taxonID	type	description	language	source
9F6A8E412308FFD9FEBB5D3BAAF2F8FC.taxon	description	Pupa (Gredleriella) von Möllendorff, 1884: 179, 180. Type species: Pupa hunana Gredler, 1881, by original designation [‘ hunanensis ’ is an incorrect subsequent spelling by von Möllendorff (1884)].	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412308FFD9FEBB5D3BAAF2F8FC.taxon	type_taxon	Type species. Pupa dorsata Ancey, 1881, by subsequent designation in Gude, 1914: 239.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412308FFD9FEBB5D3BAAF2F8FC.taxon	diagnosis	Diagnosis. Shell conic to elongate conic, brownish to chestnut-brown, 5 to 7 convex whorls. Shell surface with dense and irregular growth lines. Aperture vertical to subvertical; peristome broadly expanded; lip adnate to preceding whorl or little elevated with very short tuba. Apertural dentitions generally four or five (type species; Fig. 1 A, B) or more: one on parietal wall (in the case of angular concrescent with parietal lamella), upper and lower palatal plicae, columellar lamella, and additional dentitions (basal, palatal tubercle or infraparietal) may be present. Umbilicus narrow or perforate.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412308FFD9FEBB5D3BAAF2F8FC.taxon	discussion	Remarks. Originally, Boysidia was described as a subgenus of Pupa Röding, 1798. Subsequently, Pilsbry and Vanetta (1900: 592) transferred Boysidia to a subgenus of Hypselostoma Benson, 1856. Not long after, Gude (1914) raised Boysidia to a full genus that comprised hypselostomatid species without tuba, without spiral striation, and with a large independent or concrescent parietal lamella. This definition has been widely used and followed since then. Currently, Boysidia contains some 35 nominal species that are broadly distributed (Páll-Gergely & White, 2022; MolluscaBase, 2024). There are 19 species from China, six from Indochina, and eight from Malaysia, including Borneo (Zhang et al., 2014; MolluscaBase, 2024). In addition, there are two species with disjunct distributions: one from Timor and another from Pakistan (Rensch, 1935; Pokryszko et al., 2009). Among these, only two species, Boysidia pahpetensis Saurin, 1953 and Boysidia houaphanica, new species, are now known from Laos. When examining the specimens from China, von Möllendorff (1884) described Pupa (Gredleriella) with Pupa hunana Gredler, 1881 as the type species. Later, in a taxonomic revision of Boysidia, Pilsbry (1917: 194) recognised three subgenera: the nominotypical subgenus Boysidia, Bensonella Pilsbry & Vanatta, 1900 and Paraboysidia Pilsbry, 1917. However, the type species of Gredleriella (Pupa hunana Gredler, 1881; Fig. 1 C) is very similar to the type of Boysidia sensu stricto (Pupa dorsata Ancey, 1881; Fig. 1 A, B) in general shell form and the apertural dentitions. Therefore Pilsbry (1917: 194) synonymised subgenus Gredleriella with the nominotypical subgenus. This subgeneric classification was widely used (e. g., Schileyko, 1998) until Páll-Gergely and White (2022) formally synonymised Paraboysidia with Bensonella and recognised Bensonella as a distinct genus based on apertural dentition. The results of this generic revision by Páll-Gergely and White (2022) are followed by Tongkerd et al. (2024) and Jirapatrasilp et al. (2024). Dasypupa was nominated as a subgenus of Boysidia with the distinguishing characters of a depressed-conical shell without spiral ridges, a concresent lamella with a small and bilobed blade, and having a small columellar lamella that is situated deep inside the aperture (Thompson & Dance, 1983). Based only on similarity in shell morphology and apertural dentition, Pokryszko et al. (2009) applied this subgeneric name to a new Boysidia species from Pakistan. Nevertheless, the published re-description of the type species of Dasypupa, Boysidia salpinx Thompson & Dance, 1983 (Fig. 1 D), in the handbook of land snails and slugs of Sabah and Labuan (Vermeulen & Liew, 2022: 338, 339) does not recognise this subgenus, and rejects applying this subgenus without apparent reason. The current status of the subgenus Dasypupa is uncertain and its position and relationship with Boysidia need further study.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412309FFDCFEB45C3DABB5FC5A.taxon	description	(Figs 2 B, 3)	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412309FFDCFEB45C3DABB5FC5A.taxon	materials_examined	Material examined. Holotype CUMZ 14420 (height 3.2 mm, width 2.8 mm, 6 whorls; Fig. 3 A – C) from limestone outcrop at Ban Lao, Samtay District, Houaphanh Province, Laos (20 ° 00 ′ 52.4 ″ N, 104 ° 40 ′ 57.0 ″ E), coll. K. Inkhavilay, A. Inthavong and C. Oulaiseng, May 2022. Paratypes CUMZ 14421 (6 shells; Figs 2 B, 3 D), CUMZ 14422 (3 shells), and NHMUK (2 shells) were all collected from the type locality together with the holotype.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412309FFDCFEB45C3DABB5FC5A.taxon	diagnosis	Diagnosis. Shell minute, heliciform, without tuba. Protoconch sculpture with weakly malleated pits; teleoconch with irregular growth lines. Apertural dentition consisting of five lamellae (parietal, upper and lower palatals, basal and columellar), and small palatal tubercle.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412309FFDCFEB45C3DABB5FC5A.taxon	description	Description. Shell elongate conical, yellowish brown to chestnut-brown, with 5 ½ to 6 ¾ convex whorls. Shell height 3.1 – 3.5 mm, shell width 2.5 – 2.8 mm. Apex rather round; protoconch about 1 ½ whorls with weak malleated pits. Teleoconch generally smooth with scattered, prominent and irregular growth lines; suture wide, deeply impressed. Last whorl large, rounded, slightly descending near aperture. Peristome attached to preceding whorl; inner surface including plicae and lamellae covered with scattered small nodules; lip slightly thickened, whitish, broadly expanded. Aperture sub-rectangular, with six apertural teeth. Parietal lamella (p) strongly developed, elevated, sinuous deeply inside aperture, consisting of two parts separated at concrescent point: outer part elevated, strong, curved, reaching peristome margin; inner part thickened, elevated, stronger than outer part. Palatal tubercle (pt) strongly developed with triangular shape. Upper palatal plica (upl) thin, flake shaped, long inside aperture, elevated to nearly touch parietal lamella, forming an almost separate chamber at upper junction of peristome; lower (lpl) palatal strong, elevated. Basal plica (b) prominent, small and low. Columellar lamella (c) large, strong, distinct horizontally. Umbilicus perforate, about one-fourth of shell width, rounded, shallow, not revealing preceding whorl.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412309FFDCFEB45C3DABB5FC5A.taxon	diagnosis	Differential diagnosis. Boysidia pahpetensis (Fig. 2 A) from Laos differs from the new species in having a conic shell, four whorls, rounded aperture, small infraparietal lamella, and a concrescent point situated close to the peristome margin (Ancey, 1881; Inkhavilay et al., 2019). In contrast, the new species possesses an elongate conical shape, five to six whorls, a sub-rectangular aperture, without infraparietal lamella, and a concrescent point situated deeper inside the aperture. Boysidia robusta Bavay & Dautzenberg, 1912 from Vietnam and Boysidia chiangmaiensis Panha & Burch, 2002 from Thailand differ from the new species in having ovate conic and large shells (height 4 – 5 mm) with basal plica. In addition, Boysidia robusta also has an ovate aperture, and with small infraparietal lamella and lower basal plica, while Boysidia chiangmaiensis has a rounded aperture (Bavay & Dautzenberg, 1912; Panha & Burch, 2002 a, 2005). In comparison, Boysidia houaphanica, new species, possesses an elongate conical, smaller shell (height 3.1 – 3.5 mm), subrectangular aperture, and with basal plica. For further comparison, Boysidia tholos Panha & Burch, 2002 b from Thailand can be distinguished from this new species by having a dome-shaped shell, five whorls, rounded aperture, and infrapalatal lamella. In contrast, Boysidia houaphanica, new species has an elongated conic shell, 5 – 6 whorls, subrectangular aperture, and without infrapalatal lamella (Panha & Burch, 2002 b, 2005).	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412309FFDCFEB45C3DABB5FC5A.taxon	etymology	Etymology. The species name ‘ houaphanica ’ is derived from the type locality, Houaphanh Province, where the type specimens were collected.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412309FFDCFEB45C3DABB5FC5A.taxon	distribution	Distribution. This species, Boysidia houaphanica, new species, is known only from the type locality in Houaphanh Province, Laos. The snails were found under leaf litter in limestone rock crevices.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412309FFDCFEB45C3DABB5FC5A.taxon	discussion	Remarks. This new species occurs sympatrically with Bensonella novemdentata. The occurrence of multiple hypselostomatid species in the same locality is not uncommon (e. g., Inkhavilay et al., 2016; Vermeulen et al., 2019; Páll-Gergely & White, 2022; Sutcharit et al., 2023; Jirapatrasilp et al., 2024).	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E41230CFFDCFF0F58DBA926F8F3.taxon	type_taxon	Type species. Pupa plicidens Benson, 1849, by original designation.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E41230CFFDCFF0F58DBA926F8F3.taxon	diagnosis	Diagnosis. Shell conic to ovate conic, brownish to chestnut-brown and corneous. Shell sculptured with growth lines and very fine spiral striation. Aperture vertical; peristome expanded; lip generally adnate to preceding whorl. Apertural dentition normal (not hooked) or hooked, and generally with many plicae and lamellae; parietal and angular lamellae separated; additional dentitions (basal, columellar, infraparietal, infrapalatal, interpalatal, lower palatal, palatal tubercle, suprapalatal, supracolumellar, subcolumellar, or upper palatal) usually present. Umbilicus narrow or perforate.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E41230CFFDCFF0F58DBA926F8F3.taxon	discussion	Remarks. Currently, the genus Bensonella consists of about thirty species, which are widely distributed, ranging from southern Himalaya to Indochina, China, and Japan (Páll-Gergely & White, 2022; MolluscaBase, 2024).	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E41230CFFD2FF045C05AAB4FA7A.taxon	description	(Fig. 4)	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E41230CFFD2FF045C05AAB4FA7A.taxon	materials_examined	Material examined. Syntype MNHN-IM- 2000 - 33881 (1 shell; Fig. 4 A) from Pah Hia. CUMZ 14426 (1 shell; Fig. 4 B) from limestone outcrop at Ban Lao, Samtay District, Houaphanh Province, Laos (20 ° 00 ′ 52.4 ″ N, 104 ° 40 ′ 57.0 ″ E), coll. K. Inkhavilay, A. Inthavong, and C. Oulaiseng, May 2022.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E41230CFFD2FF045C05AAB4FA7A.taxon	diagnosis	Diagnosis. Shell elongate-conic, yellowish brown in colour. Cervical crest situated far from peristome. Aperture with eight teeth (i. e., parietal, infraparietal, angular, columellar, basal, and upper, inter and lower palatals), not hooked.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E41230CFFD2FF045C05AAB4FA7A.taxon	description	Description. Shell conical, yellowish brown to reddish brown with 5 ¼ to 6 widely convex whorls. Apex rather blunt; protoconch about 1 ½ whorls with wrinkles, very weak spiral ridges. Teleoconch with strong, irregular growth lines and conspicuous; suture impressed and deep. Last whorl large, rounded. Peristome thickened, slightly expanded; constriction weak; lip reddish brown. Cervical crest rather sharp, situated close to peristome. Aperture subquadrate with many apertural dentitions. Parietal lamella (p) strongly developed, rectangular. Infraparietal lamella (ip) small and blunt. Angular lamella (a) slightly higher than parietal lamella, sinuous: outer part short, low, reaching peristome; inner part strongly elevated. Palatal tubercle (pt) strongly developed with triangular shape; one upper palatal plica (upl) with strong folding; one inter- (itpl) and one lower (lpl) palatal relatively large, nearly equal in size, strong plica. Basal plica (b) small, low, tubercle-like. Columellar lamella (c) large, strong, distinct horizontally. Umbilicus narrowly perforate, about one-fifth of shell width, rounded, deep.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E41230CFFD2FF045C05AAB4FA7A.taxon	distribution	Distribution. This species was previously known only from the type locality (Saurin, 1953; Inkhavilay et al., 2019) and is newly recorded from Houaphanh Province in northern Laos. The snails live on limestone karsts and can be found on the limestone wall together with Boysidia houaphanica, new species.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E41230CFFD2FF045C05AAB4FA7A.taxon	discussion	Remarks. The recently collected specimens are from Houaphanh Province, which is about 200 km east of the “ type locality, which is stated to be ‘ Pahai’ ”. However, the morphology of our specimens deviates slightly from that of the syntype (Fig. 4 A) in having a more rounded whorl and deeper suture (Fig. 4 B). Otherwise, the number and arrangement of the apertural dentitions are practically identical with the syntype. Among the morphologically similar species recorded from Laos, Bensonella novemdentata differs from Bensonella paviei (Bavay & Dautzenberg, 1912) and Bensonella anguloobtusa (Inkhavilay & Panha, 2016 in Inkhavilay et al., 2016) by having a prominent cervical crest and palatal plicae forming a short tubercle and situated near the aperture. In comparison, the latter two species have very weak to no cervical crest and palatal plicae extending far inside the aperture; Bensonella anguloobtusa further possesses a subcolumellar plica without lower parietal lamella. In addition, Bensonella novemdentata can be differentiated from Bensonella paralella (Inkhavilay & Panha, 2016) by having a subrectangular aperture and prominent interpalatal and cervical crest, while Bensonella paralella possesses an ovate-shaped aperture, and without interpalatal plica and cervical crest (Inkhavilay et al., 2016; Páll-Gergely & White, 2022; Tongkerd et al., 2024). The two species, Bensonella wangviangensis (Panha & Tongkerd, 2003) and Bensonella gittenbergeri (Maassen, 2008), previously known from Laos, clearly differ from this species in both shell shape and apertural dentition. For comparison, Bensonella wangviangensis has a small shell (height ≤ 2 mm) with 4 whorls, without basal plica, the parietal wall at the posterior corner elevated, and an aperture elliptical with prominent posterior corner balloonshaped, formed by angular lamella and upper palatal plica. Meanwhile, Bensonella gittenbergeri has a large shell (height 4 – 6 mm), ovate conic shape, inconspicuous palatal tubercle, and lower palatal long, reaching expanded lip (Panha et al., 2003; Maassen, 2008; Inkhavilay et al., 2016). Regardless of their phylogenetic relationships, the traditional diagnostic characters set by their respective type species, Boysidia and Bensonella, share a peristome being adnate to the last whorl and lacking the free portion of the last whorl (tuba). In contrast, Boysidia has one parietal lamella (without angular lamella), while Bensonella has separate parietal and angular lamellae. Based on that criterion of generic classification, the species ‘ Boysidia novemdentata ’ is placed under the Boysidia (Saurin, 1953; Inkhavilay et al., 2019; MolluscaBase, 2024). However, the syntype (Fig. 4 A) has well-separated parietal and angular lamellae and peristome adnate to the preceding whorl; these are the unique characteristics of Bensonella (Pilsbry & Vanatta, 1900; Pilsbry, 1917; Schileyko, 1998; Páll-Gergely & White, 2022). Therefore, we have transferred this species into a more appropriate genus, Bensonella.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412302FFD2FC1A5BDAAFB3F8B1.taxon	description	(Fig. 5)	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412302FFD2FC1A5BDAAFB3F8B1.taxon	materials_examined	Material examined. Holotype MNHN-IM- 2000 - 31746 (Fig. 5 A) from Pah Hia. CUMZ 14425 (9 shells; Fig. 5 B, C) from Tham Piew Cave, Muang Kham District, Xiangkhouang Province, Laos (19 ° 40 ′ 24.8 ″ N, 103 ° 34 ′ 05.7 ″ E) coll. K. Inkhavilay and C. Sutcharit, August 2015.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412302FFD2FC1A5BDAAFB3F8B1.taxon	description	Description. Shell small, conical, slightly higher than width, reddish brown, 5 – 5 ½ convex whorls. Apex round; protoconch about 1 – 1 ½ whorl with weak wrinkles and spiral ridgelets. Teleoconch with strong, irregular growth lines, and sometimes growth lines conspicuously white. Suture impressed, deep. Last whorl large, rounded to weak angular (stronger on earlier whorls). Aperture wide, rounded, without any dentition, reddish brown inside. Peristome pale yellowish, little thickened, weakly expanded. Columellar whitish, straight, dilated, overhanging umbilicus. Umbilicus narrowly perforate, about one-fifth of shell width, rounded, deep.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412302FFD2FC1A5BDAAFB3F8B1.taxon	distribution	Distribution. This species was previously known only from the type locality (Saurin, 1953; Páll-Gergely et al., 2015; Inkhavilay et al., 2019), and a recent record is from Xiangkhouang Province.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412302FFD2FC1A5BDAAFB3F8B1.taxon	discussion	Remarks. Although the syntype is small (width about 2 mm), with around four whorls, and a simple and sharp peristomal lip, it probably represents an early or immature shell stage. We considered our newly collected specimens to be this species, although with slightly weaker angular whorl (on spire) and irregular shell surface. In addition, the new collection locality in Xiangkhouang Province is geographically proximal, roughly 90 km northeast of the type locality, ‘ Pahai’ [roughly in present-day Xaisomboun Province]. This species is very similar to K. clerxi Maassen, 2008 from Luang Namtha Province, about 350 km northwest of the type locality, in most shell characters. However, K. laosensis has a slightly ascending last whorl near the peristome, while K. clerxi has a slightly descending last whorl near the peristome (Maassen, 2008; Páll-Gergely et al., 2015; Inkhavilay et al., 2019). Nevertheless, we regard the two species as distinct until more detailed studies to clarify their systematic relationships become available.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412302FFD2FF615EBBA933F7E0.taxon	type_taxon	Type species. Krobylos pomjuk Panha & Burch, 2002, by original designation.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412302FFD2FF615EBBA933F7E0.taxon	diagnosis	Diagnosis. Shell depressed to elevated conic, brownish to light brown, whorl more or less angular, and suture deep. Shell surface with dense and irregular growth lines, and very weak spiral striae may be present. Aperture subvertical; peristome simple (not expanded) to weakly expanded; apertural dentition absent. Umbilicus narrow and rimate.	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
9F6A8E412302FFD2FF615EBBA933F7E0.taxon	discussion	Remarks. Currently, Krobylos consists of nine species, six of which are known exclusively in Thailand, two from Laos, and only one from southern China (Panha & Burch, 2005; Páll-Gergely et al., 2015; MolluscaBase, 2024).	en	Inkhavilay, Khamla, Sutcharit,, Chirasak (2024): A new pulmonate microsnail species in the genus Boysidia, with remarks on Bensonella and Krobylos species (Pupilloidea: Hypselostomatidae) from northern Laos. Raffles Bulletin of Zoology 72: 438-449, DOI: 10.26107/RBZ-2024-0032
