other two  Philalanka species by having more than seven spiral ribs on the penultimate whorls.</p></div>	https://treatment.plazi.org/id/5D8C50C8E41E5FDDA6B03A760DABE37F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
F65149DDAC7D5B87BEBEEB29B50E64EA.text	F65149DDAC7D5B87BEBEEB29B50E64EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pseudoplecta bijuga (Stoliczka 1873) BOR	<div><p>Pseudoplecta bijuga (Stoliczka, 1873)</p><p>Figs 17 B, 22 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2669&amp;materialsCitation.latitude=5.4248" title="Search Plazi for locations around (long 100.2669/lat 5.4248)">Research trail, Plot. 01</a>; 5.4248 ° N, 100.2669 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15114  •  same data as for preceding; BOR/MOL 15114 •  same data as for preceding; BOR/MOL 15159 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2649&amp;materialsCitation.latitude=5.4237" title="Search Plazi for locations around (long 100.2649/lat 5.4237)">Research trail, Plot. 02</a>; 5.4237 ° N, 100.2649 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15113  •  same data as for preceding; BOR/MOL 15197 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2645&amp;materialsCitation.latitude=5.4226" title="Search Plazi for locations around (long 100.2645/lat 5.4226)">Research trail, Plot. 03</a>; 5.4226 ° N, 100.2645 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15106  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15116  •  same data as for preceding; BOR/MOL 15128 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15125  •  same data as for preceding; BOR/MOL 15133 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15082  •  same data as for preceding; BOR/MOL 15083 •  same data as for preceding; BOR/MOL 15138 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15086  •  same data as for preceding; BOR/MOL 15089 •  same data as for preceding; BOR/MOL 15146 •  same data as for preceding; BOR/MOL 15431 •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2648&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2648/lat 5.4223)">Habitat nature trail, Plot. 08</a>; 5.4223 ° N, 100.2648 ° E; 760 m a. s. l.; 6 Sept. 2022; J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15095  •  same data as for preceding; BOR/MOL 15155 •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2913&amp;materialsCitation.latitude=5.4305" title="Search Plazi for locations around (long 100.2913/lat 5.4305)">Moon Gate Station 5 trail, Plot. 10</a>; 5.4305 ° N, 100.2913 ° E; 250 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15123  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2921&amp;materialsCitation.latitude=5.4341" title="Search Plazi for locations around (long 100.2921/lat 5.4341)">Moon Gate Station 5 trail, Plot. 11</a>; 5.4341 ° N, 100.2921 ° E; 110 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15093  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.286&amp;materialsCitation.latitude=5.4396" title="Search Plazi for locations around (long 100.286/lat 5.4396)">Plot. 12</a>; 5.4396 ° N, 100.2860 ° E; 85 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15090  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.22&amp;materialsCitation.latitude=5.416" title="Search Plazi for locations around (long 100.22/lat 5.416)">Tropical fruit farm, Plot. 15</a>; 5.4160 ° N, 100.2200 ° E; 250 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15276  •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2212&amp;materialsCitation.latitude=5.4429" title="Search Plazi for locations around (long 100.2212/lat 5.4429)">Taman Rimba Teluk Bahang, Trail, Simpang 6, Plot. 16</a>; 5.4429 ° N, 100.2212 ° E; 120 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15268  •  same data as for preceding; BOR/MOL 15274 •   Penang Hill, Teluk Bahang, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2218&amp;materialsCitation.latitude=5.442" title="Search Plazi for locations around (long 100.2218/lat 5.442)">Taman Rimba Teluk Bahang, Trail Stesen 3, Plot. 18</a>; 5.4420 ° N, 100.2218 ° E; 150 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15236  •  same data as for preceding; BOR/MOL 15280 •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2265&amp;materialsCitation.latitude=5.4199" title="Search Plazi for locations around (long 100.2265/lat 5.4199)">Bukit Kerajaan ForestReserve next to Lam durian farm, Plot. 20</a>; 5.4199 ° N, 100.2265 ° E; 330 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15217  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2517&amp;materialsCitation.latitude=5.4203" title="Search Plazi for locations around (long 100.2517/lat 5.4203)">Trail to Western hill, Km 3.6, Plot. 23</a>; 5.4203 ° N, 100.2517 ° E; 750 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15218  •  same data as for preceding; BOR/MOL 15286 •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15241  •  same data as for preceding; BOR/MOL 15288 •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2575&amp;materialsCitation.latitude=5.39" title="Search Plazi for locations around (long 100.2575/lat 5.39)">Forest near Dam, Plot. 25</a>; 5.3900 ° N, 100.2575 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15237  •  same data as for preceding; BOR/MOL 15293 •  same data as for preceding; BOR/MOL 15432 •   Penang Hill, Trail from Viaduct to Claremont, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">after shelter, Plot. 27</a>; 5.4205 ° N, 100.2724 ° E; 500 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15234  •  same data as for preceding; BOR/MOL 15297 •   Penang Hill, Trail - Moniot Road East, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15231  •  same data as for preceding; BOR/MOL 15300 •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2574&amp;materialsCitation.latitude=5.4165" title="Search Plazi for locations around (long 100.2574/lat 5.4165)">Moniot trail, Plot. 32</a>; 5.4165 ° N, 100.2574 ° E; 695 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15733  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15734  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4216" title="Search Plazi for locations around (long 100.2674/lat 5.4216)">Path B, Random 1. PathB</a>; 5.4216 ° N, 100.2674 ° E; 730 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15150  •   Penang Hill, Bukit Laksamana, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2362&amp;materialsCitation.latitude=5.4239" title="Search Plazi for locations around (long 100.2362/lat 5.4239)">Random 5</a>.; 5.4239 ° N, 100.2362 ° E; 710 m a. s. l.; 3 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15770  .</p><p>Remark.</p><p>One of the most common large land snails at Penang Hill, with a shell width of up to 22 mm. It is also known from other localities in Peninsular Malaysia and Thailand (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/F65149DDAC7D5B87BEBEEB29B50E64EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
AF8E6001549E57E7B7D37F0DC7563377.text	AF8E6001549E57E7B7D37F0DC7563377.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pupina aureola Stoliczka 1872	<div><p>Pupina aureola Stoliczka, 1872</p><p>Figs 7 A, 19 E</p><p>Material examined.</p><p>Malaysia • Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2913&amp;materialsCitation.latitude=5.4305" title="Search Plazi for locations around (long 100.2913/lat 5.4305)">Moon Gate Station 5 trail, Plot. 10</a>; 5.4305 ° N, 100.2913 ° E; 250 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15124  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2921&amp;materialsCitation.latitude=5.4341" title="Search Plazi for locations around (long 100.2921/lat 5.4341)">Moon Gate Station 5 trail, Plot. 11</a>; 5.4341 ° N, 100.2921 ° E; 110 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15094  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Trail - Moniot Road East, Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15225  .</p><p>Remark.</p><p>This species is also recorded from the northern part of Peninsular Malaysia and southern Thailand (Maassen 2001; Jirapatrasilp et al. 2022).</p></div>	https://treatment.plazi.org/id/AF8E6001549E57E7B7D37F0DC7563377	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
A2B868E459E85DD4A1909610E7BF6452.text	A2B868E459E85DD4A1909610E7BF6452.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pupisoma orcella (Stoliczka 1873)	<div><p>Pupisoma orcella (Stoliczka, 1873)</p><p>Remark.</p><p>Previous described and recorded by Stoliczka (1873) from the base of Penang Hill but not found in this study.</p></div>	https://treatment.plazi.org/id/A2B868E459E85DD4A1909610E7BF6452	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
EBE84A12B7815E71B504B7ECF299F136.text	EBE84A12B7815E71B504B7ECF299F136.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Quantula striata (J. E. Gray 1834) BOR	<div><p>Quantula striata (J. E. Gray, 1834)</p><p>Fig. 17 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15732  •  same data as for preceding; BOR/MOL 15767 •   Penang Hill, Air Itam Farm, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2664&amp;materialsCitation.latitude=5.3891" title="Search Plazi for locations around (long 100.2664/lat 5.3891)">Batu Panay road, Random 3. Farm</a>; 5.3891 ° N, 100.2664 ° E; 270 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15301  .</p><p>Remarks.</p><p>One of the most common large land snails at vegetable farms of Penang Hill with a shell width of up to 28 mm. It is also known from other localities in Peninsular Malaysia, Sundaland, and the Philippines (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/EBE84A12B7815E71B504B7ECF299F136	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
ABBEFC35FCFB52AD88A20913BCADC1C0.text	ABBEFC35FCFB52AD88A20913BCADC1C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhaphaulus lorraini L. Pfeiffer 1856	<div><p>Rhaphaulus lorraini L. Pfeiffer, 1856</p><p>Figs 7 B, 19 F</p><p>Material examined.</p><p>Malaysia • Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2517&amp;materialsCitation.latitude=5.4203" title="Search Plazi for locations around (long 100.2517/lat 5.4203)">Trail to Western hill, Km 3.6, Plot. 23</a>; 5.4203 ° N, 100.2517 ° E; 750 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15226  •  same data as for preceding; BOR/MOL 15285 .</p><p>Remarks.</p><p>This species was described from Penang Hill (Pfeiffer 1856) but was not recorded by Stoliczka (1872, 1873). It is a widespread species in Malaysia and southern Thailand (Laidlaw 1928; Pall-Gergely et al. 2014; Jirapatrasilp et al. 2022).</p></div>	https://treatment.plazi.org/id/ABBEFC35FCFB52AD88A20913BCADC1C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
D14BDCBA5E4A5825BFE2E56B02D887C6.text	D14BDCBA5E4A5825BFE2E56B02D887C6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Semperula birmanica (Theobald 1864)	<div><p>Semperula birmanica (Theobald, 1864)</p><p>Remark.</p><p>Old record by Stoliczka (1873).</p></div>	https://treatment.plazi.org/id/D14BDCBA5E4A5825BFE2E56B02D887C6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
6A4FBFE0863C5345B177B31BA40C36C7.text	6A4FBFE0863C5345B177B31BA40C36C7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Subulina octona (Bruguiere 1789) BOR	<div><p>Subulina octona (Bruguière, 1789)</p><p>Fig. 7 F</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2263&amp;materialsCitation.latitude=5.4218" title="Search Plazi for locations around (long 100.2263/lat 5.4218)">Lam durian farm, Plot. 19</a>; 5.4218 ° N, 100.2263 ° E; 280 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15422  •   Penang Hill, Trail to Western hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2494&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2494/lat 5.4223)">Junction to Teluk Bahang from Penang Hill, Plot. 22</a>; 5.4223 ° N, 100.2494 ° E; 760 m a. s. l.; 24 Oct. 2022; T. S. Liew, J, Dulipat, M. Y. Lee leg.; BOR/MOL 15545  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15748  .</p><p>Remarks.</p><p>New record from Penang Hill. This species is also recorded from the mainland of Peninsular Malaysia (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/6A4FBFE0863C5345B177B31BA40C36C7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
D721DBF702FB556D8DE7623627CA8C40.text	D721DBF702FB556D8DE7623627CA8C40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanychlamys indica (Godwin Austen 1883) BOR	<div><p>Tanychlamys indica (Godwin Austen, 1883)</p><p>Figs 10 B, 20 E</p><p>Material examined.</p><p>Malaysia • Penang Hill, Along Jalan Tunku Yahya Petra, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2662&amp;materialsCitation.latitude=5.4225" title="Search Plazi for locations around (long 100.2662/lat 5.4225)">Random 6</a>.; 5.4225 ° N, 100.2662 ° E; 730 m a. s. l.; 20 Jun. 2023; T. S. Liew leg.; BOR/MOL 15771  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. This species is widespread in Malaysia and might have been introduced to this location (Maassen 2001).</p></div>	https://treatment.plazi.org/id/D721DBF702FB556D8DE7623627CA8C40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
CA7A7A6D46095FF58FDFA13171375AC4.text	CA7A7A6D46095FF58FDFA13171375AC4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanychlamys stephoides (Stoliczka 1873) BOR	<div><p>Tanychlamys stephoides (Stoliczka, 1873)</p><p>Fig. 11 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2669&amp;materialsCitation.latitude=5.4248" title="Search Plazi for locations around (long 100.2669/lat 5.4248)">Research trail, Plot. 01</a>; 5.4248 ° N, 100.2669 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15164  .</p><p>Remark.</p><p>This species was previously described by Stoliczka (1873) from Penang Hill.</p></div>	https://treatment.plazi.org/id/CA7A7A6D46095FF58FDFA13171375AC4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
5FE692C91D545E0196A53004540242E7.text	5FE692C91D545E0196A53004540242E7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanychlamys tersa (Issel 1874) BOR	<div><p>Tanychlamys tersa (Issel, 1874)</p><p>Figs 11 C, 21 B</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2187&amp;materialsCitation.latitude=5.4148" title="Search Plazi for locations around (long 100.2187/lat 5.4148)">Tropical fruit farm, Plot. 14</a>; 5.4148 ° N, 100.2187 ° E; 260 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15553  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.22&amp;materialsCitation.latitude=5.416" title="Search Plazi for locations around (long 100.22/lat 5.416)">Tropical fruit farm, Plot. 15</a>; 5.4160 ° N, 100.2200 ° E; 250 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15246  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15250  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2866&amp;materialsCitation.latitude=5.44" title="Search Plazi for locations around (long 100.2866/lat 5.44)">Plot. 29</a>; 5.4400 ° N, 100.2866 ° E; 75 m a. s. l.; 2 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15758  .</p><p>Remarks.</p><p>This is a new record for Penang Hill. This species is widespread in Malaysia and might have been introduced to this location (Maassen 2001).</p></div>	https://treatment.plazi.org/id/5FE692C91D545E0196A53004540242E7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
4BA934A2EE63517EBF6386846F7BAB60.text	4BA934A2EE63517EBF6386846F7BAB60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanychlamys undetermined-1	<div><p>Tanychlamys Penang sp. 1</p><p>Figs 10 C, 20 F</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2669&amp;materialsCitation.latitude=5.4248" title="Search Plazi for locations around (long 100.2669/lat 5.4248)">Research trail, Plot. 01</a>; 5.4248 ° N, 100.2669 ° E; 720 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15160  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2645&amp;materialsCitation.latitude=5.4226" title="Search Plazi for locations around (long 100.2645/lat 5.4226)">Research
trail, Plot. 03</a>; 5.4226 ° N, 100.2645 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.;, BOR/MOL 15198  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat
nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15130  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2653&amp;materialsCitation.latitude=5.4228" title="Search Plazi for locations around (long 100.2653/lat 5.4228)">Habitat
nature trail, Plot. 05</a>; 5.4228 ° N, 100.2653 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15132  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2642&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2642/lat 5.421)">Habitat
nature trail, Plot. 07</a>; 5.4210 ° N, 100.2642 ° E; 760 m a. s. l.; 6 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15145  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2648&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2648/lat 5.4223)">Habitat
nature trail, Plot. 08</a>; 5.4223 ° N, 100.2648 ° E; 760 m a. s. l.; 6 Sept. 2022; J. Dulipat, S. M. Goh, M. Y. Lee leg.; BOR/MOL 15098  •  same data as for preceding; BOR/MOL 15154 •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2921&amp;materialsCitation.latitude=5.4341" title="Search Plazi for locations around (long 100.2921/lat 5.4341)">Moon Gate Station 5 trail, Plot. 11</a>; 5.4341 ° N, 100.2921 ° E; 110 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15153  •   Penang Hill, Botanical garden, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.286&amp;materialsCitation.latitude=5.4396" title="Search Plazi for locations around (long 100.286/lat 5.4396)">Plot. 12</a>; 5.4396 ° N, 100.2860 ° E; 85 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15413  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2187&amp;materialsCitation.latitude=5.4148" title="Search Plazi for locations around (long 100.2187/lat 5.4148)">Tropical
fruit farm, Plot. 14</a>; 5.4148 ° N, 100.2187 ° E; 260 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15247  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.22&amp;materialsCitation.latitude=5.416" title="Search Plazi for locations around (long 100.22/lat 5.416)">Tropical
fruit farm, Plot. 15</a>; 5.4160 ° N, 100.2200 ° E; 250 m a. s. l.; 21 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15302  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2637&amp;materialsCitation.latitude=5.3908" title="Search Plazi for locations around (long 100.2637/lat 5.3908)">Forest
near farm, Plot. 21</a>; 5.3908 ° N, 100.2637 ° E; 340 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15248  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2517&amp;materialsCitation.latitude=5.4203" title="Search Plazi for locations around (long 100.2517/lat 5.4203)">Trail to 
Western
hill, Km 3.6, Plot. 23</a>; 5.4203 ° N, 100.2517 ° E; 750 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15549  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest
near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15244  •  same data as for preceding; BOR/MOL 15303 •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2575&amp;materialsCitation.latitude=5.39" title="Search Plazi for locations around (long 100.2575/lat 5.39)">Forest near Dam, Plot. 25</a>; 5.3900 ° N, 100.2575 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15249  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3977" title="Search Plazi for locations around (long 100.2612/lat 5.3977)">Forest
near Dam, Plot. 26</a>; 5.3977 ° N, 100.2612 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15245  •  same data as for preceding; BOR/MOL 15304 •   Penang Hill, Trail from Viaduct to Claremont, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">after shelter, Plot. 27</a>; 5.4205 ° N, 100.2724 ° E; 500 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15305  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15740  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4216" title="Search Plazi for locations around (long 100.2674/lat 5.4216)">Path B, Random 1. PathB</a>; 5.4216 ° N, 100.2674 ° E; 730 m a. s. l.; 4 Sept. 2022; T. S. Liew, J. Dulipat leg. ; •  same data as for preceding; BOR/MOL 15151 .</p><p>Remark.</p><p>This is a new record for Penang Hill. It is one of the most common large land snails found there.</p></div>	https://treatment.plazi.org/id/4BA934A2EE63517EBF6386846F7BAB60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
7978E9F0D9DE5B008FA559C8F0D7346D.text	7978E9F0D9DE5B008FA559C8F0D7346D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tanychlamys undetermined-2	<div><p>Tanychlamys Penang sp. 2</p><p>Figs 11 A, 21 A</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2645&amp;materialsCitation.latitude=5.4226" title="Search Plazi for locations around (long 100.2645/lat 5.4226)">Research trail, Plot. 03</a>; 5.4226 ° N, 100.2645 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15424  •   Penang Hill, Teluk Bahang- Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2265&amp;materialsCitation.latitude=5.4199" title="Search Plazi for locations around (long 100.2265/lat 5.4199)">Bukit Kerajaan ForestReserve next to Lam durian farm, Plot. 20</a>; 5.4199 ° N, 100.2265 ° E; 330 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat, M. Y. Lee leg.; BOR/MOL 15552  •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2637&amp;materialsCitation.latitude=5.3908" title="Search Plazi for locations around (long 100.2637/lat 5.3908)">Forest near farm, Plot. 21</a>; 5.3908 ° N, 100.2637 ° E; 340 m a. s. l.; 23 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15550  •   Penang Hill, Trail to Western hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2494&amp;materialsCitation.latitude=5.4223" title="Search Plazi for locations around (long 100.2494/lat 5.4223)">Junction to Teluk Bahang from Penang Hill, Plot. 22</a>; 5.4223 ° N, 100.2494 ° E; 760 m a. s. l.; 24 Oct. 2022; T. S. Liew, J, Dulipat, M. Y. Lee leg.; BOR/MOL 15551  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.242&amp;materialsCitation.latitude=5.4211" title="Search Plazi for locations around (long 100.242/lat 5.4211)">Western hill to Teluk Bahang Trail Stetion 4, Plot. 30</a>; 5.4211 ° N, 100.2420 ° E; 570 m a. s. l.; 3 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15759  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2334&amp;materialsCitation.latitude=5.4291" title="Search Plazi for locations around (long 100.2334/lat 5.4291)">Western hill to Teluk Bahang Trail Stetion 7, Plot. 31</a>; 5.4291 ° N, 100.2334 ° E; 490 m a. s. l.; 3 Feb. 2023; T. S. Liew, M. Y. Lee, A. Yusni leg.; BOR/MOL 15757  .</p><p>Remark.</p><p>This is a new record for Penang Hill.</p></div>	https://treatment.plazi.org/id/7978E9F0D9DE5B008FA559C8F0D7346D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
5CFE43EA054B5FA19B9C1ACBF373E9D0.text	5CFE43EA054B5FA19B9C1ACBF373E9D0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Trichochloritis penangensis (Stoliczka 1873) BOR	<div><p>Trichochloritis penangensis (Stoliczka, 1873)</p><p>Fig. 11 F</p><p>Material examined.</p><p>Malaysia • Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15291  .</p><p>Remarks.</p><p>This species was previously described by Stoliczka (1873) from Penang Hill. It can also be found in Perak (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/5CFE43EA054B5FA19B9C1ACBF373E9D0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
8AC0A91B8E8C51B986F362F7A68756D6.text	8AC0A91B8E8C51B986F362F7A68756D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Videna cantoriana (W. H. Benson 1861)	<div><p>Videna cantoriana (W. H. Benson, 1861)</p><p>Remarks.</p><p>This species was previously recorded and described from Penang Hill (at ca 2000 feet a. s. l.) by Stoliczka (1873). It was not found in this study.</p></div>	https://treatment.plazi.org/id/8AC0A91B8E8C51B986F362F7A68756D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
FE852B2DB53A5F8BB63AB1CDF1A30D2E.text	FE852B2DB53A5F8BB63AB1CDF1A30D2E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Videna castra (Benson 1852) BOR	<div><p>Videna castra (Benson, 1852)</p><p>Fig. 17 C</p><p>Material examined.</p><p>Malaysia • Penang Hill, Teluk Bahang-Balik Pulau, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2159&amp;materialsCitation.latitude=5.418" title="Search Plazi for locations around (long 100.2159/lat 5.418)">Entrance of a durian farm, near Boulder Valley, Old rubber tree, Plot. 17</a>; 5.418 ° N, 100.2159 ° E; 180 m a. s. l.; 22 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15279  .</p><p>Remarks.</p><p>This species was also previously recorded by Stoliczka (1873). It is similar to  Videna cantoriana but has more regularly expanded whorls. This species is also recorded from mainland Peninsular Malaysia, India, and Thailand (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/FE852B2DB53A5F8BB63AB1CDF1A30D2E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
314D032FDF355160973C50248E09BC9D.text	314D032FDF355160973C50248E09BC9D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Videna timorensis (E. von Martens 1867) BOR	<div><p>Videna timorensis (E. von Martens, 1867)</p><p>Fig. 17 D</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15129  .</p><p>Remarks.</p><p>This species was also previously recorded by Stoliczka (1873). It differs from the other two  Videna species of Penang Hill by having a wider, more open umbilicus, approximately one-third of the shell width.</p></div>	https://treatment.plazi.org/id/314D032FDF355160973C50248E09BC9D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
B9D21DBE8B5E56A688F1BA90AD0886CF.text	B9D21DBE8B5E56A688F1BA90AD0886CF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Vitrinopsis nucleata (Stoliczka 1873) BOR	<div><p>Vitrinopsis nucleata (Stoliczka, 1873)</p><p>Figs 16 B, 21 F</p><p>Material examined.</p><p>Malaysia • Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2668&amp;materialsCitation.latitude=5.4231" title="Search Plazi for locations around (long 100.2668/lat 5.4231)">Habitat nature trail, Plot. 04</a>; 5.4231 ° N, 100.2668 ° E; 770 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15115  •   Penang Hill, The Habitat, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2674&amp;materialsCitation.latitude=5.4247" title="Search Plazi for locations around (long 100.2674/lat 5.4247)">Research trail, Plot. 06</a>; 5.4247 ° N, 100.2674 ° E; 750 m a. s. l.; 5 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh, M. Y. Lee, M. Y. Wua leg.; BOR/MOL 15479  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2921&amp;materialsCitation.latitude=5.4341" title="Search Plazi for locations around (long 100.2921/lat 5.4341)">Moon Gate Station 5 trail, Plot. 11</a>; 5.4341 ° N, 100.2921 ° E; 110 m a. s. l.; 7 Sept. 2022; T. S. Liew, J. Dulipat, S. M. Goh leg.; BOR/MOL 15122  •  same data as for preceding; BOR/MOL 15477 •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3904" title="Search Plazi for locations around (long 100.2612/lat 5.3904)">Forest near Dam, Plot. 24</a>; 5.3904 ° N, 100.2612 ° E; 380 m a. s. l.; 24 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15242  •  same data as for preceding; BOR/MOL 15292 •   Penang Hill, Air Itam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2612&amp;materialsCitation.latitude=5.3977" title="Search Plazi for locations around (long 100.2612/lat 5.3977)">Forest near Dam, Plot. 26</a>; 5.3977 ° N, 100.2612 ° E; 290 m a. s. l.; 25 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15478  •   Penang Hill, Trail from Viaduct to Claremont, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2724&amp;materialsCitation.latitude=5.4205" title="Search Plazi for locations around (long 100.2724/lat 5.4205)">after shelter, Plot. 27</a>; 5.4205 ° N, 100.2724 ° E; 500 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15238  •   Penang Hill, Trail - Moniot Road East, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2746&amp;materialsCitation.latitude=5.424" title="Search Plazi for locations around (long 100.2746/lat 5.424)">Plot. 28</a>; 5.4240 ° N, 100.2746 ° E; 530 m a. s. l.; 26 Oct. 2022; T. S. Liew, J. Dulipat leg.; BOR/MOL 15233  •   Penang Hill, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.2701&amp;materialsCitation.latitude=5.421" title="Search Plazi for locations around (long 100.2701/lat 5.421)">By Path H, Plot. 33</a>; 5.4210 ° N, 100.2701 ° E; 610 m a. s. l.; 4 Feb. 2023; T. S. Liew, A. Yusni leg.; BOR/MOL 15743  .</p><p>Remark.</p><p>This species was described and recorded by Stoliczka (1873) from Penang Hill, and it can also be found in Perak (Maassen 2001; Foon et al. 2017).</p></div>	https://treatment.plazi.org/id/B9D21DBE8B5E56A688F1BA90AD0886CF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Goh, Soo-Mun;Jasrul, Dulipat;Lee, Mei-Yi;Vermeulen, Jaap J.;Liew, Thor-Seng	Goh, Soo-Mun, Jasrul, Dulipat, Lee, Mei-Yi, Vermeulen, Jaap J., Liew, Thor-Seng (2025): From 153 - year-old records to contemporary discoveries: land snail (Mollusca, Gastropoda) diversity in Penang Hill, Malaysia. ZooKeys 1230: 99-154, DOI: 10.3897/zookeys.1230.136906
