taxonID	type	description	language	source
914C87C7241AFFE56AA7F914FC60FDB8.taxon	type_taxon	Type species: Diestrammena palpata Rehn, 1906, by original designation	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7241AFFE56AA7F914FC60FDB8.taxon	description	The genus Diestramima was defined by Storozhenko (1990) and has 42 recognised species divided among a number of subgenera (Cigliano et al. 2025). The general characteristics that unite the species within this genus, such as the projection from 7 th abdominal tergite of males are described here. HEAD. Oval in shape. Face smooth and shiny, and usually with short hair on the dorsal surface of head, maxillary palps and antennae. Fastigium vertici always projecting forward, dividing into apical half of conical tubercles. Bases of tubercles drawn together with obtuse apices. Eyes oval, almost the size of antennal scapi. Median ocellus oval to oblong, and lateral ocelli oval or almost circular situated on lateral surface of basal fastigium vertici. THORAX. Body medium in size with abundant fine black and brown hairs (Table 2). Anterior margin of the pronotum straight, posterior margin protruding and obtusely rounded, posterolateral margins almost rounded. The posterior margin of mesonotum obviously protruding backward, lateral margins angular; posterior margin of metanotum rather straight with almost circular lateral margins. Posteromedian projection of 7 th abdominal tergite variable from short to long covering distal half or entire part of paraproct (Fig. 7 & 8). The shape of the 7 th abdominal tergite projection is described under each species account. The paraproct variable with short to long almost finger-like structure (Fig. 7 & 8). The cerci usually slender, conical, and have acute apices. LEGS. Legs long compared to Rhaphidophora (Table 2). Fore femora with single short, retrolateral, apical spine and mid femora with a pair of prominent apical spines (one prolateral and one retrolateral). Hind femora with 5 – 13 inferior retrolateral spines (Table 3). Fore and mid tibiae each with four inferior linear spines (two prolateral and two retrolateral), three inferior apical spines (one prolateral, one ventral and one retrolateral) and with one superior apical spine on fore tibiae (retrolateral) and two superior apical spines on mid tibiae (one prolateral and one retrolateral). Hind tibiae with variable number of linear, short, fixed spines on superior surface (in two rows of prolateral and retrolateral) (Table 3), and typical eight, articulated apical spines. The longest pair of these (sometimes referred to as spurs) are shorter than 1 st and / or 2 nd segments of hind tarsi combined. The 1 st tarsal segment of hind tibiae have between one to four superior spines. COLOURATION. Head with short brown or black hairs. Face white and glabrous always with 4 vertical black stripes below eyes and inner sides of antennal scapes, and a pair of lateral marks on clypeus (Fig. 8 A). Body usually black with small brown or yellowish spots on thoracic (dorsolateral) and abdominal tergites, covered in black and brown short hairs. Legs dark and pale speckled, hind femur dark to pale speckled especially at proximal half.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72418FFEA6AA7FAE9FA39F844.taxon	description	(Figure 6 A – K) Thirteen specimens in our collections were identified as D. tsongkhapa. These were collected from four sites in central Bhutan, increasing the known distribution of this species (Fig. 4). Here we present images and summary of the key traits of this species (Fig. 6).	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72418FFEA6AA7FAE9FA39F844.taxon	materials_examined	Material examined. BHUTAN • 5 adult females; Philligang, Nabji, Korphoog, Trongsa; 27.178 ° N, 90.535 ° E; 1381 m asl; 30 Jul. 2022; Sanjit Rai leg.; Cool broadleaf forest: under fallen tree branches and leaf litter; MPN _ CW 5516, MPN _ CW 5520, MPN _ CW 5517, MPN _ CW 5518, MPN _ CW 5519. • 4 adult males; Philligangju, Nabji, Korphoog, Trongsa; 27.180 ° N, 90.535 ° E; 1305 m asl; 30 Jul. 2022; Sanjit Rai leg.; Cool broadleaf forest: under fallen tree branches and leaf litter; MPN _ CW 5522, MPN _ CW 5521, MPN _ CW 5523, MPN _ CW 5524. • 2 male and 2 female adults; Namkye, Nabji, Korphoog, Trongsa; 27.165 ° N, 90.562 ° E; 2252; m asl; 22 Jul. 2022; Sanjit Rai leg.; Cool broadleaf forest: under fallen tree branches and leaf litter; MPN _ CW 5525, MPN _ CW 5526, MPN _ CW 5527, MPN _ CW 5528. Male. Body medium in size (~ 20 mm). Median ocellus oval and lateral ocelli circular situated on lateral surface of basal fastigium vertici. Posteromedian projection of 7 th abdominal tergite rather high in profile and long (Fig. 6 E), obviously surpassing apices of paraproct, gradually narrowing towards apex when viewed dorsally, apex slightly extended with distinct notch (Fig. 6 D). Paraproct with broad proximal half tapering abruptly and curved upward, apex obtuse (Fig. 6 C & D). Subgenital plate large, curved, posterior edge slightly crimped, apex rounded (Fig. 6 C). Cerci slender, conical, apices almost acute. Linear measures of body elements (Table 2). Fore and mid leg spine details as per the generic description. Hind leg spine details (Table 3). Head dorsally brown with abundant short brown hair. Body black with many small brown or yellowish spots on thoracic and abdominal tergites, covered in black and brown short hairs. A pale, thin median line present but inconspicuous (Fig. 6 F). Paraproct black when viewed dorsally. Cerci white with black spots on dorsal side with short yellowish-brown hair and longer hair.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72418FFEA6AA7FAE9FA39F844.taxon	description	Female. The general appearance is similar to male. Posteromedian projection of 7 th abdominal tergite short and angular with posteromedial apex obtusely rounded completely covering 8 th abdominal tergite, 9 th tergite broader and more rounded. Subgenital plate almost translucent, simple, triangular, apex obtusely rounded, slightly curved upward (Fig. 6 K). Ovipositor slender and longer than body, slightly curved, apices acute, distal half of lower valves crenulated, upper valves smooth (without hair along the margins) (Fig. 6 I & J). Size of body elements (Table 2) and combination of leg spines (Table 3) are similar to male.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72416FFE86AA7FF19F8B9FC84.taxon	description	(Figure 7 A – K)	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72416FFE86AA7FF19F8B9FC84.taxon	materials_examined	Holotype. BHUTAN • 1 adult male; Guru Drakphu Nye, Wangphu, Gomdar, Samdrup Jongkhar; 26.982 ° N, 91.584 ° E; 1028 m asl; 29 May 2022; C. Dorji & Jigme leg.; Subtropical forest: limestone cave entrance; MPN _ CW 5486; NBCB 146. Paratype. BHUTAN • 1 adult female; Guru Drakphu Nye, Wangphu, Gomdar, Samdrup Jongkhar; 26.982 ° N, 91.584 ° E; 1028 m asl; 29 May 2022; C. Dorji & Jigme leg.; Subtropical forest: limestone cave entrance; MPN _ CW 5487; NBCB 152. Other material. BHUTAN • 2 adult females; Guru Drakphu Nye, Wangphu, Gomdar, Samdrup Jongkhar; 26.982 ° N, 91.584 ° E; 1028 m asl; 29 May 2022; C. Dorji & Jigme leg.; Subtropical forest: limestone cave entrance; MPN _ CW 5485, MPN _ CW 5488. • 1 adult male; Yurung bridge, Khar, Pema Gatshel, 27.040 ° N, 91.380 ° E; 571 m asl; 31 May 2022; C. Dorji, Tshewang Namgay & Jigme leg.; Subtropical forest: cliff base under leaf litter and small rocks; MPN _ CW 5489. • 6 male and 5 female adults; Murang bridge, Khar, Pema Gatshel; 27.011 ° N, 91.420 ° E; 923 m asl; 25 May 2022, 31 May 2022; C. Dorji, Tshewang Namgay & Jigme leg.; Subtropical forest: cliff base under leaf litter and small rocks; MPN _ CW 5479, MPN _ CW 5480, MPN _ CW 5481, MPN _ CW 5482, MPN _ CW 5536, MPN _ CW 5537, MPN _ CW 5538, MPN _ CW 5539, MPN _ CW 5540, MPN _ CW 5490, MPN _ CW 5491. • 3 male and 5 female adults; Khelaphung, Kharungla, Brekha, Trashigang; 27.155 ° N, 91.628 ° E; 2039 m asl; 1 Jul. 2022; C. Dorji & Jigme leg.; Cool broadleaf forest: surrounding huge boulders and nearby vegetation; MPN _ CW 5492; MPN _ CW 5493, MPN _ CW 5494, MPN _ CW 5495, MPN _ CW 5496, MPN _ CW 5497, MPN _ CW 5498, MPN _ CW 5499.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72416FFE86AA7FF19F8B9FC84.taxon	description	Description. Male. Body small in size (<17 mm). Median ocellus oblong and lateral ocelli circular. Posteromedian projection of 7 th abdominal tergite very short never extending beyond tenth abdominal tergite, gradually narrowing when viewed from above (Fig. 7 B & D), truncate or slightly convex apex (Fig. 7 D). Paraproct rather short and almost triangular, apices acute (Fig. 7 B & C). Cerci conical, apices acute. Subgenital plate similar to D. tsongkhapa. Linear measures of body elements (Table 2). Fore and mid leg spine details as per the generic description. Hind leg spine details (Table 3). Female. General appearance is similar to male but larger (body> 14 mm long, hind legs 7 mm longer than male’s; Table 2). Posteromedian projection of 7 th abdominal tergite short and angular with posteromedial apex obtusely rounded covering 8 th abdominal tergite. Ovipositor narrow and almost straight (slightly curved upward at distal half), and as long as body (Fig. 7 I). Lower valves are finely serrated at distal end, with acute apices (Fig. 7 J). Upper valves with equally spaced short hair along dorsal margins (almost starting from the base) (Fig. 7 J), gradually narrowing with slightly acute apex (Fig. 7 I). Subgenital plate broad and triangular with obtusely angular apical part (Fig. 7 K). Linear measures of body elements (Table 2). Fore and mid leg spine details as per the generic description. Hind leg spine details (Table 3). Colouration. Head dorsally brown with abundant short brown-yellow hairs. Body brown with few small yellowish spots along posterior edge thoracic and abdominal tergites, covered in black and brown short hairs. Paraproct black dorsally, and white ventrally. Cerci white with brown spots on dorsal side. The distal half of cerci are covered by short yellowish-brown hair and fewer longer hairs at proximal and mid areas.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72416FFE86AA7FF19F8B9FC84.taxon	etymology	Etymology. “ Matermagna ” is Latin for ‘ great mother’ – because adult females have larger bodies and longer hind legs than males.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72415FFE96AA7FC21F9D0FDA2.taxon	description	(Figure 8 A – H)	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72415FFE96AA7FC21F9D0FDA2.taxon	materials_examined	Holotype. BHUTAN • 1 adult male; Khandro Yeshi Tshogyel Nye, Minjiwoong peg, Serthi, Samdrup Jongkhar; 26.988 ° N, 92.012 ° E; 833 m asl; 28 May 2022; C. Dorji, Tshering Dorji & Jigme leg.; Subtropical forest: cave entrance; MPN _ CW 5484; NBCB 147.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72415FFE96AA7FC21F9D0FDA2.taxon	description	Description. Male. Body medium in size (~ 12 mm). Median ocellus oval, and lateral ocelli almost circular situated on lateral surface of basal fastigium vertici. Posteromedian projection of 7 th abdominal tergite is short covering only distal half of paraproct with lateral sides nearly parallel (Fig. 8 E), lateral and apical margins slightly curved inwards with truncate apex (Fig. 8 D). Paraproct rather short and triangular on lateral view, apices acute. Cerci slender, conical, apices acute. Linear measures of body elements (Table 2). Fore and mid leg spine details as per the generic description. Hind leg spine details (Table 3). Colouration. Head dorsally black with abundant short brown-black hairs. Body black with small brown or yellowish spots on thoracic (dorsolateral) and abdominal tergites, covered in black and brown hair. A pale, thin median line is present but inconspicuous (Fig. 8 F). Paraproct white with black spots at the distal end when viewed dorsally (Fig. 8 C). Cerci with two-thirds end black with dense short yellowish-brown hairs and fewer longer hairs in the mid region. Female. Unknown.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72415FFE96AA7FC21F9D0FDA2.taxon	etymology	Etymology. The new species is named for the type locality Minjiwoong under Serthi geog, Samdrup Jongkhar district.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72414FFE96AA7FD43F9E0F8BE.taxon	description	(Figure 9 A – K)	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72414FFE96AA7FD43F9E0F8BE.taxon	materials_examined	Holotype. BHUTAN • 1 adult male; Bawoong, Ramjar, Trashi Yangtse; 27.416 ° N 91.586 ° E; 1582 m asl; 2 Jul. 2022; C. Dorji & Jigme leg.; Temperate forest: under boulders and surrounding vegetation; MPN _ CW 5500; NBCB 148. Paratype. BHUTAN • 1 adult female; Bawoong, Ramjar, Trashi Yangtse, 27.416 ° N, 91.586 ° E; 1582 m asl; 2 Jul. 2022; C. Dorji & Jigme; leg.; Temperate forest: under boulders and surrounding vegetation; MPN _ CW 5534; NBC 153. Other material. BHUTAN • 3 male and 5 female adults; Bawoong, Ramjar, Trashi Yangtse; 27.416 ° N 91.586 ° E; 1582 m asl; 2 Jul. 2022; C. Dorji & Jigme leg.; Temperate forest: under boulders and surrounding vegetation; MPN _ CW 5501, MPN _ CW 5502, MPN _ CW 5503, MPN _ CW 5504, MPN _ CW 5531, MPN _ CW 5532, MPN _ CW 5533, MPN _ CW 5535.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72414FFE96AA7FD43F9E0F8BE.taxon	description	Description. Male. Body medium in size (~ 16 mm). Median ocellus oval and lateral ocelli circular situated on lateral surface of basal fastigium verticis. Posteromedian projection of 7 th abdominal tergite short covering only two-thirds of paraproct (Fig. 9 B & E), narrowing to apex (Fig. 9 D), lateral margins not curved inwards, apex truncate with slight downward bend (Fig. 9 E). Paraproct rather short and triangular on lateral view, apices acute (Fig. 9 C & E). Cerci conical, apices acute. This species is morphologically similar to D. tsongkhapa but differs by its smaller body (15.6 – 16.2 mm), shorter 7 th abdominal tergite with truncated apex, and relatively shorter paraproct. This species is not phylogenetically sister to D. tsongkhapa (Fig. 7). Linear measures of body elements (Table 2). Fore and mid leg spine details as per the generic description. Hind leg spine details (Table 3). Colouration. Head dorsally brown covered by brown and black hairs. A pale, thin median line present but inconspicuous (Fig. 9 F). Rostral tubercles black from above and white on lateral sides and apices. Body brown with small brown spots along posterior edges of thoracic abdominal tergites, covered in black and brown short hairs. Paraproct black when viewed dorsally. Cerci white with black colouration on dorsal part with short yellowish-brown hairs and fewer longer at proximal half. Female. Body colour much lighter than males with few larger dark black spots along posterior edge of thoracic and abdominal tergites (Fig. 9 I). Same body size and leg length as male (Table 2). Posteromedian projection of 7 th abdominal tergite short and angular with posteromedial apex obtusely rounded completely covering 8 th and 9 th abdominal tergite. Subgenital plate membranous, broad and short, almost triangular, apex obtusely rounded, slightly curved upward (Fig. 9 K). Ovipositor narrow and as long as body (Fig. 9 J). Combination of leg spines is similar to males.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72414FFE96AA7FD43F9E0F8BE.taxon	etymology	Etymology. The name refers to the very large number of tiny linear spines on the superior surface of the hind tibiae.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72413FFEF6AA7FA22FB8CF8B3.taxon	description	(Figure 10 A – H and Figure 11 A – F)	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72413FFEF6AA7FA22FB8CF8B3.taxon	materials_examined	Holotype. BHUTAN • 1 adult male; Tshangkha, Tangsibji, Trongsa; 27.440 ° N, 90.420 ° E; 2161 m asl; 6 Jul. 2022; C. Dorji, Karma Chorten & Jigme leg.; Cool broadleaf forest: under fallen tree branches and leaf litter; MPN _ CW 5509; NBCB 149. Paratype. BHUTAN • 1 adult female; Tshangkha, Tangsibji, Trongsa; 27.440 ° N, 90.420 ° E; 2161 m asl; 6 Jul. 2022; C. Dorji, Karma Chorten & Jigme leg.; Cool broadleaf forest: under fallen tree branches and leaf litter; MPN _ CW 5510; NBCB 154. Other material. BHUTAN • 1 adult male; Tshangkha, Tangsibji, Trongsa; 27.440 ° N, 90.420 ° E; 2161 m asl; 6 Jul. 2022; C. Dorji, Karma Chorten & Jigme leg.; Cool broadleaf forest: under fallen tree branches and leaf litter.; MPN _ CW 5508. • 1 adult female; Philligang, Nabji, Korphoog, Trongsa; 27.178 ° N, 90.535 ° E; 1381 m asl; 30 Jul. 2022; Sanjit Rai leg.; Cool broadleaf forest: under fallen tree branches and leaf litter; MPN _ CW 5515.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72413FFEF6AA7FA22FB8CF8B3.taxon	description	Description. Male. Body small (10 – 13 mm). Median ocellus oval and lateral ocelli circular situated on lateral surface of basal fastigium verticis. Posteromedian projection of 7 th abdominal tergite long, obviously surpassing apices of paraproct (Fig. 10 E & F; Fig. 11 A). The first half concaved laterally from the sides (Fig. 10 E; Fig. 11 A), second half flat and parallel curving downwards. The apex slightly curved inwards or downward (Fig. 10 E; Fig. 11 C), apical area of posteromedian projection dichotomous and acute when viewed from ventral side (Fig. 10 C & E; Fig. 11 C). Paraproct rather long (not as long as D. tsongkhapa), proximal half broad, narrowing towards apex, obtuse, slightly curved from middle directing backward. Subgenital plate similar to sister species D. tsongkhapa (Fig. 6). Cerci slender, conical, apices obtuse. Linear measures of body elements (Table 2). Fore and mid leg spine details as per the generic description. Hind leg spine details (Table 3). Colouration. Head dorsally yellowish or white and black. The body is mostly very dark to black with or without small pale spots and numerous short black and yellow hairs. A pale dorsomedial stripe (think or thin) extends from the pronotum to the 2 nd or 6 th abdominal tergite (Fig. 10 F). White spots possible on metanotum and 1 st tergite (Fig. 10 F). Paraproct black. Cerci black with white on ventral and basal parts, and with dense short yellowish-brown hairs and few long pale hairs at the base (Fig. 10 C). Female. Body colour darker than male without white spots on metanotum and 1 st abdominal tergite. Median line is rather thin extending only until 2 nd abdominal tergite. Posteromedian projection of 7 th abdominal tergite short and angular with posteromedial apex rounded completely covering 8 th abdominal tergite (Fig. 11 D). Subgenital plate almost translucent, broad and short, almost convex with obtusely rounded apex (Fig. 11 F). Ovipositor slightly longer than body length, thin. Upper valves broad and apex acute. Lower valves with numerous fine teeth from below the apex (Fig. 11). Size of body elements (Table 2) and leg spine combination (Table 3) are similar to male.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72413FFEF6AA7FA22FB8CF8B3.taxon	etymology	Etymology. The new species is named in honour of late Dr. Phub Dorji, the first president of the College of Natural Resources, Royal University of Bhutan.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72411FFF26AA7FA9AFD54FE50.taxon	description	(Figures 12 A – K)	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72411FFF26AA7FA9AFD54FE50.taxon	materials_examined	Holotype. BHUTAN • 1 adult male; Samkhar bridge, Gelephu, Sarpang; 27.009 ° N, 90.581 ° E; 942 m asl; 22 May 2022; C. Dorji & Jigme leg.; Subtropical forest: under boulders above bridge; MPN _ CW 5541; NBCB 150. Paratype. BHUTAN • 1 adult female; Samkhar bridge, Gelephu, Sarpang; 27.009 ° N, 90.581 ° E; 942 m asl; 22 May 2022; C. Dorji & Jigme leg.; Subtropical forest: under boulders above bridge; MPN _ CW 5478; NBCB 155. Other material. BHUTAN • 2 adult females; Samkhar bridge, Gelephu, Sarpang; 27.009 ° N, 90.581 ° E; 942 m asl; 22 May 2022; C. Dorji & Jigme leg.; Subtropical forest: under boulders above bridge; MPN _ CW 5542, MPN _ CW 5543.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72411FFF26AA7FA9AFD54FE50.taxon	description	Description. Male. Body small (<14 mm). Median ocellus oblong and lateral ocelli circular. Posteromedian projection of 7 th abdominal tergite short covering only proximal part of paraproct (Fig. 12 B & E), shallowly grooved from dorsomedial until apex (Fig. 12 C), barely wider, almost parallel lateral edges and truncate with small notch at median part of apex (Fig. 12 C). Paraproct rather short and almost triangular on lateral view, apices acute (Fig. 12 D & E). Cerci conical, apices acute, Subgenital plate similar to D. tsongkhapa. Linear measures of body elements (Table 2). Fore and mid leg spine details as per the generic description. Hind leg spine details (Table 3). Colouration. Head dorsally brown covered by brown-yellow hairs. Rostral tubercles brown from above. Maxillary palps brown with white in mid region of each segment. Body brown with few small brown / white spots along posterior edges of thoracic and abdominal tergites, covered in black and brown short hairs. A pale, thin median line present but inconspicuous extending only up to 2 nd or 3 rd tergites (Fig. 12 F). Paraproct black when viewed dorsally. Cerci white with black spots on dorsal side of distal half, and with short yellowish-brown hair and fewer longer hair at proximal half. Female. General appearance is similar to male, but body colour can be lighter with reddish brown on dorsal parts of thoracic tergites. White spots dorsomedial or along posterior edges of thoracic and abdominal tergites (Fig. 12 J). Posteromedian projection of 7 th abdominal tergite short and angular with posteromedial apex obtusely rounded completely covering 8 th abdominal tergite. Ovipositor narrow and almost straight, and shorter than body, slightly curved at apex. Lower valves without serration, acute. Upper valves with hair along the margins, slightly acute at apex. The size of body elements and combination of leg spines are similar to male.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C72411FFF26AA7FA9AFD54FE50.taxon	etymology	Etymology. The species is named after the type locality Samkhar, Sarpang district.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240FFFF26AA7FE71F8C8F84C.taxon	type_taxon	Type species Rhaphidophora picea Serville, 1838	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240FFFF26AA7FE71F8C8F84C.taxon	description	The genus Rhaphidophora was defined in 1838 with a single female specimen of one species and now includes 129 species (Cigliano et al. 2025). The general characteristics of the genus are agreed (Gorochov, 1999, 2012) and here we summarise the appearance of Rhaphidophora species considering traits that are shared. HEAD. Oval, smooth and shiny except for maxillary palps and antennae covered in dense short hair. Fastigium projecting forward with longitudinal furrow along midline on dorsal surface, partially dividing apical half into a pair of rostral tubercles. Eyes oval, slightly bigger than antennal scapi. Median ocellus situated in between ventral margins of antennal scapi and, lateral ocelli occupying basal two-thirds of lateral surface of rostral tubercles. THORAX. Body medium size and with abundant fine black or brown hairs. Anterior margin of pronotum generally straight and posterior margin obtusely convex or nearly straight. The posterolateral margin of pronotum straight or concave while posterolateral margins of mesonotum and metanotum are slightly oblique or almost rounded. LEGS. Legs moderately long (Table 4). Fore femora with single short, prolateral, apical spine and mid femora with a pair of prominent apical spines (one prolateral and one retrolateral). Hind femora unarmed but with apical inferior spine (retrolateral). Fore tibiae with three inferior linear spines (one prolateral and two retrolateral), and two apical spines (one prolateral and one retrolateral). Mid tibiae with four superior linear spines (two prolateral and two retrolateral) and five inferior linear spines (three prolateral and two retrolateral), and two apical spines on inferior and superior sides (one prolateral and one retrolateral). Hind tibiae with variable number of linear, short, fixed spines on superior surface (in two rows of prolateral and retrolateral) (Table 4), and typical eight, articulated apical spines. The longest pair of these (= spurs) are longer than 1 st and / or 2 nd segments of hind tarsi combined. The 1 st tarsal segment of hind tibiae have variable number of superior spines. MALE TERMINALIA. Adult male terminalia show significant differences among species and are widely used as a key morphological trait for diagnosing species within this genus. A description of terminalia is provided for each newly described species. Abdominal tergites usually without distinct posterior projections. The epiproct is weakly specialised, broad always projecting behind body. The posterior margin may bear one to two sclerotised spines of variable size and shape. Genitalia membranous without any sclerotization. Cerci slender, conical and usually longer than epiproct, covered in both short and long hairs. Subgenital plate is broad and fairly similar. Styli variable from stout to somewhat long in short hairs. COLOURATION. Head and pronotum dorsally dark brown, face and mouthparts brownish. Rostral tubercles usually dark brown. Compound eyes small and black, brown or sometimes with maculation of black and white. Maxillary palps usually pale or with black and white distinct band on each segment. Antennae longer than body mostly pale brown in colour. Body dark brown or yellowish brown with irregular white patches along the posterior of abdominal tergites from above and slightly yellowish brown from beneath. Legs are general yellowish with somewhat darker colouration in joints. Hind femur paler speckled at proximal end and darker to distal end. Cerci and styli either black or white or both. NYMPHS. General appearance of the nymph is usually similar to adults but terminalia used for distinguishing species are not well developed.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240EFFF36AA7FBC9FD2CF88F.taxon	description	(Figure 14 A – G)	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240EFFF36AA7FBC9FD2CF88F.taxon	materials_examined	Holotype. BHUTAN • 1 adult male; Marung bridge, Khar, Pema Gatshel; 27.011 ° N, 91.420 ° E; 923 m asl; 25 May 2022; C. Dorji, Tshewang Namgay & Jigme leg.; Subtropical forest: cliff base under leaf litter and small rocks; MPN _ CW 5483; NBCB 145. Other material. BHUTAN • 1 nymph female; Bali-Pothey, Kilkhorthang, Tsirang; 27.464 ° N, 90.059 ° E; 1266 m asl; 1 May 2022; C. Dorji & Yeshi Phuntsho leg.; Temperate forest: under two huge boulders; MPN _ CW 5470.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240EFFF36AA7FBC9FD2CF88F.taxon	description	Description. Male. Body medium in size (~ 25 mm). Median and lateral ocelli oval. Epiproct oval when viewed dorsally, not strongly curved but deeply grooved with a conspicuous pair of projections on the distal end directed slightly backwards (Fig. 14 E). Ventral part of epiproct triangular, with a pair of large swollen lobes directed upward and triangular sclerotised apical lobe that usually remains inserted into genital cavity (Fig. 14 E & F). Cerci almost twice as long as epiproct. Paraproct obtusely rounded (Fig. 14 F). Subgenital plate with two ovoid lobs not completely divided (Fig. 14 E). Styli stout (as short as the length of subgenital plate), more cylindrical than conical (Fig. 14 F & G). Linear measures of body elements (Table 4). Fore and mid leg spine details as per the generic description except mid tibiae with two linear superior spines (one prolateral and one retrolateral). Hind leg spine details (Table 5). Colouration. Body brownish from above and yellowish brown from beneath. Legs light to medium brown. Head brownish dorsally, face yellowish with 4 short vertical black stripes on epicarnium below eyes and antennae scapi (Fig. 14 C). Epiproct white / yellowish with light black lining along groove. Cerci and styli white with dense yellowish brown short hair and fewer longer hair along cerci. Female. Unknown.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240EFFF36AA7FBC9FD2CF88F.taxon	etymology	Etymology. The new species is named in recognition of the Bhutan nation.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240DFFF66AA7F9BAF891FF20.taxon	description	(Figure 15 A – G)	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240DFFF66AA7F9BAF891FF20.taxon	materials_examined	Holotype. BHUTAN • 1 adult male; Botanical Park, Lamperi, Thimphu; 27.498 ° N, 89.767 ° E; 2761 m asl; 22 Aug. 2022; C. Dorji leg.; Cool broadleaf forest: forest floor under leaf litter; MPN _ CW 5529; NBCB 144.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240DFFF66AA7F9BAF891FF20.taxon	description	Description. Male. Body small in size (<15 mm). Median and lateral ocelli almost oval and same in size. Lateral surface of a pronotum with central dark brown spots but inconspicuous (oval) (Fig. 15 A). All abdominal tergites without distinct posterior projections. Posterior margin of tenth abdominal tergite concave. Epiproct simple, broad, oval, narrowing towards apex with carinated margin bearing a pair of sclerotised ventral spines (Fig. 15 F & G). Ventral basal plate of epiproct triangular, forming a large single sclerotised apical spine (Fig. 15 E). Cerci slender, conical and three times longer than epiproct. Styli stout and cylindrical with obtuse apex. Linear measures of body elements (Table 4). Fore and mid leg spine details as per the generic description. Hind leg spine details (Table 5). Colouration. Body yellowish brown with dark brown irregular patches along the posterior of abdominal tergites dorsally (Fig. 15 A & B) and slightly yellowish brown from beneath. Cerci and styli with dense yellowish brown short hair and fewer longer hair along the cerci. Female. Unknown.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240DFFF66AA7F9BAF891FF20.taxon	etymology	Etymology. “ Bicuspis ” is Latin for ‘ two sides’ or ‘ two cusps’ – derived from the paired spines on the ventral margin of male epiproct.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240BFFF76AA7FEC4FB36FAA1.taxon	description	(Figure 16 A – K)	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240BFFF76AA7FEC4FB36FAA1.taxon	materials_examined	Holotype. BHUTAN • 1 adult male; Tshangkha, Tangsibji, Trongsa; 27.440 ° N, 90.420 ° E; 2161 m asl; 6 Jul. 2022; C. Dorji, Karma Chorten & Jigme leg.; Cool broadleaf forest: under fallen tree branches and leaf litter; MPN _ CW 5505; NBCB 142. Paratype. BHUTAN • 1 adult female; Namkye, Nabji, Korphoog, Trongsa; 27.166 ° N, 90.562 ° E; 2252 m asl; 11 Aug. 2022; Sanjit Rai leg.; Cool broadleaf forest: under fallen tree branches and leaf litter; MPN _ CW 5544; NBCB 151. Other material. BHUTAN • 2 nymph males; Tshangkha, Tangsibji, Trrongsa; 27.440 ° N, 90.420 ° E; 2161 m asl; 6 Jul. 2022; C. Dorji, Karma Chorten & Jigme leg.; Cool broadleaf forest: under fallen tree branches and leaf litter.; MPN _ CW 5506, MPN _ CW 5507. BHUTAN • 1 female; Namkye, Nabji, Korphoog, Trongsa; 27.166 ° N, 90.562 ° E; 2252 m asl; 11 Aug. 2022; Sanjit Rai leg.; Cool broadleaf forest: under fallen tree branches and leaf litter; MPN _ CW 5545	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240BFFF76AA7FEC4FB36FAA1.taxon	description	Description. Male. Body small in size (<16 mm). Median ocellus oval and lateral ocelli slightly oblong or oval, occupying basal two-thirds of lateral surface of the rostral tubercles. Posterior median margin of 9 th abdominal tergite less rounded, embedded within 8 th abdominal tergite (Fig. 16 F). Tenth abdominal tergite with obtuse projection at the posterolateral margin with more convex shaped projection in the middle (Fig. 16 F). Epiproct broad, less strongly curved but with two distinct grooves on the dorsal surface (Fig. 16 F & G), with a pair of distal projections on the posterior median curved slightly upwards (Fig. 16 E & G). Ventral basal plate of epiproct quadrate, with a pair of large lobes directed upward away from each other (usually backward when inserted to genitalia cavity) (Fig. 16 E). These lobes are as long as the length of ventral basal plate of epiproct, with a pair of short sclerotised apical margin spines (Fig. 16 E & H). Cerci slender and cylindrical, apices obtuse. Subgenital plate much wider than long forming oval lobes with wide space in between. Styli short, conical, arcuate with sclerotised acute apices (Fig. 16 H). Linear measures of body elements (Table 4). Fore and mid leg spine details as per the generic description except mid tibiae with three superior linear spines (one prolateral and two retrolateral). Hind leg spine details (Table 5). Colouration. Head and pronotum mostly dark brown or reddish and black dorsally. Body brown with numerous small pale brown or yellowish spots along posterior edges of thoracic and abdominal tergites (Fig. 16 E), and slightly yellowish brown from beneath. Rostral tubercles dark brown. Cerci blackish in the centre and white on the ends. Female. Larger than male (Table 4), body medium in size (> 25 mm). Median and lateral ocelli same as male holotype. Subgenital plate much wider than long, simple, more convex than triangular, basal with small depression in the centre (Fig. 16 J). Ovipositor base not so broad, long and gradually narrowing towards apex, curved, apices acute. Lower valves serrated on distal area of ventral margin, and upper valves smooth. The general colouration is similar to male except body with more small brown / yellowish spots along posterior edges of thoracic and abdominal tergites. Linear measures of body elements (Table 4). Fore and mid leg spine details as per the generic description except mid tibiae with only three inferior linear spines (retrolateral). Hind leg spine details (Table 5).	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240BFFF76AA7FEC4FB36FAA1.taxon	etymology	Etymology. “ Bilobata ” from the Latin for ‘ two lobes’ – referring to the specialised ventral basal plate of epiproct, which forms a pair of long lobes.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240AFFF46AA7FA44FDE8F886.taxon	description	(Figure 17 A – H)	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240AFFF46AA7FA44FDE8F886.taxon	materials_examined	Holotype. BHUTAN • 1 adult male; Mahadev Dham, Tashiding, Dagana; 26.899 ° N, 89.975 ° E; 915 m asl; 21 May 2022; Jigme Wangchuk & Galey leg.; Subtropical forest: surrounding limestone cave under leaf litter; MPN _ CW 5475; NBCB 143. Other material. BHUTAN • 2 male and 1 female nymphs; Mahadev Dham, Tashiding, Dagana; 26.899 ° N, 89.975 ° E; 915 m asl; 21 May 2022; Jigme Wangchuk & Galey leg.; Subtropical forest: surrounding limestone cave; MPN _ CW 5474, CW 5476, CW 5477.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240AFFF46AA7FA44FDE8F886.taxon	description	Description. Male. Body medium in size (<20 mm). Median ocellus elliptic or oblong (Fig. 17 C) slightly longer than its width; lateral ocelli oval, occupying basal two-thirds of lateral surface of the rostral tubercles. Posterolateral margins on left side of mesonotum and metanotum are different from the right possibly because of incomplete ecdysis during nymph stages or early-stage injuries (Fig. 17 A &. B). Epiproct broad and long, fishtail shaped when view dorsally. The dorsal surface of epiproct is shallowly grooved (Fig. 17 G), slightly curved laterally, projecting behind body. In about proximal half gradually narrowing with apical margin obtusely excised from the centre forming two lateral obtuse apices (Fig. 17 G). Ventral surface of basal plate of epiproct is almost quadrate with slightly angled from middle on lateral view with a pair of small sclerotised marginal spines slightly projecting outwards, separated by traverse ridge (Fig. 17 E). Cerci slender, conical and slightly longer than epiproct. Paraproct acutely triangular (Fig. 17 F). Subgenital plate width as wide as length, with two distinct large lobes (Fig. 17 E). Styli stout and simple, conical and apical obtuse without sclerotization. Linear measures of body elements (Table 4). Fore and mid leg spine details as per the generic description except mid tibiae with only three inferior linear spines (retrolateral). Hind leg spine details (Table 5). Colouration. Head dorsally back with brownish face and mouthparts. Rostral tubercles dark black. Eyes silver white and black forming maculation of white and black connecting spots. Body dark brownish from above and slightly yellowish brown from beneath when compared to other species. Epiproct brownish. Cerci dark brown with one-third end white. Cerci and styli white with dense yellowish brown short setae, and fewer longer hairs along first half of the cerci. Female. Adult unknown.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
914C87C7240AFFF46AA7FA44FDE8F886.taxon	etymology	Etymology. Name is derived from fishtail shape of male epiproct, when viewed from above.	en	Dorji, Cheten, Morgan-Richards, Mary, Trewick, Steven A. (2025): Little-known wingless crickets of Bhutan (Rhaphidophoridae): discovery and description of nine new species. Zootaxa 5653 (1): 1-32, DOI: 10.11646/zootaxa.5653.1.1, URL: https://doi.org/10.11646/zootaxa.5653.1.1
