identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
F1763122DBFE5093903B67A818528ED9.text	F1763122DBFE5093903B67A818528ED9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Iotacyphidae Shcherbakov, Bashkuev, & Shmakov 2024	<div><p>3.3. Family Iotacyphidae Shcherbakov, Bashkuev, &amp; Shmakov, 2024 stat. nov.</p><p>Iotacyphinae Shcherbakov, Bashkuev, &amp; Shmakov, 2024: 416. Type genus: Iotacypha Shcherbakov, Bashkuev, &amp; Shmakov, 2024.</p><p>Diagnosis.</p><p>These insects are well characterized by the synapomorphic presence in the forewing of large median cells and strong angles in the courses of R and M, all absent in the Lophioneuridae (sensu Nel et al. 2012, 2014) and Moundthripidae Nel, Azar, &amp; Nel, 2007 . These three families have a CuA re-emerging first from R + M + CuA but CuA is not orthogonal to R + M in the Moundthripidae versus orthogonal in Iotacyphidae and Lophioneuridae (see below).</p><p>Included genera.</p><p>Burmacypha Zherikhin, 2000 (mid-Cretaceous Kachin amber); Iotacypha Shcherbakov, Bashkuev, &amp; Shmakov, 2024 (mid-Cretaceous Kachin amber); Retiptera Cumming, Engel, Lian, &amp; Ulitzka, 2024 (mid-Cretaceous Kachin amber); Triassocypha gen. nov. (Middle Triassic, Monte San Giorgio, Switzerland).</p></div>	https://treatment.plazi.org/id/F1763122DBFE5093903B67A818528ED9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Boderau, Mathieu;Montagna, Matteo;Magnani, Fabio;Magoga, Giulia;Shmakov, Alexey;Engel, Michael S.;Nel, Patricia;Nel, André	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
22E6595FCC9E5AF98C6C59F6598F1A19.text	22E6595FCC9E5AF98C6C59F6598F1A19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lophioneuroptera Engel, Montagna, Boderau, P. Nel, & A. Nel	<div><p>3.2. Clade Lophioneuroptera Engel, Montagna, Boderau, P. Nel, &amp; A. Nel nov.</p><p>(= “ Lophioneurida Tillyard, 1921 ” sensu auctorum)</p><p>Remarks.</p><p>We reject the name “ Lophioneurida ” as applied to this clade on the grounds that it and other names like it above the family group, represent an attempt to impose a rogue nomenclatural system by way of changes to the Principle of Coordination relative to the ICZN family-group series. In brief, there is no such thing as “ Lophioneurida Tillyard, 1921 ” as names above the family group are not coordinate with the family-group series of names. Tillyard (1921) established a family-group name, Lophioneuridae, based on the type genus Lophioneura Tillyard, 1921 and therefore his name is coordinate throughout the ICZN-regulated family group (ICZN 1999), but not further through names outside of the family group. Efforts to either typify and / or make names above the level of the family group coordinate with those of families (e. g., Rohdendorf 1977; Rasnitsyn 1982) have never been adopted nor sanctioned by the ICZN (1999). Accordingly, such efforts to make them part of the natural coordination and typification of family-group names are operating outside of the bounds of the ICZN (1999). We therefore reject the use of “ Lophioneurida ” and other rogue attempts to circumvent the ICZN and force typified coordination of family-group names with those clearly outside of the ICZN-regulated system of names. The same problem applies to the family-group name Thripidae Fallén, 1814 as extended to a superorder as “ Thripida ”.</p><p>Diagnosis.</p><p>Forewing with CuA re-emerging orthogonally from R + M + CuA.</p><p>Included families.</p><p>Lophioneuridae Tillyard, 1921 (spectral thrips) and Iotacyphidae Shcherbakov, Bashkuev, &amp; Shmakov, 2024 (shielded thrips).</p></div>	https://treatment.plazi.org/id/22E6595FCC9E5AF98C6C59F6598F1A19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Boderau, Mathieu;Montagna, Matteo;Magnani, Fabio;Magoga, Giulia;Shmakov, Alexey;Engel, Michael S.;Nel, Patricia;Nel, André	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
AF2C7FF3A7105D6C85FAD56905D9004A.text	AF2C7FF3A7105D6C85FAD56905D9004A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Holothysanoptera Engel, Montagna, Boderau, P. Nel, & A. Nel	<div><p>3.1. Superorder Holothysanoptera Engel, Montagna, Boderau, P. Nel, &amp; A. Nel nov.</p><p>(Carboniferous Westphalothripidesidae P. Nel, Azar, Prokop, Roques, Hodebert, &amp; A. Nel, 2012 to crown-Thysanoptera)</p><p>(= “ Thripida Fallén, 1814 ” sensu P. Nel et al. 2014)</p><p>Diagnosis.</p><p>Left mandibular stylet, right absent; piercing lacinial stylets; pretarsus with eversible arolium; pretarsal claws reduced in adults; wings with fringe of long setae and CuA simple.</p><p>Included clades.</p><p>Westphalothripidesidae, Lophioneuroptera (spectral and shielded thrips), Betathysanoptera Engel ( Zoropsocidae Tillyard, 1935, beta thrips), Thysanoptera (true thrips).</p></div>	https://treatment.plazi.org/id/AF2C7FF3A7105D6C85FAD56905D9004A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Boderau, Mathieu;Montagna, Matteo;Magnani, Fabio;Magoga, Giulia;Shmakov, Alexey;Engel, Michael S.;Nel, Patricia;Nel, André	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
13725FD7043C51B98F4BAF5807C83B3B.text	13725FD7043C51B98F4BAF5807C83B3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triassocypha Boderau & Montagna & Magnani & Magoga & Shmakov & Engel & Nel & Nel 2025	<div><p>3.5. Triassocypha gen. nov.</p><p>Type species.</p><p>Triassocypha prima sp. nov.</p><p>Etymology.</p><p>The name is an artificial combination of Triassic, referring to period from which the fossils originate, and the Ancient Greek adjective κῡφός (kūphós, meaning, “ bent forward ” or “ curved ”; feminine κῡφή). The gender of the name is feminine.</p><p>Diagnosis.</p><p>Forewing ScP re-emerging from R as a short, straight veinlet perpendicular to C; a closed cell between MA and MP; two longitudinal cells between R and M / MA; CuP + PCu well separated from A.</p><p>Included species.</p><p>Triassocypha prima sp. nov. and T. secunda sp. nov., both from the Triassic of Monte San Giorgio.</p></div>	https://treatment.plazi.org/id/13725FD7043C51B98F4BAF5807C83B3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Boderau, Mathieu;Montagna, Matteo;Magnani, Fabio;Magoga, Giulia;Shmakov, Alexey;Engel, Michael S.;Nel, Patricia;Nel, André	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
8033400C167853E18DD29D3F80C6CB52.text	8033400C167853E18DD29D3F80C6CB52.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triassocypha prima Boderau & Montagna & Magnani & Magoga & Shmakov & Engel & Nel & Nel 2025	<div><p>3.6. Triassocypha prima sp. nov.</p><p>Figures 2 A, B, D, 3 A</p><p>Material.</p><p>Holotype, MCSN 8715 (a body with two forewings and one hind wing, legs, and one antenna apparently complete), stored in Museo cantonale di storia naturale, Lugano, Switzerland.</p><p>Locality and horizon.</p><p>Upper Kalkschieferzone member of Meride Limestone (transition interval to Archelaus and Regoledanus Ammonoid Zones, upper Ladinian), Middle Triassic; VM 12 site (DMS geographical coordinates: 45°53’26’’N; 8°56’49’’E), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=8.946944&amp;materialsCitation.latitude=45.890556" title="Search Plazi for locations around (long 8.946944/lat 45.890556)">Val Mara near Meride</a>, Monte San Giorgio, Switzerland.</p><p>Etymology.</p><p>The specific epithet is the Latin adjective prīmus, meaning, “ first ” or “ early ”, and refers to this as the first described species of the genus.</p><p>Diagnosis.</p><p>Forewing characters only: ma-mp longer than r-ma; distal-most cell between R and MA three times as long as wide.</p><p>Description.</p><p>Body 1.3 mm long, presence of long macrosetae on the thorax and some wing veins. Head: head apparently orthognathous to prognathous, with a relatively long buccal cone; compound eyes rounded with diameter 0.1 mm; antenna elongate, with flagellomeres elongate, distal most flagellomeres shorter than other flagellomeres, scape large, pedicel apparently small. Thorax: thorax 0.5 mm long; all legs preserved and of similar shapes, apparently only two tarsomeres, basal one light brown and distal one dark brown. Abdomen: abdomen elongate and broad, 0.7 mm long, 0.55 mm wide; cerci absent. Forewing venation: forewing 1.6 mm long, 0.6 mm wide, pear-shaped, rounded apically, narrow basally; a common stem ScP + R + M + CuA from which CuA separates 0.2 mm from wing base, perpendicular to ScP + R + M and short before cua-cup terminates into vein; cua-cup 0.07 mm long, straight, aligned with distal part of CuA; CuA simple and curved distally, ending at wing margin 0.5 mm from wing base; CuP + PCu straight, well separated from A; A short, weakly curved; ScP re-emerging from R 0.8 mm from wing base, perpendicular to C and R, straight, 0.12 mm long; M separating from R 0.1 mm distal of base of CuA; stem of M curved, 0.25 mm long, separating into MA and MP and delimiting a broad elongate closed cell, 0.5 mm long, 0.2 mm wide; both MA and MP with a strong angle distally and ending on posterior margin at right angle; a long ma-mp perpendicular to MA and MP; two elongate, broad cells between R and MA, separated by r-ma perpendicular to R and MA, basal-most cell 0.35 mm long, 0.07 mm wide; distal-most cell 0.6 mm long, 0.2 mm wide; R forming an angle distal of ScP and straight distally; separating into RA and RP 0.45 mm distal of ScP; RA straight, short, rather oblique, nearly orthogonal with RP; RP forming a strong angle and reaching wing apex; straight, elongate rp-ma, 0.1 mm long.</p></div>	https://treatment.plazi.org/id/8033400C167853E18DD29D3F80C6CB52	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Boderau, Mathieu;Montagna, Matteo;Magnani, Fabio;Magoga, Giulia;Shmakov, Alexey;Engel, Michael S.;Nel, Patricia;Nel, André	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
05146F1C0D605D0C87B67D81E8B3237C.text	05146F1C0D605D0C87B67D81E8B3237C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triassocypha secunda Boderau & Montagna & Magnani & Magoga & Shmakov & Engel & Nel & Nel 2025	<div><p>3.7. Triassocypha secunda sp. nov.</p><p>Figures 2 C, E, 3 B</p><p>Material.</p><p>Holotype MCSN 8689 (a body with two forewings and fragments of legs), stored in Museo cantonale di storia naturale, Lugano, Switzerland.</p><p>Locality and horizon.</p><p>Upper Kalkschieferzone member of Meride Limestone (transition interval to Archelaus and Regoledanus Ammonoid Zones, upper Ladinian), Middle Triassic; VM 12 site (DMS geographical coordinates: 45°53’26’’N; 8°56’49’’E), <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=8.946944&amp;materialsCitation.latitude=45.890556" title="Search Plazi for locations around (long 8.946944/lat 45.890556)">Val Mara near Meride</a>, Monte San Giorgio, Switzerland .</p><p>Etymology.</p><p>The specific epithet is the Latin numeral secundus, meaning “ second ”, and refers to this as the second described species of the genus.</p><p>Diagnosis.</p><p>Forewing characters only: ma-mp as long as r-ma; distal-most cell between R and MA two times as long as wide.</p><p>Description.</p><p>Body 1.4 mm long, presence of long macrosetae on the head, thorax, and some wing veins. Head: head apparently orthognathous to prognathous, with a relatively long buccal cone; compound eyes rounded with diameter 0.1 mm; [antenna not preserved]. Thorax: 0.37 mm long, 0.4 mm high; legs partly preserved. Abdomen: broad, 0.9 mm long, 0.7 mm wide; cerci absent. Forewing venation: forewing 1.7 mm long, 0.8 mm wide, pear-shaped, rounded apically, narrow basally; a common stem ScP + R + M + CuA; CuA, cua-cup, and CuP not visible; ScP re-emerging from R 0.8 mm from wing base, perpendicular to C and R, straight, 0.15 mm long; M separating from R 0.6 mm distal of base of CuA; stem of M curved, 0.5 mm long, separating into MA and MP and delimiting a broad elongate closed cell, 0.4 mm long, 0.19 mm wide; both MA and MP with a strong angle distally and ending on posterior margin at right angle; a long ma-mp perpendicular to MA and MP; two elongate, broad cells between R and MA, separated by r-ma perpendicular to R and MA, basal-most cell 0.4 mm long, 0.15 mm wide; distal-most cell 0.46 mm long, 0.23 mm wide; R forming an angle distal of ScP and straight distally, separating into RA and RP 0.5 mm distal of ScP; RA straight, short, rather oblique, nearly orthogonal with RP; RP forming a strong angle, reaching wing apex; straight, elongate rp-ma, 0.12 mm long.</p></div>	https://treatment.plazi.org/id/05146F1C0D605D0C87B67D81E8B3237C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Boderau, Mathieu;Montagna, Matteo;Magnani, Fabio;Magoga, Giulia;Shmakov, Alexey;Engel, Michael S.;Nel, Patricia;Nel, André	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
CF6F0A2E69C4546DBB9958ECC8B0F5CB.text	CF6F0A2E69C4546DBB9958ECC8B0F5CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Triassocyphinae Boderau & Montagna & Magnani & Magoga & Shmakov & Engel & Nel & Nel 2025	<div><p>3.4. Triassocyphinae subfam. nov.</p><p>Type genus.</p><p>Triassocypha gen. nov.</p><p>Diagnosis.</p><p>Two closed cells between R and M and a closed cell between the branches of M.</p></div>	https://treatment.plazi.org/id/CF6F0A2E69C4546DBB9958ECC8B0F5CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Boderau, Mathieu;Montagna, Matteo;Magnani, Fabio;Magoga, Giulia;Shmakov, Alexey;Engel, Michael S.;Nel, Patricia;Nel, André	Boderau, Mathieu, Montagna, Matteo, Magnani, Fabio, Magoga, Giulia, Shmakov, Alexey, Engel, Michael S., Nel, Patricia, Nel, André (2025): Triassic iotacyphids shed light on the venation of crown Thysanoptera and their stem relatives (Condylognatha: Holothysanoptera). Arthropod Systematics & Phylogeny 83: 521-530, DOI: 10.3897/asp.83.e161465
