taxonID	type	description	language	source
9E60878A9B58FF9A74BEFC1DD25244A7.taxon	description	New species of Scottmoria, the size and shape of leaf blades, short inflorescences on terminal leafy branches and size of flowers resemble those of S. integrifolia (Ruiz & Pav. ex Miers) Cornejo in Vargas et al. (2024: 174), however, S. austroccidentalis differs from the latter species by the verruculose-papillose petioles (vs. smooth petioles) and calyx lobes imbricate along the lower half, with ciliolate margins (vs. calyx lobes valvate, with glabrous margins).	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
9E60878A9B58FF9A74BEFC1DD25244A7.taxon	materials_examined	Type: — ECUADOR. El Oro: El Guabo, El Retiro, bosque nublado, pastizales abandonados, 3 ° 12 ’ S 79 ° 49 ’ W, 1755 m, 6 Jul 2017 (fl), E. Freire 12222 (holotype: QCNE- 0241389!). Tree to 20 m tall and ca. 20 cm DBH, the bark and trunk unknown. Stem glabrous, lenticellate, the lenticels orbicular to elliptic. Leaves petiolate, the petioles channeled adaxially, rounded abaxially, hemispherical in cross section, 5 – 10 × 1 – 2 mm, verruculose-papillose, glabrous; blades narrowly elliptic to oblong-elliptic, 7 – 15 × 2.5 – 6 cm, chartaceous (dry), with abundant black punctations abaxially, the base cuneate to broadly obtuse, revolute, the margins entire, the apex shortly acuminate; venation eucamptodromous towards base and brochidodromous towards apex, the midrib verruculose-papillose, and sulcate to impressed at base, costate to prominent for most of length adaxially, salient abaxially, glabrous, the secondary veins in 8 – 12 pairs, sulcate to impressed and prominulous adaxially, salient abaxially, the tertiary veins densely reticulate. Inflorescences axillary on leafy branches, a raceme, the rachis 1 – 10 cm × 1 – 2 mm at lower half, straight to slightly flexuose, glabrous, lenticellate; pedicels ca. 5 × 0.7 – 1.5 mm, lenticellate, glabrous, blackish (dry); floral bracts 2, triangular to deltoid, 1 – 2 mm long, deciduous; flower buds purple (fresh). Flowers ca. 4 cm in diam. (fresh), 2.5 – 3 cm in diam. (dry); hypanthium minutely verruculose, glabrous, truncate at summit, tapering at base, 5 – 6 mm long to articulation; calyx with 6 lobes, the lobes broadly ovate to deltoid, 1.5 – 3 × 1.5 – 3 mm (dry), divergent to patent at anthesis, smooth abaxially, the margins ciliolate, the bases imbricate, arising from fused calyx rim; petals 6, elliptic to obovate, 1 – 1.5 × 0.6 – 1 cm, pink or purplish red, the lower half not costate; androecial hood with three coils, 1 – 1.5 cm across, pink or purplish red (fresh), inner cleft present; vestigial stamens on outside of coils; staminal ring with ca. 110 stamens, the filaments ca. 1 mm long (fresh), clavate, the anthers ca. 0.5 mm long (fresh); the ovary summit slightly raised over hypanthium, surrounded by a plane intrastaminal ring scar, shallow depression between ring scar and summit of ovary present; the style ca. 1.5 × 1.5 mm, stout, distinguished from ovary summit, a stylar collar absent, the stigma pore transversely linear-oblong. Fruits and seeds not seen.	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
9E60878A9B58FF9A74BEFC1DD25244A7.taxon	discussion	Discussion: — Scottmoria austroccidentalis resembles S. integrifolia, a species that occurs from western Ecuador to southwestern Colombia, but the new species differs from the latter by the characters discussed above in the diagnosis. The calyx lobes, which are imbricate in the lower half, and with ciliolate margins, are quite distinct from those of congenerics on the Pacific coast of northwestern South America.	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
9E60878A9B58FF9A74BEFC1DD25244A7.taxon	etymology	Etymology: — The epithet austroccidentalis refers to the southern range of this new species within western Ecuador. Common names: — Not recorded. Habitat and distribution: — A medium-sized tree, persistent as a solitary individual surrounded by pastures or in small fragmented patches of secondary wet forest, in well-drained soils. Known only from the type locality and in the Buenaventura ecological reserve, both located in the province of El Oro, 1300 – 1755 m elevation, in evergreen southwestern Andean montane forests.	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
9E60878A9B58FF9A74BEFC1DD25244A7.taxon	description	Phenology: — Flowers have been observed in May and July. No flower visitors have been observed in the field; however, it is expected that the flowers of this and those of the other new species of Scottmoria in this article are visited and pollinated by native bees. Conservation status: — Based on georeferenced herbarium collections, the occurrence of Scottmoria austroccidentalis in two localities allows us to assign here the provisional category of Critically Endangered (CR B 2 a, b) following the IUCN (2022) criteria. Paratypes: — ECUADOR. El Oro: Piñas, buffer zone / border región of Reserva Ecológica Buenaventura; 11 km (air km) north of the Entrada ¨ La Virgin ¨ or road towards Viron 3 ° 33 ’ 28 ” S 79 ° 46 ’ 12 ” W, 1300 – 1700 m, 13 May 2003 (fl), J. L. Clark, M. Mailloux & S. Seger 7984 (NY!, QCNE!, US!).	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
9E60878A9B5DFF9974BEFC8BD48C44A7.taxon	description	New species of Scottmoria, the size, shape of leaf blades and number of lateral veins resemble those of S. antioquiensis (Dugand & Daniel) Cornejo in Vargas et al. (2024: 172), however, S. petalocostata differs from the latter species by the triangular, valvate calyx lobes (vs. broadly ovate to hemiorbicular, imbricate in the lower half) and the four outer petals with a prominent costa along the lower half and slightly pustulate abaxially (vs. petals not costate and smooth abaxially).	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
9E60878A9B5DFF9974BEFC8BD48C44A7.taxon	materials_examined	Type: — ECUADOR. Carchi: San Marcos de los Coaiqueres and surrounding perhumid forest on trail Chical-Tobar Donoso, 1 ° 06 ’ N 78 ° 16 ’ W, 800 m, 8 Feb 1985 (fl), B. Øllgaard, J. Korning, K. Thomsen & T. Illum 57654 (holotype: NY- 728178!; isotypes: AAU!, QCA- 54434!). Tree to 30 m tall and 30 cm DBH, the bark and trunk unknown. Stems glabrous, lenticellate, the lenticels circular to elliptic. Leaves petiolate, the petioles shallowly channeled adaxially, rounded abaxially, hemispherical in cross section, 12 – 20 × 2 – 3 mm, lenticellate, glabrous; blades elliptic, 10 – 15 × 5.5 – 8 cm, chartaceous (dry), with abundant black punctations abaxially, the base broadly obtuse to rounded, revolute, the margins entire, the apex rounded to shortly acuminate; venation eucamptodromous towards base and brochidodromous towards apex, the midrib rather smooth, sulcate adaxially, prominent, more or less finely veined and with abundant dark short streaks abaxially, glabrous, the secondary veins in 7 – 12 pairs, impressed to prominulous adaxially, salient abaxially, the tertiary veins reticulate. Inflorescences axillary and terminal, a panicle on leafy branches, the rachis 5.5 – 14 cm × 4 – 8 mm at lower half, straight to slightly flexuose, lenticellate, glabrous; pedicels 5 – 8 × 3 – 5 mm, richly verruculose-lenticellate, glabrous, blackish (dry); floral bracts absent; flower buds dark purple (fresh). Flowers ca. 3 cm in diam. (fresh), ca. 2.5 cm in diam. (dry); hypanthium glabrous, densely pustulate or verruculose-lenticellate, truncate at summit, abruptly contracted onto pedicel at base, 2 – 3 mm long, inconspicuously articulate; calyx with 6 lobes, the lobes triangular, 3 – 4 × 2 – 3 mm (fresh), broadly divergent to patent at anthesis, pustulate or verruculose-lenticellate (fresh) abaxially, the margins entire, the bases valvate, arising from fused calyx rim; petals 6, broadly obovate to ± orbicular or broadly ovate, 1.5 – 2.3 × 1.3 – 2 cm, dark purple, the 4 outer petals somewhat pustulate abaxially and with a prominent costa along the lower half; androecial hood with three coils, ca. 1.3 cm across, yellow (fresh), inner cleft present; vestigial stamens on outside of coils; staminal ring with ca. 120 stamens, the filaments 0.5 – 1.5 mm long (fresh), clavate, the anthers ca. 0.5 mm long (fresh); the ovary summit slightly raised over hypanthium, surrounded by a prominent intrastaminal ring scar, ca. 0.3 mm high, with a shallow circular depression between intrastaminal ring scar and summit of ovary; the style arising ca. 2 × 1.5 mm, stout, distinguished from ovary summit, a stylar collar absent, the stigma pore transversely linear-oblong. Fruits and seeds not known.	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
9E60878A9B5DFF9974BEFC8BD48C44A7.taxon	discussion	Discussion: — Scottmoria petalocostata resembles S. antioquiensis that is restricted to the departments of Antioquia, Chocó and Cundinamarca, in Colombia, and the state of Mérida in NW Venezuela (Mori and Prance 1990, Mori et al. 2015, Vargas et al. 2024). For differences, see the diagnosis. Furthermore, the other species in the genus are not known to have abaxially costate four outer petals.	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
9E60878A9B5DFF9974BEFC8BD48C44A7.taxon	etymology	Etymology: — The epithet petalocostata refers to the outer petals with a conspicuous prominent costa. Common names: — Not recorded. Habitat and distribution: — A canopy tree of mature wet forests, in well drained soils. Known from northwestern Ecuador at 800 m elevation. This new species may also occur in the department of Nariño, southwestern Colombia.	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
9E60878A9B5DFF9974BEFC8BD48C44A7.taxon	description	Phenology: — Flowers have been collected in February. Conservation status: — Scottmoria petalocostata is only known from the type collection gathered in 1985 from a single locality, it is assigned here the provisional category of Critically Endangered (CR B 2 a, b) following IUCN (2022) criteria. After a full revision of the genus Scottmoria we have found that the following species placed in Eschweilera exhibit the morphological characters of Scottmoria and therefore need to be transferred to that genus. The defining characters are the androecial hood with three or four inner coils as seen in medial longitudinal section, a short, stout, cylindric style, and two kinds of seed covering: 1) a thin white sarcotesta over the entire or most of the mature seed that is infiltrated within testa or 2) a thin white sarcotesta present over the entire or most of the young seeds, this mostly dissolving at maturity and leaving sarcotesta remains mainly near to the funicular lateral arils, that are thick, yellow to white, mostly half I-beam shaped, with rubber-like texture and with overlapping ends on the seed; seeds with funicle forked at insertion point from apex of seed, the funicle apical branches soon anastomosing into an elongate filiform vascular bundle as long as the whole seed, that is longitudinally arranged on top of seed lateral crest, and the funicle fully surrounded by the lateral aril, inserted at the inner base of the fruit (Vargas et al. 2024, Cornejo & Prance 2024).	en	Cornejo, Xavier, Prance, Ghillean T. (2025): Two new species of Scottmoria (Lecythidaceae) from western Ecuador and six new combinations in the genus. Phytotaxa 696 (3): 253-261, DOI: 10.11646/phytotaxa.696.3.5, URL: https://doi.org/10.11646/phytotaxa.696.3.5
