identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
857E879E8E32965FFF40FA9CFD98F9A8.text	857E879E8E32965FFF40FA9CFD98F9A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Scarabaeini Latreille 1802	<div><p>SCARABAEINI Latreille, 1802</p><p>*  Escarabaeus furcatus (de Castelnau, 1840) (Pl. 1, Fig. 1-2)</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 2 ex.</p><p>- P. N. du W, Mamassi-Gourma, 30 IX 2015, lumière, 3 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E32965FFF40FA9CFD98F9A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E32965FFC6FFC87FA82FB38.text	857E879E8E32965FFC6FFC87FA82FB38.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Chalconotus aurescens (Bates 1888)	<div><p>Chalconotus aurescens (Bates, 1888)</p><p>(Pl. 3, Fig. 1)</p><p>- Houyéyogbé, VII 2013, 1 ex., U.V.</p><p>- Bohicon, 14 IV 2012, lumière, 1 ex.</p><p>- P. N. Pendjari, Batia, 18-25 VIII 2010, piège au sol, 4 ex.</p><p>- P. N. Pendjari, VI 2005, nombreux exemplaires.</p><p>- P. N. du W, IX 2003, nombreux exemplaires.</p><p>- P.N.duW,Mamassi-Gourma, 29 IX 2015,lumière, 7 ex.; 30 IX 2015,lumière, 1 ex.</p><p>- [Boucomont, 1933: cercle de Djoungou Kouandé; Kandy. Boucomont, 1933 (sub  variété hopei Kolbe): cercle de Djoungou Kouandé].</p></div>	https://treatment.plazi.org/id/857E879E8E32965FFC6FFC87FA82FB38	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E32965FFC1EFB68FB74FADF.text	857E879E8E32965FFC1EFB68FB74FADF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Chalconotus convexus (Boheman 1857)	<div><p>Chalconotus convexus (Boheman, 1857) (Pl. 3, Fig. 2)</p><p>- P.N.duW,Mamassi-Gourma, 29 IX 2015,lumière, 4 ex.; 30 IX 2015,lumière, 2 ex.; - P. N. Pendjari, VI 2005, nombreux exemplaires.</p><p>- P. N. du W, IX 2003, nombreux exemplaires.</p><p>- [Boucomont, 1933: Haut Dahomey].</p></div>	https://treatment.plazi.org/id/857E879E8E32965FFC1EFB68FB74FADF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E32965FFF6BF9F8FD34F7F2.text	857E879E8E32965FFF6BF9F8FD34F7F2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Kheper festivus subsp. festivus (von Harold 1868)	<div><p>Kheper festivus festivus (von Harold, 1868) (Pl. 1, Fig. 3)</p><p>- Abomey-Calavi, Togba, 15 V 2002, 1 ex.; VIII 2003, 1 ex.</p><p>- Cové Toué, 19 VI 2018, 1 ex.</p><p>- Lokossa, XI 2015, à vue, 2 ex.</p><p>- Attogon, Tokopa, 30 V 2016, 4 ex., sous excrément de zébu.</p><p>- Attogon, forêt de Niaouli, 18 XI 2015, 15 ex., sous excrément de zébu, 10 VII 2016, sous excrément de zébu, 5 ex.</p><p>- Houéyogbé, 06 VII 2009, sous excrément humain, 1 ex.</p><p>- [Janssens, 1940: Dahomey. Moretto, 1998: plateau de Zagnanado et Ketou, région forestière des Hollis d’Adia - Ouéré Illémou; Porto Novo]</p><p>*  Kheper subaeneus (von Harold, 1869) (Pl. 1, Fig. 4)</p><p>- Houéyogbé, 24 V 2009,piège au sol, 1 ex.</p><p>- Pénéssoulou,forêt de Goutansoukpa, 15 VIII 2011, 1 ex., sous excrément de zébu.</p><p>- P.N. Pendjari, VI 2005,commun dans les crottins d’éléphants,nombreux spécimens.</p></div>	https://treatment.plazi.org/id/857E879E8E32965FFF6BF9F8FD34F7F2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E32965FFC61FF0FFA7EFE80.text	857E879E8E32965FFC61FF0FFA7EFE80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Parateuchus palemo (Olivier 1789)	<div><p>Parateuchus palemo (Olivier, 1789)</p><p>(Pl. 2, Fig. 1 &amp; Pl. 73, Fig. 1)</p><p>- [Moretto, 2016: plateau de Zagnanado, 1910; Kétou, 1900; cercle de Djougou-Kouandé, 1908; Tanguiéta-Kosso, 15 V 1996; Kétou].</p></div>	https://treatment.plazi.org/id/857E879E8E32965FFC61FF0FFA7EFE80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E32965FFC6BFE92FC1FFD97.text	857E879E8E32965FFC6BFE92FC1FFD97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Scarabaeus goryi (de Castelnau 1840)	<div><p>Scarabaeus goryi (de Castelnau, 1840)</p><p>(Pl. 2, Fig. 2-3 &amp; Pl. 73, Fig. 2)</p><p>-   P. N. du W, IX 2003, 1 ex., piège lumineux,  Tapoa Djerma à moins d’ 1 km de la mare aux Lions, côté Burkina Faso  .</p><p>- Houéyogbé, forêt de Houéyogbé, VII 2013, 2 ex., ultra-violet.</p><p>- [Moretto &amp; Deschamps, 2023: Batran [Baffan]; Savalou, Dassa Zoumé; Savé].</p></div>	https://treatment.plazi.org/id/857E879E8E32965FFC6BFE92FC1FFD97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E32965FFC6BFD9FFBDEFD50.text	857E879E8E32965FFC6BFD9FFBDEFD50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Scarabaeus isidis (de Castelnau 1840)	<div><p>Scarabaeus isidis (de Castelnau, 1840)</p><p>(Pl. 2, Fig. 4)</p><p>- P.N.duW, Mamassi-Gourma, 29 IX 2015, lumière, 1 ex.</p><p>- [Moretto &amp; Deschamps, 2023: Malanville; Haut Dahomey, entre Parakou et Djougou].</p></div>	https://treatment.plazi.org/id/857E879E8E32965FFC6BFD9FFBDEFD50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E32965EFC16FA48FEB0FEE0.text	857E879E8E32965EFC16FA48FEB0FEE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Chalconotus cupreus (Fabricius 1775)	<div><p>Chalconotus cupreus (Fabricius, 1775)</p><p>(Pl. 3, Fig. 3)</p><p>- Abomey-Calavi, Togba, VIII 2003, sous excrément, 1 ex.</p><p>- Houéyogbé,forêtdeHouéyogbé, 24V 2009,piège ausol, 4 ex.; 06 VII 2009,lumière, 3 ex.; 18 XI 2010,lumière, 2 ex.; 17 VIII  2011,U.V., 7 ex.; 25IX 2012,lumière, 2 ex.</p><p>- Lokossa, XI 2015, lumière, 1 ex.</p><p>- Allada,Tokpota, 24 IX 2013,sous excrément, 1 ex.; 30 V 2016,sous excrément, 1 ex.</p><p>- Attogon,forêtde Niaouli, 25 V 2016, lumière, 1 ex.; 26 V 2016, au sol, 1 ex.; 29 X 2014,au sol, 1 ex.;  18 XI 2015,U. V., 3 ex.; 07VI 2016,au sol, 2 ex.;  X 2016,U. V., 4 ex.</p><p>- Pobé, forêt de Pobé, 26 X 2014, au sol, 1 ex.</p><p>- Dan, bord du Zou,  13 IV 2012, U. V., 1 ex.; 07 V  2013, U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, VIII 2011, lumière, 5 ex.; 21 X 2011, lumière, 1 ex.; 29 X 2011,lumière, 1 ex.</p><p>- Pénéssoulou, centre ville, lumière, 13 VI 2013, 2 ex.; 28 VI 2013,lumière, 1 ex.</p><p>- Pénéssoulou,forêtde Pénélan, 14V  2013,U.V., 1 ex.; 26V  2013,U.V., 2 ex.; 26 V 2013,sousexcrément humain, 1 ex.; 27V  2013,U.V., 3 ex.; 03VII  2013,U.V., 1 ex.; 07 VIII  2013,U.V., 1 ex.;  08 IX 2013,U. V., 3 ex.; 16 XI 2013, sous excrément humain, 3 ex.; 16 VI  2016,U.V., 3 ex.</p><p>-  Kouandé, antenne ORTB, 05 XI 2015,  U. V., 2 ex.</p><p>- P. N. de la Pendjari, Batia, 26 VIII 2010, lumière, 2 ex.</p><p>- [Boucomont, 1933: Ouidah: plateau de Zagnando et Ketou, rég. forestière desHollis, 1910; cercle de Djongou. Janssens, 1938: Dahomey. Boucomont, 1933 (sub forme H): cercle de Djougou Kouandé].</p><p>*  Chalconotus suturalis (Janssens, 1938) (Pl. 3, Fig. 4)</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V., 23 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 17 VI 2013, sous excrément humain, 1 ex.; 2014, sous excrément humain, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E32965EFC16FA48FEB0FEE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E33965EFC7BFE4FFC77FD40.text	857E879E8E33965EFC7BFE4FFC77FD40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Garreta laetus subsp. laetus (Hope 1842)	<div><p>Garreta laetus laetus (Hope, 1842)</p><p>(Pl. 5, Fig. 1)</p><p>- Attogon, forêt de Niaouli, 04 V 2013, au sol, 1 ex.; 22 II 2014, au sol, 1 ex.</p><p>- Zogbodomey, forêt de la Lama, 11 VI 2009, sous excrément humain, 9 ex.; 08 VII 2009, sous excrément, 2 ex.; 05 VIII 2009, sous excrément humain, 9 ex.; 05 VIII 2009, sous excrément humain, 1 ex.; 08 VIII 2009, à vue, 2 ex.</p><p>- Pobé, forêt de Pobé, 29 VI 2013, lumière, 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 12 V 2013, sous excrément divers, 1 ex.; 02 VII 2013, sous excrément, 2 ex.; 04 VII 2013, lumière, 3 ex.;  2014, U. V., 2 ex.; 16 VI  2016, U.V., 1 ex.</p><p>- Pénéssoulou, centre ville, 17 VI 2013, lumière, 1 ex.; 28 VI 2013,lumière, 1 ex.</p><p>- [Moretto &amp; Génier, 2015: Gourma &amp; Mossi; Ketou; Reg. De Zagnanado].</p></div>	https://treatment.plazi.org/id/857E879E8E33965EFC7BFE4FFC77FD40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E33965EFF69FE4CFBE9FEE7.text	857E879E8E33965EFF69FE4CFBE9FEE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gymnopleurini Lacordaire 1856	<div><p>GYMNOPLEURINI Lacordaire, 1856</p><p>*  Allogymnopleurus aeneus (von Harold, 1875) (Pl. 4, Fig. 1)</p><p>-  Kouandé, 05 XI 2015, U. V., 1 ex.</p><p>- P. N. de la Pendjari, Batia, 27 VII 2009, piège au sol, 5 ex.; 18-25 VIII 2010, piège au sol, 11 ex.; 26 VIII 2010, piège excrément, 19 ex.</p><p>- P. N. Pendjari, VI 2005, quelques spécimens, bouse de zébu.</p><p>* A llogymnopleurus  olivieri (de Castelnau, 1840) (Pl. 4, Fig. 2 &amp; Pl. 73, Fig. 3)</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, piège lumineux et dans les bouses de buffles.</p><p>*  Allogymnopleurus umbrinus (Gerstaecker, 1871) (Pl. 4, Fig. 3 &amp; Pl. 73, Fig. 4)</p><p>- P.N.Pendjari, VI 2005, plusieurs spécimens sous bouses de zébus et de buffles.</p><p>*  Allogymnopleurus youngai Endrödi, 1973 (Pl. 4, Fig. 4)</p><p>- P.N. dela Pendjari,Batia, 24 VII 2009,piège au sol, 2 ex.; 27VII 2009, piège au sol, 2 ex.; 29 VII 2009, piège au sol, 3 ex.</p><p>- P. N. Pendjari, Batia, VI 2004, bouse de zébu, nombreux exemplaires; 18 VIII 2010, lumière, 1 ex.</p><p>- P.N.Pendjari, VI 2005, plusieurs exemplaires dans crottins d’éléphants.</p><p>- P.N. duW,pistedeKérémou-Koudou, IX 2003,nombreux exemplaires, bouse de zébu. Les deux formes cohabitent dans le parc de la Pendjari: thorax rouge, élytres bleus et thorax vert, élytres bleus.</p></div>	https://treatment.plazi.org/id/857E879E8E33965EFF69FE4CFBE9FEE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E339651FC47FCD2FE98F8EA.text	857E879E8E339651FC47FCD2FE98F8EA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Garreta nitens (Olivier 1789)	<div><p>Garreta nitens (Olivier, 1789)</p><p>(Pl. 5, Fig. 2)</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013,sous bouse zébu sèche, 4 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 19 X 2011, sous excrément, 2 ex.; 30 V 2012, 1 ex.</p><p>- Pénéssoulou, forêt de Pénéssoulou, 12 V 2013, sous excréments divers, 1 ex.; 26 V 2013, sous excréments divers, 1 ex.</p><p>- P. N. de la Pendjari, Batia, 24 VII 2009, piège excrément, 4 ex.; 27 VII 2009, piège excrément, 1 ex.; 18 VIII 2010, piège au sol, 1 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, nombreux exemplaires, sous bouses de buffles et crottins d’éléphants.</p><p>- P. N. du W, IX 2003, nombreux exemplaires sous bouses de zébus et crottins d’éléphants.</p><p>- [Moretto &amp; Génier, 2015: Kosso; Tanguiéta; Mts Kouffè;Alafiarou: Ft de WariMaro].</p><p>*  Gymnopleurus coerulescens (Olivier, 1789) (Pl. 5, Fig. 3)</p><p>- Bohicon, 15 VI 2014, lumière, 1 ex.</p><p>-   Dan, bord du Zou, 13 IV 2012, sous excrément, 23 ex. ; 17 VII 2013, sous excrément, 2 ex.</p><p>- Attogon, forêt de Niaouli, 15 X 2016, sous excrément, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu sèche, 1 ex.</p><p>- P. N. de la Pendjari, VI 2005, 1 ex.</p><p>*  Gymnopleurus fulgidus (Olivier, 1789)</p><p>(Pl. 5, Fig. 4)</p><p>- Lokossa, 24 III 2006, 2 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 14 II 2020, 2 ex., U. V .</p><p>-  Kouandé, antenne ORTB, 05 XI 2015, sous excrément, 2 ex.</p><p>- P. N. Pendjari, 07 XI 2012, mare Bali, sous bouse de buffle, 1 ex.</p><p>- P. N. Pendjari, Batia, 24 VII 2009, sous excrément humain, 1 ex., 29 VII 2009, sous excrément humain, 9 ex.; 18 VIII 2010, sous excrément humain, 3 ex.; 08 XI 2012, sous excrément humain, 29 ex.; 11 XI 2012, sous excrément humain, 20 ex.; 08 XI 2015, sous bouse, 29 ex.; 09 XI 2015, sous excrément, 5 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, nombreux exemplaires.</p><p>- Malanville, 01 X 2015, lumière, 4 ex.</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 2 ex.</p><p>- P. N. du W, IX 2003, plusieurs exemplaires dans divers excréments.</p><p>*  Gymnopleurus nyankpalaensis Endrödi, 1976 (Pl. 6, Fig. 1)</p><p>-  Kouandé, antenne ORTB, 06 XI 2015, lumière, 1 ex.</p><p>- P. N. de la Pendjari, Batia, 18-25 VIII 2010, piège excrément, 10 ex.; 26 VIII 2010, piège excrément, 5 ex.</p><p>- P. N. Pendjari,Tanougou, VI 2005, plusieurs exemplaires dans excréments de Babouins.</p><p>- P. N. du W, chutes de Koudou, IX 2003, plusieurs exemplaires dans excrément humain.</p><p>*  Gymnopleurus profanus (Fabricius, 1792)</p><p>(Pl. 6, Fig. 2)</p><p>- Attogon, forêt de Niaouli, 2014, piège fruit, 1 ex.</p><p>-  P. N. du W, Tapoa Djerma (côté Burkina Faso), IX 2003, 1 ex. sous excrément humain .</p><p>*  Gymnopleurus puncticollis Gillet, 1909</p><p>(Pl. 6, Fig. 3)</p><p>- P. N. de la Pendjari, Batia, 27 VII 2009, sous excrément, 6 ex.; 18 VIII 2010, piège excrément, 3 ex.</p><p>- VersTanougou, VI 2005, nombreux exemplaires sous excrément de Babouins et de Mustelidés.</p><p>- P.N.duW, chaînedel’Atakora, VI 2005,nombreux exemplaires sous excrément de babouins et bouses de zébus.</p></div>	https://treatment.plazi.org/id/857E879E8E339651FC47FCD2FE98F8EA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3C9651FEADF8B9FA40FE1A.text	857E879E8E3C9651FEADF8B9FA40FE1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphini Mulsant 1842	<div><p>SISYPHINI Mulsant, 1842</p><p>*  Sisyphus (Neosisyphus) armatus Gory, 1833</p><p>(Pl. 6, Fig. 4)</p><p>- P. N. de la Pendjari, Batia, 24 VIII 2010, pièges crevettes, 3 ex.; 20-26 VIII 2010, piège au sol, 2 ex.</p><p>*  Sisyphus (Neosisyphus) baoule (Cambefort, 1984) (Pl. 7, Fig. 1)</p><p>-  P. N. de la Pendjari, Batia, 07 XI 2015, U. V., 2 ex.</p><p>*  Sisyphus (Neosisyphus) gladiator Arrow, 1927</p><p>(Pl. 7, Fig. 2-3 &amp; Pl. 73, Fig. 5-6)</p><p>- P. N. Pendjari, VI 2005, crottins d’éléphants, nombreux exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E3C9651FEADF8B9FA40FE1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3C9651FCF1FE0AFA76FD68.text	857E879E8E3C9651FCF1FE0AFA76FD68.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Neosisyphus) paschalidisae (Cambefort 1984)	<div><p>Sisyphus (Neosisyphus) paschalidisae (Cambefort, 1984) (Pl. 7, Fig. 4)</p><p>- Savalou, forêt de Logozohé, 06 VIII 2007, sous bouse zébu, 4 ex.</p><p>- P.N.Pendjari, VI 2006,nombreux exemplaires dans les crottins d’éléphants.</p><p>- P. N. de la Pendjari, Batia, 20-26 VIII 2010, dans excrément, 15 ex.; 07 XI 2015, sous bouse zébu, 2 ex.</p><p>- P. N. du W, réserve de la Djonga, près de la mare24, VI 2005, nombreux exemplaires dans les crottins d’éléphants.</p><p>- [Montreuil, 2015: rés.Djona, excr.Éléphant; Batia; forêt de Logozohe].</p></div>	https://treatment.plazi.org/id/857E879E8E3C9651FCF1FE0AFA76FD68	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3C9651FC24FD3AFBA3FCE8.text	857E879E8E3C9651FC24FD3AFBA3FCE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Neosisyphus) youngai Endrodi 1973	<div><p>Sisyphus (Neosisyphus) youngai Endrödi, 1973</p><p>(Pl. 8, Fig. 1-2 &amp; Pl. 73, Fig. 7-8)</p><p>- [Montreuil, 2015: Pobé Ouémé].</p></div>	https://treatment.plazi.org/id/857E879E8E3C9651FC24FD3AFBA3FCE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3C9651FC01F880FA50F792.text	857E879E8E3C9651FC01F880FA50F792.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Parasisyphus) Barbero, Palestrini & Zunino 1991	<div><p>Sisyphus (Parasisyphus) complexe  goryi (Pl. 9, Fig. 1)</p><p>- Lokossa, 01 III 2006, 2 ex.</p><p>- Houéyogbé, 22 VIII  2011, 1 ex.; VII  2013, 1 ex., U.V.;  23 II 2020, U. V .</p><p>- Pénéssoulou, zone humide, sortie du village, 1 ex., sous bouses de zébus.</p><p>Spécimens en cours d’étude, dans le cadre d’une révision du groupe. [Hautier et al., 2003: Ina, sub  Sisyphus crispatus Gory, 1833].</p></div>	https://treatment.plazi.org/id/857E879E8E3C9651FC01F880FA50F792	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3C9651FCF3FCB8FA0BFA62.text	857E879E8E3C9651FCF3FCB8FA0BFA62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Sisyphus) africanus subsp. africanus Montreuil 2017	<div><p>Sisyphus (Sisyphus) africanus africanus Montreuil, 2017 (Pl. 8, Fig. 3)</p><p>- Lokossa, 01 III 2006, 1 ex.</p><p>- Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, piège excrément, 3 ex.</p><p>- Allada, Donou, 10 VI 2016, 5 ex.</p><p>-  Attogon, forêt de Niaouli, 24 X 2019, 1 ex., U. V .</p><p>-  Zogbodomey, forêt de la Lama, 27 V 2018, 2 ex., U. V .</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013, sous bouse zébu, 8 ex.; 25 V 2013, sous bouse zébu, 10 ex. paratypes.</p><p>- Pénéssoulou, forêt de Pénélan,  03 XII 2014, U. V., 1 ex.; 16 VI  2016, U.V., 10 ex.</p><p>- Pénéssoulou, zone humide, sortie du village, 23 II 2020, 13 ex., sous bouses de zébus.</p><p>-  Kouandé, antenne ORTB, 05 XI 2015,  U. V., 3 ex.</p><p>- P. N. Pendjari, Batia, Camp Numi, 09 XI 2015, piège poisson, 1 ex.</p><p>- P. N. Pendjari, Hôtel Pendjari, 26 VIII 2010, dans excrément, 21 ex.</p><p>- P.N.Pendjari, VI 2005, nombreux exemplaires sur crottins d’éléphants.</p><p>- [Montreuil, 2017: P. N. Pendjari, Batia, entrée du Parc, piège crevettes; Pénéssoulou, forêt de Saramanga, sous bouse zébu; Kandi,Alfakouara,Djona].</p></div>	https://treatment.plazi.org/id/857E879E8E3C9651FCF3FCB8FA0BFA62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3C9651FC0FFA37FBA7F890.text	857E879E8E3C9651FC0FFA37FBA7F890.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Sisyphus) arboreus Walter 1982	<div><p>Sisyphus (Sisyphus) arboreus Walter, 1982</p><p>(Pl. 8, Fig. 4)</p><p>- Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, piège excrément, 13 ex.</p><p>-  Attogon, forêt de Niaouli, 12 XII 2011, U. V., 1 ex.</p><p>- Attogon, Niaouli,  29 IV 2012, U. V., 3 ex.; 02 V  2013, U.V., 16 ex.; 31 X 2014, fauchage, 2 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 29 X 2011, lumière, 3 ex.</p><p>- P. N. de la Pendjari, Batia, 20-26 VIII 2010, piège au sol, 3 ex.</p><p>- [Montreuil, 2016: Djigbé Forêt; Goutansoukpa, lumière; Attogon, forêt de Niaouli; Pendjari, Batia].</p></div>	https://treatment.plazi.org/id/857E879E8E3C9651FC0FFA37FBA7F890	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3D9650FEB4FF0BFDC8FE07.text	857E879E8E3D9650FEB4FF0BFDC8FE07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Coprini Leach 1815	<div><p>COPRINI Leach, 1815</p><p>sensu lato</p><p>*  Catharsius achates (Olivier, 1789) (Pl. 9, Fig. 2)</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 3 ex.</p><p>- P. N. du W, Mamassi-Gourma, 29 IX 2015, lumière, 7 ex.</p><p>- Zogbodomey, Domé, 13 XII 2014, lumière, 4 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E3D9650FEB4FF0BFDC8FE07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3D9653FF71FE2EFF2EFD3A.text	857E879E8E3D9653FF71FE2EFF2EFD3A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Catharsius eteocles (de Castelnau 1840)	<div><p>Catharsius eteocles (de Castelnau, 1840)</p><p>(Pl. 9, Fig. 3 &amp; Pl. 73, Fig. 9)</p><p>-  Bohicon, quartier Soglogon, 28 IV 2013, U. V., 1 ex.</p><p>- P. N. Pendjari, Batia, 18-24 VIII 2010, piège au sol, 1 ex.; 24 VIII 2010, lumière, 3 ex.</p><p>- P.N.duW,TapoaDjerma(côtéBurkinaFaso), IX 2003,plusieurs exemplaires.</p><p>- [Hautier et al., 2003: Ina].</p><p>*  Catharsius guineensis Ferreira, 1963 (Pl. 9, Fig. 4; Pl. 10, Fig. 1 &amp; Pl. 73, Fig. 10)</p><p>-  Attogon, forêt de Niaouli, 06 IV 2012, U. V., 3 ex.</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, lumière, 1 ex.;  18 IV 2012, U. V., 1 ex.;  20 IV 2012, U. V., 2 ex.;  29 IV 2012, U. V., 2 ex.; 24 V  2012, U.V., 2 ex.; 06 VI  2016, U.V., 3 ex.;  01 XII 2016, U. V., 2 ex.; 08 VI  2018, U.V., 2 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse zébu sèche, 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 08 IX 2013, 1 ex.</p><p>- P. N. Pendjari, Batia, 24 VII 2009, piège au sol, 2 ex.; 26 VIII 2010, piège au sol, 1 ex.;  07 XI 2015, U. V., 1 ex.;  09 XI 2015, U. V., 1 ex.</p><p>- P. N. du W, Mamassi-Gourma, 29 IX 2015, lumière, 2 ex.</p><p>*  Catharsius complexe polynices Kolbe, 1893 (Pl. 10, Fig. 2-3)</p><p>-   Dan, bord du Zou, 13 IV 2012, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 21 X 2011, sous excrément, 15 ex.; 29 X 2011, sous excrément, 6 ex.</p><p>- Goutansoulpa, 19 X 2011, lumière, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013,sous bouse zébu sèche, 1 ex.</p><p>-  Pénéssoulou, stade de foot, 18 X 2012, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 17 VI 2013, sous excrément, 2 ex.; 15 V  2013, U.V., 1 ex.; 18 V  2013, U.V., 1 ex.;  11 X 2012, U. V., 1 ex.; VIII  2013, U.V., 4 ex.;  08 IX 2013, U. V., 5 ex.; 2014, à vue, 2 ex.</p><p>*  Catharsius pseudolycaon Ferreira, 1960</p><p>(Pl. 10, Fig. 4; Pl. 11, Fig. 1 &amp; Pl. 73, Fig. 11)</p><p>-  Pénéssoulou, forêt de Pénélan, 02 VII 2013, U. V., 1 ex.</p><p>-  Kouandé, 05 XI 2015, U. V., 6 ex.</p><p>-  P. N. Pendjari, Batia, 07 XI 2015, U. V., 1 ex.</p><p>- P. N. du W, Point Triple, IX 2003, plusieurs exemplaires dans excrémens humains;</p><p>-  Tanguiéta, chaîne de l’Atakora, nord du Bénin, VI 2005, sous excrément humain .</p><p>*  Catharsius complexe sesostris Waterhouse, 1891 (Pl. 11, Fig. 2-3 &amp; Pl. 73, Fig. 12)</p><p>- Houéyogbé, forêt de Houéyogbé, III 2012, à vue, 2 ex.; 2010,au sol, 1 ex.; 20 IX 2011, sur excrément, 1 ex.;  20 X 2014, U. V., 1 ex.; 25 VI  2016, U.V., 1 ex.</p><p>- Lokossa, bois de la Sitex, 2008, sous excrément, 1 ex.</p><p>-  Attogon, forêt de Niaouli, 02 IV 2013, U. V., 1 ex.; 2014, sur exrément, 2 ex.</p><p>-  Bohicon, quartier Soglogon, 18 IV 2018, U. V., 2 ex.;  20 IV 2012, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, VIII 2011, lumière, 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 28 V  2013, U.V., 1 ex.; 17 VI 2013, sous excrément, 1 ex.;  08 IX 2013, U. V., 3 ex.; 16 VI  2016, U.V., 1 ex.</p><p>- Pénéssoulou, centre ville, 17 VI 2013, lumière, 1 ex.; 28 VI 2013, lumière, 1 ex.</p><p>-  Djougou, forêt de Sérou, 10 XI 2015, U. V., 1 ex.</p><p>- Ndali, forêt de Ndali, Bas fond, 11 XI 2015, sous bouse zébu, 1 ex.</p><p>- Kouandé, antenne  ORTB, 05 XI 2015, U.  V., 1 ex.; 06 XI 2015, U.  V., 1 ex.</p><p>- P. N. Pendjari, Batia, 24 VII 2009, piège au sol, 3 ex.</p><p>- P. N. Pendjari, forêt de la Boundjagou, VI 2005, plusieurs exemplaires sous ecréments humains.</p><p>- P. N. du W, Alpha Kouara, IX 2003, plusieurs exemplaires.</p><p>- P.N. delaPendjari, réserve delaDjona, VI 2005, plusieurs exemplaires sous ecréments humains.</p><p>-   P. N. Pendjari,  Pont de l’Alibori, VI 2005, plusieurs exemplaires, sous excrément humain  .</p></div>	https://treatment.plazi.org/id/857E879E8E3D9653FF71FE2EFF2EFD3A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3E9653FF37FD6AFD81FCB8.text	857E879E8E3E9653FF37FD6AFD81FCB8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris angustus Nguyen-Phung & Cambefort 1986	<div><p>Copris angustus Nguyen-Phung &amp; Cambefort, 1986 (Pl. 11, Fig. 4; Pl. 12, Fig. 1 &amp; Pl. 74, Fig. 1-2)</p><p>- [Nguyen-Phung &amp; Canbefort, 1986: Haut Dahomey].</p></div>	https://treatment.plazi.org/id/857E879E8E3E9653FF37FD6AFD81FCB8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3E9652FF5AFCEAFBD3FE4A.text	857E879E8E3E9652FF5AFCEAFBD3FE4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris carmelita (Fabricius 1801)	<div><p>Copris carmelita (Fabricius, 1801)</p><p>(Pl. 12, Fig. 2)</p><p>- Attogon, forêt de Niaouli,  01 IV 2013, U. V., 2 ex.;  02 IV 2012, U. V., 1 ex.;  02 II 2017, U. V., 1 ex.</p><p>- Zogbodomey, forêt de la Lama,  31 I 2017,U. V., 2 ex.; 3 VI  2013, U.V. 1 ex.;  12 XI 2015, U. V., 1 ex.</p><p>-  Djougou, forêt de Sérou, 01 X 2019, U. V., 1 ex.</p><p>-  Pénéssoulou, foret de Pénélan, 26 XI 2019, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 21 X 2011, sous excrément, 1 ex.; 29 X 2011, sous excrément, 1 ex.;</p><p>- P. N. Pendjari, hôtel Pendjari, 26 VIII 2010, sous bouse buffle, 1 ex.</p><p>- [Ferreira, 1961: Porto-Novo. La citation très surprenante de Porto-Novo de  Copris harrisi Waterhouse, 1891, espèce d’Afrique orientale, dans cette même publication, correspond vraissemblablement à une confusion avec  Copris carmelita].</p><p>*  Copris coriarius Gillet, 1907</p><p>(Pl. 12, Fig. 3-4 &amp; Pl. 74, Fig. 3)</p><p>- Pénéssoulou, forêt de Pénéssoulou, 12 V 2013, sous excrément, 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 08 IX 2013, U. V., 2 ex.</p><p>-  P. N. Pendjari, Batia, Camp Numi, 07 XI 2012, U. V., 2 ex.;</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, 1 ex., piège lumineux.</p><p>*  Copris gazellarum Gillet, 1918</p><p>(Pl. 13, Fig. 1)</p><p>-  Kouandé, antenne ORTB, 05 X 2019,  U. V., 2 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013,sous bouse de zébu sèche, 3 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, VIII 2011, sous excrément, 2 ex.</p><p>- Pénéssoulou, centre ville, 17 VI 2013, lumière, 1 ex.;  forêt de Pénélan, 08 VIII 2013, U. V., 1 ex.</p><p>*  Copris interioris Kolbe, 1897 (Pl. 13, Fig. 2-3 &amp; Pl. 74, Fig. 4-5)</p><p>- Houéyogbé, forêt de Houéyogbé, 24 V 2009, sous excrément, 1 ex.</p><p>-  Attogon, forêt de Niaouli, 01 IV 2013, U. V., 1 ex.; 02V  2013, U.V., 1 ex.; 2014, sous excrément, 1 ex.</p><p>-  Zogbodomey, forêt de la Lama, 31 I 2017, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, lumière, 2 ex.;  14 IV 2012, U. V., 2 ex.;  20 IV 2012, U. V., 1 ex.; 11 V  2013, U.V., 1 ex.;  IV 2016, U. V., 1 ex.; 06 VI  2016, U.V., 1 ex.;  17 XI 2017, U. V., 2 ex.; 03 VI  2018, U.V., 12 ex.; 08 VI  2018, U.V., 5 ex.; 16 VI  2018, U.V., 1 ex.; 17 V  2018, U.V., 2 ex.</p><p>- Dan, bord du Zou,  13 IV 2012, U. V., 2 ex.; 04 VI  2013, U.V., 3 ex.</p><p>- Savalou, forêt de Logozohé, 06 VIII 2007, sous excrément de zébu, 8 ex.; 04 IX 2007, sous excrément de zébu, 8 ex.</p><p>-  Kouandé, antenne ORTB, 05 X 2019,  U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, VIII 2011, lumière, 2 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013, sous bouse zébu, 1 ex.; 25 V 2013, sous bouse de zébu sèche, 15 ex.</p><p>- Pénéssoulou, centre ville; 28VI 2013, lumière, 2 ex.; 17VI 2013,lumière, 3 ex.; 22 V 2013, lumière, 3 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 15 V  2012, U.V., 1 ex.;  11 X 2012, U. V., 1 ex.; 23 V  2013, U.V., 2 ex.; 27 V  2013, U.V., 17 ex.; 28 V  2013, U.V., 2 ex.; 02 VII  2013, U.V., 8 ex.; 03 VII  2013, U.V., 1 ex.; 04 VII  2013, U.V., 1 ex.; 30 VII  2013, U.V., 3 ex.;  08 IX 2013, U. V., 8 ex.; 2014, lumière, 5 ex.; 16 VI  2016, U.V., 4 ex.</p><p>- Pénéssoulou, forêt de Pénéssoulou, 12 V 2013, sous excrément divers, 1 ex. - Ndali, forêt de Ndali, 11 XI 2015, sous bouse de zébu, 5 ex.</p><p>- Kouandé, antenne ORTB;  04 XI 2015, U. V., 1 ex.;  05 XI 2015, U. V., 2 ex. -   P. N. Pendjari,  Batia, 27 VII 2009, piège au sol, 1 ex.; 18-25 VIII 2010, piège au sol, 7 ex.; 26 VIII 2010, piège excrément, 5 ex.;  07 XI 2015, U. V., 1 ex.;  08 XI 2015, U. V., 2 ex.;  09 XI 2015, U. V., 1 ex.</p><p>- P.N.Pendjari,Batia,Camp Numi,  10 XI 2012, U. V., 1 ex.;  07 XI 2015, U. V., 2 ex.;  03 XI 2015, U. V., 1 ex.</p><p>- P. N. Pendjari, forêt de Bondjagou, VI 2005, sous excrément humain; 26 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P.N.Pendjari,versTanougou, VI 2005,plusieurs exemplaires sous bouses de zébus.</p><p>- P.N.du W,AlphaKouara,prèsdelamare, 24IX 2003,plusieursexemplaires sous excrément humain.</p><p>- P. N. du W, Tapoa Djerma, marre aux Lions (côté Burkina Faso), IX 2003, sous crottins d’éléphants.</p><p>*  Copris laius laius von Harold, 1868 (Pl. 13, Fig. 4; Pl. 14, Fig. 1 &amp; Pl. 74, Fig. 6-7)</p><p>- Pénéssoulou, forêt de Goutansoukpa, VIII 2011, au sol, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu sèche, 5 ex. -   Pénéssoulou, forêt de Pénélan , 15 V  2013, U.V., 3 ex.; 18 V  2013, U.V., 3 ex.; 23 V  2013, U.V., 1 ex.; 28 V  2013, U.V., 1 ex.; 17 VI 2013, sous excrément, 3 ex.;  08 IX 2013, U. V., 2 ex.</p><p>-  Pénéssoulou, centre ville, 22 V 2013, U. V., 1 ex.; 17 VI 2013, lumière, 5 ex.</p><p>- P. N. Pendjari, Batia, 18-25 VIII 2010, sous excrément, 3 ex.; 24 VIII 2010, sous excrément, 9 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, 28  VIII 2010, piège excrément, 1 ex.; VI 2005, plusieurs exemplaires, piège lumineux.</p><p>- P. N. du W, Alpha Kouara, réserve de la Djonga, IX 2003, plusieurs exemplaires.</p><p>*  Copris megaceratoides Waterhouse, 1891</p><p>(Pl. 14, Fig. 2-3 &amp; Pl. 74, Fig. 8-9)</p><p>-  Bohicon, quartier Soglogon, 20 IV 2012, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V., 2 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 29 X 2011, sous excrément, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013, sous bouse zébu, 3 ex; 24 VI 2013, sous bouse de zébu, 2 ex.; 25 V 2013, sous bouse de zébu, 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 15 V  2013, U.V., 1 ex.; 18 V  2013, U.V., 1 ex.; 27 V  2013, U.V., 1 ex.; 02 VII  2013, U.V., 2 ex.; 04 VII  2013, U.V., 1 ex.</p><p>-  Djougou, forêt de Sérou, 10 XI 2015, U. V., 1 ex.</p><p>-  P. N. Pendjari, Batia, Camp Numi, 11 XI 2012, U. V., 2 ex.</p><p>- P. N. du W, réserve de la Djona, Près de la mare, 24, IX 2003, sur crottins d’éléphants.</p><p>- P.N.duW,pisteKoudouaupointTriples, IX 2003, dans les bouses de buffles.</p><p>*  Copris moffartsi Gillet, 1907 (Pl. 14, Fig. 4; Pl. 15, Fig. 1 &amp; Pl. 74, Fig. 10-11)</p><p>- Pénéssoulou, forêt de Pénélan,  08 IX 2013, U. V., 3 ex.; 02 VII  2013, U.V., 3 ex.; 04 VII  2013, U.V., 2 ex.;  24 I 2017, U. V., 1 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 07 XI 2012, U. V., 1 ex.</p><p>*  Copris orion orion Klug, 1835 (Pl. 15, Fig. 2 -3 &amp; Pl. 74 &amp; Fig. 12)</p><p>-  Pobé, forêt de Pobé, 21 XII 2014, U. V., 1 ex.</p><p>- Bohicon, 15 VI 2009, lumière, 1 ex.; 03 VI  2018, U.V., 1 ex.</p><p>- Dan, bord du Zou, 04 VI  2013, U.V., 1 ex.;  10 XII 2014, U. V., 2 ex.</p><p>- Savalou, forêt de Logozohé, 06 VIII 2007, sous excrément, 1 ex.</p><p>- Bassila, Tchétou, 17 VI 2016, bouse zébu, 1 ex.</p><p>-  Bétérou, bord de l’Ouémé, 27 I 2017, U. V., 4 ex.</p><p>-  Bassila, Manigri, 15 I 2017, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 19 X 2011, à vue, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013, sous excrément, 1 ex.; 25 V 2013, sous bouse de zébu, 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 27 V  2013, U.V., 3 ex.; 02 VII  2013, U.V., 3 ex.; 30 VII  2013, U.V., 1 ex.;  01 XII 2014, U. V., 1 ex.; 16 VI  2016, U.V., 1 ex.</p><p>- Pénéssoulou, centre ville, 17 VI 2013, lumière, 1 ex.</p><p>-  Djougou, forêt de Sérou, 10 XI 2015, U. V., 1 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 07 XI 2012, U. V., 1 ex.</p><p>*  Copris orphanus Guérin-Méneville, 1847</p><p>(Pl. 15, Fig. 4 &amp; Pl. 75, Fig. 1)</p><p>-  Pénéssoulou, forêt de Pénélan 10 XI 2014, U. V., 1 ex.</p><p>*  Copris sinon Olivier, 1789 (Pl. 16, Fig. 1 &amp; Pl. 75, Fig. 2)</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 6 ex.</p><p>*  Heliocopris antenor (Olivier, 1789) (Pl. 16, Fig. 2 &amp; Pl. 75, Fig. 3)</p><p>- Pénéssoulou, forêt de Goutansoukpa, 21 IX 2011, lumière, 1 ex.</p><p>- Pénéssoulou, centre-ville, VI 2013, lumière, 2 ex.; 2014, lumière, 4 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 1 ex.</p><p>-  Kouandé, antenne ORTB, 06 XI 2015,  U. V., 1 ex.</p><p>- P. N. du W, réserve de la Djona, IX 2003, plusieurs exemplaires, piège lumineux.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, plusieurs exemplaires au piège lumineux et sous crottins d’éléphants.</p></div>	https://treatment.plazi.org/id/857E879E8E3E9652FF5AFCEAFBD3FE4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3F9652FC3CFDD8FB06FD2A.text	857E879E8E3F9652FC3CFDD8FB06FD2A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Heliocopris dilloni Guerin-Meneville 1847	<div><p>Heliocopris cf. dilloni Guérin-Méneville, 1847</p><p>(Pl. 16, Fig. 3 &amp; Pl. 75, Fig. 4-5)</p><p>-   P. N du W,  Tapoa Djerma, mare aux Lions (côté Burkina Faso), IX 2003, plusieurs exemplaires, crottins d’éléphants  .</p></div>	https://treatment.plazi.org/id/857E879E8E3F9652FC3CFDD8FB06FD2A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3F9655FC64FD7FFD8DFEB7.text	857E879E8E3F9655FC64FD7FFD8DFEB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Heliocopris myrmidon Kolbe 1893	<div><p>Heliocopris myrmidon Kolbe, 1893</p><p>(Pl. 16, Fig. 4 &amp; Pl. 75, Fig. 6)</p><p>-  Pénéssoulou, forêt de Pénélan, 18 V 2013, U. V., 1 ex.</p><p>- [Coache et al., 2014: Pénéssoulou, forêt de Goutansoukpa, à la lumière; Pénéssoulou, forêt de Pénélan, U.V.; Pénéssoulou, village, lumière; forêt de Pénéssoulou, lumière].</p><p>*  Litocopris punctiventris Waterhouse, 1891 (Pl. 17, Fig. 1)</p><p>-  Attogon, forêt de Niaouli, 01 XII 2014, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan,  11 X 2012, U. V., 1 ex.; 23 V  2013, U.V., 2 ex.; 27 V  2013, U.V., 3 ex.; 17 VI  2013, U.V., 1 ex.; 02 VII  2013, U.V., 1 ex.;</p><p>03 VII  2013, U.V., 1 ex.;  08 IX 2013, U. V., 1 ex.;  01 XII 2014, U. V., 1 ex.; 16 VI  2016, U.V., 2 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 07 XI 2012, U. V., 2 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E3F9655FC64FD7FFD8DFEB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FC49F94BFB11F7A8.text	857E879E8E389655FC49F94BFB11F7A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitini de Castelnau 1840	<div><p>ONITINI de Castelnau, 1840</p><p>*  Heteronitis tridens (de Castelnau, 1840) (Pl. 20, Fig. 4; Pl. 21, Fig. 1 &amp; Pl. 75, Fig. 9-10)</p><p>- P. N. Pendjari, 7 km de la forêt de la Bondjagou, VI 2005, sous crottins d’éléphants.</p><p>*  Onitis aeneus van Lansberge, 1875</p><p>(Pl. 21, Fig. 2 &amp; 3)</p><p>- P. N. Pendjari, Batia, Camp Numi, 26 VIII 2010, lumière, 2 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, sous bouse éléphant, 15 ex.</p><p>- P. N. du W, piste Alpha-Kouara, mare 24, IX 2003, plusieurs exemplaires, dans les corttins d’éléphants et au piège lumineux.</p></div>	https://treatment.plazi.org/id/857E879E8E389655FC49F94BFB11F7A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FF6DFEFFFDFDFB30.text	857E879E8E389655FF6DFEFFFDFDFB30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Metacatharsius abortivus Fairmaire 1891	<div><p>Metacatharsius abortivus Fairmaire, 1891 (Pl. 17, Fig. 2)</p><p>- Ouidah, forêt de Pahou, 02 IV 2014, 2 ex.</p><p>- Houéyogbé, forêt de Houéyogbé,  23 IV 2013, U. V., 1 ex.; 08 V  2013, U.V., 1 ex.; 02 VI  2013, U.V., 1 ex.</p><p>-  Attogon, forêt de Niaouli, 01 IV 2013, U. V., 1 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 13 V 2013, U. V., 3 ex.</p><p>- Bohicon, quartier Soglogon,  20 IV 2012, U. V., 2 ex.;  28 IV 2012, U. V., 2 ex.;  28 IV 2013, U. V., 5 ex.; 01 V  2013, U.V., 1 ex.; 03 VI  2018, U.V., 2 ex.</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 27 V  2013, U.V., 2 ex.; 02 VII  2013, U.V., 1 ex.</p><p>- P.N.Pendjari, VI 2005,hôtelPendjari, piège lumineux, plusieurs exemplaires.</p><p>- [Boucomont, 1928: Dahomey].</p><p>*  Metacatharsius complexe inermis sp. 1 (avec tête inerme)</p><p>(Pl. 17, Fig. 3)</p><p>- Lokossa, bois de la Sitex, 2008, battage, 1 ex.</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, lumière, 1 ex.;  18 IV 2012, U. V., 1 ex.; 03 VI  2018, U.V., 1 ex.</p><p>-   Dan, bord du Zou, 13 IV 2012, U. V., 2 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 19 X 2011, lumière, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013, sous bouse de zébu, 1 ex.</p><p>* Metacarthasius complexe inermis sp. 2 (avec tubercule céphalique) (Pl. 17, Fig. 4)</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E389655FF6DFEFFFDFDFB30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FF02FB62FD52FB5F.text	857E879E8E389655FF02FB62FD52FB5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Metacatharsius parcepunctatus Balthasar 1965	<div><p>Metacatharsius parcepunctatus Balthasar, 1965</p><p>- [Hautier et al., 2003: Ina]. Présence possible, à confirmer.</p></div>	https://treatment.plazi.org/id/857E879E8E389655FF02FB62FD52FB5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FF76FAC7FE05F8FA.text	857E879E8E389655FF76FAC7FE05F8FA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Metacatharsius peleus (Olivier 1789)	<div><p>Metacatharsius peleus (Olivier, 1789) (Pl. 18, Fig. 1)</p><p>-  Attogon, forêt de Niaouli, 06 IV 2012, U. V., 1 ex.</p><p>- Bohicon, 03 VI  2018, U.V., 1 ex.</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 1 ex.</p><p>- P. N. du W, Mamassi-Gourma, 30 IX 2015, lumière, 2 ex.</p><p>- [Hautier et al., 2003: Ina].</p><p>*  Metacatharsius renaudi Frey, 1960 (Pl. 18, Fig. 2 &amp; 3)</p><p>- P. N. du W, réserve de la Djona, IX 2003, plusieurs exemplaires, piège lumineux.</p><p>*  Metacatharsius tschadensis Balthasar, 1961 ? (Pl. 18, Fig. 4)</p><p>-  Zinvié, vallée du Sitatunga, 19 V 2016, U. V., 1 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 13 V 2013, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V., 2 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E389655FF76FAC7FE05F8FA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FF75F86FFD18F79A.text	857E879E8E389655FF75F86FFD18F79A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria coprinarum Cambefort 1982	<div><p>Pedaria coprinarum Cambefort, 1982 (Pl. 19, Fig. 1)</p><p>- Bohicon, quartier Soglogon, 03 VI  2018, U.V., 3 ex.</p><p>- [Josso &amp; Prévost, 2015: environ de Cotonou; environ de Save].</p></div>	https://treatment.plazi.org/id/857E879E8E389655FF75F86FFD18F79A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FC6CFF0FFAEEFC32.text	857E879E8E389655FC6CFF0FFAEEFC32.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria criberrima Waterhouse 1890	<div><p>Pedaria criberrima Waterhouse, 1890 (Pl. 19, Fig. 2)</p><p>-   Athiémé, rive du Mono, 31 V  2013, U. V., 3 ex.</p><p>- Houéyogbé, forêt de Houéyogbé,  02 IV 2013, U. V., 3 ex.; 01 VI  2013, U.V., 1 ex.; 27 VIII  2014, U.V., 1 ex.</p><p>-  Zogbodomey, fokoli, 10 I 2019, U. V., 1 ex.</p><p>-  Bohicon, quartier Soglogon, 28 IV 2013, U. V., 4 ex.</p><p>- Cové, VII 2009, sous excrément, 1 ex.</p><p>- Savalou, forêt de Logozohé, 06 VIII 2007, sous bouse zébu, 1 ex.</p><p>- Pénéssoulou, 23 V  2013, U.V., 1ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 02 VII 2013, U. V., 2 ex.</p><p>- Pénéssoulou, centre-ville, 17 VI 2013, lumière, 5 ex.</p><p>-  Kouandé, antenne ORTB, 05 XI 2015,  U. V., 1 ex.</p><p>- P. N. Pendjari, mare Bali, 07 XI 2012, dans bouse de buffle, 4 ex.</p><p>- P.N.duW,AlphaKouara, IX 2003,plusieurs exemplaires,sous excrément humain.</p><p>- [Josso &amp; Prévost, 2015: P. N. Pendjari; P. N. du W, piste Koudou, Keremou; réserve de la Djona, Aphakouara. Boucomont, 1922: Kandy, donnée non revalidée dans le cadre de la révision de Josso &amp; Prévost (2015)].</p><p>*  Pedaria durandi Paulian, Cambefort &amp; Mauchamp, 1982</p><p>(Pl. 19, Fig. 3)</p><p>- Bohicon, quartier Soglogon, 06 X 2009, lumière, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E389655FC6CFF0FFAEEFC32	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FC68FC62FB28FBB0.text	857E879E8E389655FC68FC62FB28FBB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria estellae Josso & Prevost 2006	<div><p>Pedaria estellae Josso &amp; Prévost, 2006 (Pl. 19, Fig. 4 &amp; Pl. 75, Fig. 7)</p><p>- [Josso &amp; Prévost, 2015: parc de la Pendjari].</p></div>	https://treatment.plazi.org/id/857E879E8E389655FC68FC62FB28FBB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FC61FBE2FA64FB58.text	857E879E8E389655FC61FBE2FA64FB58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria fernandezi Cambefort 1982	<div><p>Pedaria fernandezi Cambefort, 1982</p><p>(Pl. 20, Fig. 1)</p><p>- Savalou, forêt de Logozohé, 06 VIII 2007, sous bouse zébu, 2 ex.; 04 IX 2007, sous bouse zébu, 1 ex.</p><p>- Pénéssoulou, centre-ville, 17 VI 2013, lumière, 1 ex.</p><p>- [Josso &amp; Prévost, 2015: parc de la Pendjari, chutes de Tanougou]</p></div>	https://treatment.plazi.org/id/857E879E8E389655FC61FBE2FA64FB58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FC7DFAC8FA86FADF.text	857E879E8E389655FC7DFAC8FA86FADF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria humana Cambefort 1982	<div><p>Pedaria humana Cambefort, 1982</p><p>(Pl. 20, Fig. 2 &amp; Pl. 75, Fig. 8)</p><p>- [Josso &amp; Prévost, 2015: environs de Save, Akongbere].</p></div>	https://treatment.plazi.org/id/857E879E8E389655FC7DFAC8FA86FADF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E389655FC7CFA48FB28F9DA.text	857E879E8E389655FC7CFA48FB28F9DA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria renwarti Cambefort 1982	<div><p>Pedaria renwarti Cambefort, 1982</p><p>(Pl. 20, Fig. 3)</p><p>-  Bohicon, quartier Soglogon, 03 VI 2018, U. V., 30 ex.; 25 VI 2018, 3 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, à vue, 27 X 2011, 1 ex.</p><p>- P.N.duW,Tapoa Djerma,mare aux Lions, IX 2003, plusieurs exemplaires sous crottins d’éléphants.</p><p>- [Josso &amp; Prévost, 2015: parc de la Pendjari].</p></div>	https://treatment.plazi.org/id/857E879E8E389655FC7CFA48FB28F9DA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E399657FEA1FF0AFB47F8FF.text	857E879E8E399657FEA1FF0AFB47F8FF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis affinis Felsche 1911	<div><p>*  Onitis affinis Felsche, 1911</p><p>(Pl. 21, Fig. 4)</p><p>- P. N. Pendjari, forêt de Bondjagou, 25 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P. N. du W, réserve de la Djona, piste entre Alpha Kouara et la mare 24, IX 2003, un exemplaire au piège lumineux.</p><p>*  Onitis alexis alexis Klug, 1835</p><p>(Pl. 22, Fig. 1 &amp; 2)</p><p>- Grand Popo, 22 II 2008, dans bouse de zébu, 5 ex.</p><p>- Ouidah, forêt de Pahou,  21 XII 2014, U. V., 1 ex.;  25 XI 2015, U. V., 4 ex.;  05 I 2017, U. V., 29 ex.</p><p>-  Houéyogbé, forêt de Houéyogbé, 15 X 2014,U. V., 1 ex.;  28 XII 2016, U. V., 1 ex.</p><p>-   Athiémé, bord du Mono, 29 XII 2014, U. V., 2 ex.</p><p>-  Lokossa, quartier Vivotin, 28 XII 2011, U. V., 1 ex.</p><p>-   Ouédémé Adja, bord du Mono, 30 XII 2016, U. V., 2 ex.</p><p>- Attogon, forêt de Niaouli, 01 V  2013, U.V., 1 ex.; 02 V  2013, U.V., 1 ex.;  18 XI 2015, U. V., 1 ex.</p><p>-  Adidevo, Dévé, carrière, 07 II 2017, U. V., 1 ex.</p><p>-   Kpoba, bord du Mono, 02 I 2017, U. V., 4 ex.</p><p>-  Pobé, forêt de Pobé, 13 II 2017, U. V., 4 ex.</p><p>-   Sahé, bord du Couffo, 14 V  2013, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, lumière, 5 ex.; 1 V  2013, U.V., 1 ex.; 16 V  2013, U.V., 3 ex.; 24 VI  2013, U.V., 1 ex.;  14 XI 2014, U. V., 4 ex.; 15 VI  2014, U.V., 2 ex.;  11 XI 2014, U. V., 3 ex.;  16 XI 2014, U. V., 1 ex.;  19 XI 2014, U. V., 5 ex.;  20 XI 2014, U. V., 2 ex.;  21 XI 2014, U. V., 2 ex.;  23 XI 2014, U. V., 5 ex.;  22 XI 2014, U. V., 7 ex.;  24 XI 2014, U. V., 5 ex.;  26 XI 2014, U. V., 7 ex.;  17 XII 2014, U. V., 3 ex.; 06 VI  2016, U.V., 15 ex.;  22 X 2017, U. V., 1 ex.;  12 XI 2017, U. V., 1 ex.;  26 XI 2017, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 10 XII 2014, U. V., 3 ex.</p><p>-  Pobé, 13 II 2017, U. V., 1 ex.</p><p>-  Djidja, forêt d’Atchérigbé, 14 II 2017, U. V., 7 ex.</p><p>-  Savalou, Kpataba, Lycée technique, bas fond, 14 XII 2016, U. V., 2 ex.</p><p>-  Djougou, forêt de Sérou, 23 I 2017, U. V., 1 ex.</p><p>-  Bassila, Manigri, stade de foot, 25 I 2017, U. V., 12 ex.</p><p>- Bassila, Tchétou, 17 VI 2016, sous bouse de zébu, 1 ex.</p><p>-  Bétérou, bord de l’Ouémé, 28 XII 2011, U. V., 2 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V  2013, U.V., 6 ex.; 25 V  2013, U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 02 VII  2012, U.V., 2 ex.;  11 X 2012, U. V., 2 ex.; 27 V  2013, U.V., 1 ex.; 04 VII  2013, U.V., 1 ex.; 30 VII  2013, U.V., 1 ex.; 15 VI  2016, U.V., 1 ex.;  24 I 2017, U. V., 1 ex.</p><p>- P.N.Pendjari,Batia,Camp Numi,  07 XI 2012, U. V., 1 ex.;  08 XI 2015, U. V., 1 ex.;  26 I 2017, U. V., 2 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 15 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P. N. du W, Alpha Kouara, IX 2003, quelques exemplaires, piège lumineux.</p><p>*  Onitis bocandei Gillet, 1911 (Pl. 22, Fig. 3)</p><p>- P. N. Pendjari, forêt de Bondjagou, 26  VII 2009, sous bouse éléphant, 5 ex.; VI 2005, près de la forêt de Bondjagou, quelques exemplaires, piège lumineux.</p><p>- P. N. du W, Point Triple, IX 2003, 1 ex.</p><p>*  Onitis camerunus Gillet, 1933</p><p>(Pl. 22, Fig. 4 &amp; Pl. 23, Fig. 1)</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013, sous bouse de zébu, 6 ex.</p><p>*  Onitis cupreus de Castelnau, 1840 (Pl. 23, Fig. 2 &amp; 3)</p><p>-  Houéyogbé, forêt de Houéyogbé, 16 X 2014, U. V., 1 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 11 XI 2014, U. V., 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 02 XII 2014, U. V., 1 ex.</p><p>- P. N. Pendjari, Batia, 18-25 VIII 2010, piège au sol, 1 ex.; 26 VIII 2010, piège excrément, 1 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 03 XII 2015, U. V., 1 ex.</p><p>- P. N. du W, réserve de la Djona, Alpha Kouara, Mare 24, IX 2003, quelques exemplaires sous excrément humain.</p><p>- Tapoa Djerma, IX 2003, quelques exemplaires, sous excrément humain.</p><p>- Mare aux Lions, IX 2003, quelques exemplaires, sous crottins d’éléphants.</p><p>*  Onitis gilleti Boucomont, 1933</p><p>(Pl. 23, Fig. 4)</p><p>- P. N. Pendjari, forêt de Bondjagou, VI 2005, piège lumineux, plusieurs exemplaires.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous crottins d’éléphants, plusieurs exemplaires; 25 VII 2009, sous bouse éléphant, 2 ex.; 26 VII 2009, sous bouse éléphant, 12 ex.</p><p>*  Onitis guineensis Gillet, 1918 (Pl. 24, Fig. 1 &amp; 2 &amp; Pl. 75, Fig. 11-12)</p><p>- P. N. Pendjari, près de la forêt de Bondjagou, VI 2005, piège lumineux, plusieurs exemplaires.</p><p>- P. N. Pendjari, direction hôtelPendjari, VI 2005, sous crottins d’éléphants, plusieurs exemplaires.</p><p>- P. N. du W, Tapoa Djerma, près de la mare aux Lions, IX 2003, sous crottins d’éléphants et au piège lumineux, plusieurs exemplaires.</p><p>*  Onitis nigeriensis Ferreira, 1977</p><p>(Pl. 24, Fig. 3 &amp; 4)</p><p>- Savalou, forêt de Logozohé, 06 VIII 2007, sous bouse de zébu, 1 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 2 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 28 V  2013, U.V., 2 ex.; 17 VI 2013, sous excrément, 2 ex.; 15 VI  2016, U.V., 3 ex.; 16 VI  2016, U.V., 1 ex.</p><p>- Pénéssoulou, centre ville, 17 VI  2013, U.V., 2 ex.</p><p>- P. N. Pendjari, direction hôtel Pendjari, VI 2005, sous crottins d’éléphants, plusieurs exemplaires; 26 VIII 2010, sous bouse buffle, 2 ex.</p><p>- P. N. du W, Tapoa Djerma, IX 2003, sous excrément humain, plusieurs exemplaires.</p><p>*  Onitis reichei reichei van Lansberge, 1875 (Pl. 25, Fig. 1 &amp; 2)</p><p>-  Ouidah, forêt de Pahou, 06 I 2017, U. V., 4 ex.</p><p>- Houéyogbé, forêt de Houéyogbé, 22 VIII  2011, U.V., 1 ex.;  15 X 2014, U. V., 5 ex.;  16 X 2014, U. V., 1 ex.; X 2015, lumière, 1 ex.;  23 XI 2015,U. V., 2 ex.;  24 XI 2015, U. V., 4 ex.; 25 VI  2016, U.V., 7 ex.;  28 XII 2016, U. V., 12 ex.</p><p>- Tokay, Lobogo, 14 VI  2013, U.V., 2 ex.</p><p>- Athiémé bord du Mono, 11 V  2013, U.V., 4 ex.; 29 XII 2014, 3 ex.;  30 I 2017, U. V., 2 ex.;  10 II 2017, U. V., 1 ex.</p><p>-   Ouédéme-Adja, bord du Mono, 30 XII 2016, U. V., 3 ex.</p><p>- Attogon, forêt de Niaouli,  30 IV 2013, U. V., 1 ex.;  17 XI 2015, U. V., 1 ex.;  18 XI 2015, U. V., 2 ex.;  02 II 2017, U. V., 1 ex.;  03 II 2017, U. V., 1 ex.</p><p>-   Kpoba, bord du Mono, 02 I 2017, U. V., 12 ex.</p><p>- Pobé, forêt de Pobé, 18 VI  2013, U.V., 1 ex.;  20 XI 2015, U. V., 1 ex.;  13 II 2017, U. V., 3 ex.</p><p>-  Zogbodomey, forêt de la Lama, 26 IV 2013, U. V., 1 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 05 V 2013, U. V., 1 ex.</p><p>- Savalou, forêt de Logozohé, 06 VIII 2007, 1 ex.</p><p>- Bohicon, quartier Soglogon, 14 IV 2012, 1 ex.;  11 XI 2014, U. V., 1 ex.</p><p>- Dan, bord du Zou, 04 VI  2013, U.V., 1 ex.;  10 XII 2014, U. V., 2 ex.</p><p>-  Djidja, forêt d’Atchérigbé, 14 II 2017, U. V., 2 ex.</p><p>-  Bassila, Manigri, stade de foot, 25 I 2017, U. V., 2 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 17 X 2011, lumière, 1 ex.; 19 X 2011, lumière, 1 ex.; 21 X 2011, lumière, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013, sous bouse zébu, 2 ex.</p><p>-   Saramanga village, 24 V  2013, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 12 V  2013, U.V., 1 ex.; 23 V  2013, U.V., 1 ex.; 27 V  2013, U.V., 2 ex.; 28 V  2013, U.V., 4 ex.; 02 VII  2013, U.V., 2 ex.; 03 VII  2013, U.V., 6 ex.; 04 VII  2013, U.V., 1 ex.; 30 VII  2013, U.V., 3 ex.; 15 VI  2016, U.V., 2 ex.</p><p>- Pénéssoulou, sortie du village, zone humide, 23 II 2020 1 ex.</p><p>- P. N. Pendjari, forêt de Bondjagou, 18 XI 2015, sous bouse éléphant, 1 ex.; 25 VII 2009, sous bouse éléphant, 5 ex.</p><p>- P.N.Pendjari,vershôtelPendjari, VI 2005,piège lumineux,plusieurs exemplaires.</p><p>- P. N. du W, Mamassi-Gourma, 30 IX 2015, lumière, 3 ex.</p><p>- P. N. du W, réserve de la Djona, Alpha kouara, IX 2003, sous bouses de zébus, plusieurs exemplaires.</p><p>*  Onitis sibutensis Janssens, 1943 (Pl. 25, Fig. 3)</p><p>- P. N. Pendjari, forêt de Bondjagou, 26 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P.N.duW,TapoaDjerma, IX 2003, sous excrément humain.</p><p>- Tanguiéta, chaîne de l’Atakora, IX 2003, au piège lumineux et sous excrément humain.</p><p>- P. N. Pendjari, chutes de Koudou, IX 2003, piège lumineux et excrément humain.</p><p>*  Onitis unguiculatus (Olivier, 1789)</p><p>(Pl. 69, Fig. 4)</p><p>- P. N. du W, Tapoa Djerma (côté Burkina Faso), IX 2003, piège lumineux, plusieurs exemplaires.</p><p>*  Onitis vanderkelleni van Lansberge, 1886 (Pl. 25, Fig. 4)</p><p>- Houéyogbé, forêt de Houéyogbé, III 2012, lumière, 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 30 VII 2013, U. V., 1 ex.</p><p>*  Onitis violaceus van Lansberge, 1875</p><p>(Pl. 26, Fig. 1)</p><p>- P. N. Pendajri, forêt de Bondjagou, 25 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P.N.Pendajri,hôtelPendjari, VI 2005, piège lumineux, plusieurs exemplaires. - P.N.duW,TapoaDjerma, IX 2003, piège lumineux et sous excrément humain.</p><p>- Tanguiéta,chaîne de l’Atakora, IX 2003, piège lumineux, plusieurs exemplaires.</p><p>*  Pleuronitis boucomonti Janssens, 1937 (Pl. 26, Fig. 2)</p><p>- P. N. Pendjari, Batia, 24 VII 2009, piège au sol, 1 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous crottins d’éléphants.</p><p>- P. N. Pendjari, versTanougou etau nord de Batia, VI 2005, sous bouses de zébus, plusieurs exemplaires.</p><p>- P. N. du W, Tapoa Djerma, IX 2003, 1 ex., au vol en fin de matinée.</p><p>*  Pleuronitis fulgidus (de Castelnau, 1840)</p><p>(Pl. 70, Fig. 1)</p><p>- P. N. du W, Tapoa Djerma, à 1 km de la mare aux Lions, IX 2003, sous excrément humain, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E399657FEA1FF0AFB47F8FF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3A9656FC03F868FE82FEE0.text	857E879E8E3A9656FC03F868FE82FEE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Clypeodrepanus strigatus (Janssens 1953)	<div><p>Clypeodrepanus strigatus (Janssens, 1953) (Pl. 26, Fig. 3 &amp; Pl. 76, Fig. 1)</p><p>- [Janssens, 1953: Wydah]</p><p>*  Cyptochirus distinctus (Janssens, 1953) (Pl. 26, Fig.4)</p><p>- P. N. Pendjari, Batia, 27 VII 2009, piège au sol, 1 ex.</p><p>- 7 km de la forêt de Boundjagou en direction de l’hôtel Pendjari, VI 2005, dans crottins d’éléphants.</p></div>	https://treatment.plazi.org/id/857E879E8E3A9656FC03F868FE82FEE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3B9656FF72FEB7FDC3FC92.text	857E879E8E3B9656FF72FEB7FDC3FC92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Eodrepanus bechynei (Janssens 1953)	<div><p>Eodrepanus bechynei (Janssens, 1953)</p><p>(Pl. 27, Fig. 1)</p><p>- Bohicon, 15 VI 2009, sous bouses de zébus, 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 23 XI 2020, sous bouse de zébu, 2 ex.</p><p>- Pénéssoulou, zone humide, sortie du village, 23 II 2020, 2 ex.</p><p>- [Barbero et al., 2009: Parakou; west of Benin N. P., N Kaobagou].</p><p>*  Euoniticellus capnus Cambefort, 1996 (Pl. 27, Fig. 2)</p><p>- P. N. Pendjari, forêt de Bondjagou, 26 VII 2009, sous bouse éléphant, 3 ex.</p><p>- P.N.Pendjari,vers hôtel Pendjari, 7 km de la forêt de Bondjagou, VI 2005, sous crottins d’éléphants, quelques exemplaires.</p><p>- P. N. du W, réserve de la Djona, piste Alpha Kouara, mare 24, IX 2003, sous crottins d’éléphants, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E3B9656FF72FEB7FDC3FC92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E3B9649FF63FC88FE7DFE77.text	857E879E8E3B9649FF63FC88FE7DFE77.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Euoniticellus intermedius (Reiche 1849)	<div><p>Euoniticellus intermedius (Reiche, 1849)</p><p>(Pl. 27, Fig. 3)</p><p>- Grand Popo, 22 II 2008, sous bouse de zébu, 31 ex.</p><p>- Lokossa, Djotokpa, 03 IV 2006, sous de zébu, 2 ex.</p><p>- Lokossa, quartier Agonvé, 12 VIII 2007, sous bouse de zébu, 1 ex.; 03 IV 2006, sous bouse de zébu, 6 ex.</p><p>- Bohicon, 15 VI 2009, 1 ex.;  22 XI 2017, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 03 XII 2014, 2 ex.</p><p>- Pénéssoulou, zone humide, sortie du village, 23 II 2020, sous bouse zébu, 2 ex.</p><p>- P. N. Pendjari, Batia, 24 VII 2009, 1 ex.; 18 VIII 2010, 7 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26 VIII 2010,sous excrément de buffle, 2 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 11 XI 2012, U. V., 2 ex.</p><p>- P. N. du W, réserve de la Djona, piste Alpha Kouara, mare 24, IX 2003, sous crottins d’éléphants, quelques exemplaires.</p><p>- [Boucomont, 1921: Wydah].</p><p>*  Euoniticellus kawanus Janssens, 1939 (Pl. 27, Fig. 4)</p><p>- P. N. Pendjari, forêt de Bondjagou, à vue sur bouse éléphants, 26 VII 2009, 1 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous crottins d’éléphants, nombreux exemplaires.</p><p>*  Euoniticellus nasicornis (Reiche, 1849)</p><p>(Pl. 70, Fig. 2 &amp; Pl. 76, Fig. 2)</p><p>- P. N. du W, réserve de la Djona, piste Alpha Kouara, mare 24, IX 2003, sous bouses de zébus, quelques exemplaires.</p><p>*  Euoniticellus parvus (Kraatz, 1895) (Pl. 28, Fig. 1)</p><p>-  Zogbodomey, forêt de Lokoli, 14 II 2020, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, lumière, 1 ex.</p><p>-  Pobé, forêt de Pobé, 19 XI 2015, U. V., 1 ex.</p><p>- Savalou, forêt de Logozohé, 04 IX 2007, sous excrément de zébu, 3 ex.</p><p>-  Savalou, Kpataba, lycée technique, bas fond, 14 XII 2016, U. V., 2 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 4 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 03 XII 2014, U. V., 5 ex.</p><p>- Pénéssoulou, zone humide, sortie du village, 23 II 2020, 8 ex.</p><p>- Ndali, forêt de Ndali, 11 XI 2015, bouse zébu, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 24 VIII 2010, sous bouse de buffle, 2 ex.</p><p>- P. N. Pendjari, mare Bali, 07 XI 2012, sous bouse de buffle, 5 ex.</p><p>- P. N. Pendjari,forêtde Bondjagou, VI 2005, sous crottins d’éléphants, quelques exemplaires.</p><p>- P. N. du W, réserve de la Djona, piste Alpha Kouara, 24, IX 2003, sous bouses de zébus, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E3B9649FF63FC88FE7DFE77	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E249649FF18FE20FD47FA82.text	857E879E8E249649FF18FE20FD47FA82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Euoniticellus triangulatus (von Harold 1873)	<div><p>Euoniticellus triangulatus (von Harold, 1873) (Pl. 28, Fig. 2 &amp; Pl. 76, Fig. 3)</p><p>- [Boucomont, 1921: Wydah].</p><p>*  Ixodina bos (Endrödi, 1976)</p><p>(Pl. 28, Fig. 3)</p><p>- Savalou, forêt de Logozohé, sous bouse de zébu, 6 VIII 2007, 1 ex.</p><p>*  Ixodina endroedyi (Endrödi, 1976) (Pl. 28, Fig. 4 &amp; Pl. 76, Fig. 4)</p><p>-  Zogbodomey, forêt de Lokoli, 14 II 2020, U. V., 2 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 25 V 2013, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 23 XI 2019, sous bouse de zébu, 1 ex.</p><p>- Pénéssoulou, zone humide, sortie du village, 23 II 2020, 1 ex.</p><p>- P. N. Pendjari, Batia, piège crevettes, 24 VIII 2010, 3 ex.</p><p>- 7 km de la forêt de Bondjagou.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005,sous crottins d’éléphants,plusieurs exemplaires;</p><p>- P. N. du W, piste Kérémou-Koudou, IX 2003, sous bouses de zébus, plusieurs exemplaires.</p><p>- Zogbodomey, forêt de Lokoli, 14 XI 2019, sous excrément zébu, 2 ex.</p><p>*  Ixodina szunyoghyi occidentalis (Endrödi, 1976)</p><p>(Pl. 29, Fig. 1 &amp; Pl. 76, Fig. 5)</p><p>- Bassila, Tchétou, 17 XI 2016, sous bouse de zébu, 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 03 XII 2014, U. V., 2 ex.; 23 XI 2020, dans bouse de zébu, 6 ex.</p><p>- Pénéssoulou, zone humide, sortie du village, 23 II 2020, 9 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E249649FF18FE20FD47FA82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E249649FF13FA97FB7BFEFA.text	857E879E8E249649FF13FA97FB7BFEFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Latodrepanus simonisi (Barbero 2009)	<div><p>Latodrepanus simonisi (Barbero et al., 2009) (Pl. 29, Fig. 2)</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, lumière, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu, 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 03 XII 2014, U. V., 2 ex.</p><p>- Ndali, forêt de Ndali, bouse zébu, 11 XI 2015, 1 ex.</p><p>- [Boucomont, 1921 et Janssens, 1953: Ouidah (sub laticollis)].</p><p>*  Liatongus militaris (de Castelnau, 1840)</p><p>(Pl. 29, Fig. 3 &amp; Pl. 76, Fig. 6)</p><p>- P. N. du W, réserve de la Djona, piste Alpha Kouara, IX 2003, sous crottins d’éléphants, quelques exemplaires.</p><p>*  Oniticellus formosus Chevrolat, 1830 (Pl. 29, Fig. 4)</p><p>- Lokossa, 12 VIII 2007, sous bouse de zébu, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013, sous bouse de zébu, 1 ex.</p><p>- Pénéssoulou, zone humide, sortie du village, 23 II 2020, 1 ex.</p><p>- Tanguiéta, P.N. Pendjari, Batia, 18 VIII 2010, lumière, 1 ex.; 09 XI 2015, dans excrément, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005,sous crottins d’éléphants,quelques exemplaires.</p><p>- P.N.duW,pistede KoudouauPointTriple, IX 2003,bouses de buffles,quelques exemplaires.</p><p>Tibiodrepanus sulcicollis (de Castelnau, 1840)</p><p>(Pl. 30, Fig. 1-2 &amp; Pl. 76, Fig. 7-8)</p><p>- Lokossa, Djetopka, 12 VIII 2007, 3 ex.</p><p>- [Barbero et al., 2011: Djona réserve]</p></div>	https://treatment.plazi.org/id/857E879E8E249649FF13FA97FB7BFEFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E249649FC1DFE90FA61FC37.text	857E879E8E249649FC1DFE90FA61FC37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiniocellus praetermissus Branco 2010	<div><p>Tiniocellus praetermissus Branco, 2010</p><p>(Pl. 30, Fig. 3)</p><p>- Lokossa, 24 III 2006, sous bouse zébu, 3 ex.</p><p>- Savalou, forêt de Logozohé, sous bouse zébu, 06 VIII 2007, sous bouse zébu, 1 ex.; 04 IX 2007, sous bouse de buffle, 2 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 27 X 2011, sous excrément, 1 ex.; 29 X 2011, lumière, 1 ex.;  29 X 2013, U. V., 1 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 1 ex.; 25 V 2013, sous bouse zébu, 7 ex.</p><p>- P.N.Pendjari,Batia, 24 VIII 2008, piège au sol, 2 ex.; 24 VII 2009, piège au sol, 7 ex.; 27 VII 2009, piège au sol, 1 ex.; 24 VIII 2010, sous excrément humain, 14 ex.</p><p>-  P. N. Pendjari, Batia, Camp Numi, 11 XI 2012, U. V., 6 ex.; 09 XI 2015, sous excrément, 3 ex.</p><p>- P. N. Pendjari, mare Bali, 07 XI 2012, sous bouse de buffle, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26 VIII 2010, sous bouse de buffle, 9 ex.</p><p>- P. N. Pendjari, forêt de Bondjagou, VI 2005, sous crottins d’éléphants, quelques exemplaires.</p><p>- P.N.duW,chutesdeKoudou,PointTriple, IX 2003,sous excrément humain.</p><p>- [Branco, 2010: Alfakouara, Res. Djona; Kaobagou (Parc du W)].</p></div>	https://treatment.plazi.org/id/857E879E8E249649FC1DFE90FA61FC37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E249649FC71FC60FB2DFAE7.text	857E879E8E249649FC71FC60FB2DFAE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiniocellus setifer (Kraatz 1895)	<div><p>Tiniocellus setifer (Kraatz, 1895)</p><p>(Pl. 30, Fig. 4)</p><p>- Grand Popo, 22 II 2008, 8 ex.</p><p>-  Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, U. V., 1 ex.</p><p>- Lokossa, 01 III 2006, 1 ex., sous bouse de zébu; 24 III 2006, sous bouse de zébu, 5 ex.; 12 VIII 2007, sous bouse de zébu, 1 ex.</p><p>- Allada, Donou, 10 VI 2016, 3 ex.</p><p>- Savalou, forêt de Logozohé, 04 IX 2007, 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 29 X 2011, lumière, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu, 2 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 28 V  2013, U.V., 1 ex.; 03 XII 2014, 1 ex.; 16 VI  2016, U.V., 3 ex.</p><p>- Pénéssoulou, zone humide, sortie du village, 23 II 2020, 11 ex.</p><p>- [Branco, 2010: Bembereke, 2 km W Gando].</p></div>	https://treatment.plazi.org/id/857E879E8E249649FC71FC60FB2DFAE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E249648FC18FAB3FDECFC98.text	857E879E8E249648FC18FAB3FDECFC98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagini Burmeister 1846	<div><p>ONTHOPHAGINI Burmeister, 1846</p><p>*  Alloscelus paradoxus Boucomont, 1923 sensu lato</p><p>(Pl. 31, Fig. 1)</p><p>-  Houéyogbé, forêt de Houéyogbé, 15 X 2014, U. V., 1 ex.</p><p>*  Cambefortius bidentiger (d’Orbigny, 1905) (Pl. 31, Fig. 2)</p><p>-  Pénéssoulou, forêt de Pénélan, 16 VI 2016, U. V., 1 ex.</p><p>*  Heteroclitopus sp.</p><p>(Pl. 31, Fig. 3)</p><p>-  Pénéssoulou, forêt de Pénélan, 3 VII 2013, U. V., 1 ex.</p><p>*  Pinacotarsus dohrni von Harold, 1875 (Pl. 31, Fig. 4)</p><p>-  Pénéssoulou, forêt de Pénélan, 23 V 2013, U. V., 2 ex.</p><p>*  Pseudosaproecius comatus (d’Orbigny, 1902)</p><p>(Pl. 32, Fig. 1)</p><p>- Zogbodomey, forêt de la Lama, 06 VI  2013, U.V., 1 ex. -   Pénéssoulou, forêt de Pénélan ,  11X 2012,U. V., 1 ex.; 27V  2013,U.V., 1 ex.; 28 V  2013,U.V., 2 ex.; 04VII  2013,U.V., 1 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 09 XI 2012, U. V., 1 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2005,piège lumineux,quelques exemplaires.</p><p>*  Pseudosaproecius cf cylindroides (d’Orbigny, 1908) (Pl. 32, Fig. 2)</p><p>- Ouidah, forêt de Pahou,  21 XI 2015,U. V., 1 ex.;  25 XI 2015, U. V., 3 ex.;  06 I 2017, U. V., 1 ex.</p><p>- Houéyogbé, forêt de Houéyogbé,  22 X 2012,U. V., 2 ex.; 01 VI  2013,U.V., 3 ex.;  15 X 2014, U. V., 1 ex.; 25 VI  2016,U.V., 1 ex.; 26 VI  2018, U.V., 4 ex.</p><p>- Tokay,Lobogo, 14 VI  2013, U.V., 3 ex.</p><p>-  Zinvié, vallée du Sitatunga, 19 V 2016, U. V., 1 ex.</p><p>- Attogon,forêtde Niaouli, 07VI 2013, 4 ex.;  01IV 2013,U. V., 1 ex.;  30 IV 2013, U. V., 1 ex.; 26 VI  2013, U.V., 1 ex.;  16XII 2014, U. V., 1 ex.;  18 XI 2015, U. V., 2 ex.; 07 VI  2016,U.V., 1 ex.; 21V  2016,U.V., 1 ex.; 14VI  2016,U.V., 1 ex.; 18VI  2016,U.V., 1 ex.</p><p>- Zogbodomey, forêt de la Lama, 15 V  2013, U.V., 1 ex.; 02 VI  2018, U.V., 1 ex.</p><p>-  Bohicon, quartier Soglogon, IV 2012, U. V., 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 02 VII 2013, U. V., 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E249648FC18FAB3FDECFC98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E259648FCECFE07FAA6FB80.text	857E879E8E259648FCECFE07FAA6FB80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pseudosaproecius Balthasar 1941	<div><p>Pseudosaproecius complexe validicornis (Quedenfeldt, 1884) (Pl. 33, Fig. 2-3 &amp; Pl. 76, Fig. 10)</p><p>-  Abomey-Calavi, Zinvié, vallée du Sitatunga, 26 V 2016, U. V., 2 ex.</p><p>- Attogon, Niaouli, 07 VI  2016, U.V., 2 ex.</p><p>- Zogbodomey, forêt de la Lama,  26 IV 2013, U. V., 12 ex.; 03 VI  2013,U.V., 1 ex.; 06 VI 2013, 8 ex.,  U. V., 1 ex.; 07 VI  2013, U.V., 8 ex.; 15 V  2013, U.V., 1 ex.; 16 VI  2013, U.V., 1 ex.; 22 VI  2018, U.V., 1 ex.</p><p>-   Sahé, bord du Couffo, 14 V  2013, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V. 35 ex.</p><p>- Savalou, Kpataba, montagne, 17 VI  2016, U.V., 1 ex.</p><p>-  Savalou, Kpataba, lycée technique, bas fond, 14 XII 2016, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 12 V  2013, U.V., 1 ex.; 27 V  2013, U.V., 1 ex.; 03 VII  2013, U.V., 3 ex.; 04 VII  2013, U.V., 2 ex.</p><p>- Pénéssoulou, centre ville, stade de foot, 22 V  2013, U.V., 1 ex.; 17 VI  2013, U.V., 1 ex.</p><p>- P.N.Pendjari,Batia,Camp Numi,  03 XI 2015, U. V., 1 ex.;  07 XI 2015, U. V., 1 ex.;  08 XI 2015, U. V., 2 ex.;  9 XI 2015, U. V., 1 ex.</p><p>- P.N.duW,pistedeTapoaDjerma, IX 2003,piège lumineux,quelques exemplaires.</p><p>- [Orbigny, 1902: Ouidah (citation reprise dans BRANCO, 1995)].</p></div>	https://treatment.plazi.org/id/857E879E8E259648FCECFE07FAA6FB80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E259648FF6EFC8FFD8AFC1A.text	857E879E8E259648FF6EFC8FFD8AFC1A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pseudosaproecius ocellatus (Frey 1958)	<div><p>*  Pseudosaproecius ocellatus (Frey, 1958)</p><p>(Pl. 32, Fig. 3)</p><p>-  Zogbodomey, forêt de la Lama, 06 VI 2013, U. V., 2 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E259648FF6EFC8FFD8AFC1A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E259648FF0FFC0FFA04FE12.text	857E879E8E259648FF0FFC0FFA04FE12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pseudosaproecius portentosus (Felsche 1907)	<div><p>*  Pseudosaproecius portentosus (Felsche, 1907) (Pl. 32, Fig. 4; Pl. 33, Fig. 1 &amp; Pl. 76, Fig. 9)</p><p>-  Attogon, forêt de Niaouli, 14 VI 2016, U. V., 1 ex.</p><p>-  Bohicon, quartier Soglogon, 27 IV 2013, U. V., 4 ex.</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V. 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 12 V  2013, U.V., 1 ex.; 23 V  2013,U.V., 7 ex.; 27 V  2013, U.V., 2 ex.; 28 V  2013, U.V., 1 ex.; 02 VII  2013, U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénéssoulou, 12 V 2013, sous excrément, 1 ex.</p><p>-  P.N. delaPendjari, Batia, 07 XI 2015, U. V., 2 ex.</p><p>- P.N.duW,pistedeTapoaDjerma, IX 2003,piège lumineux,quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E259648FF0FFC0FFA04FE12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFF0EF97FFD36F8AA.text	857E879E8E26964BFF0EF97FFD36F8AA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobiomorphus megaponerae Brauns 1914	<div><p>*  Caccobiomorphus megaponerae Brauns, 1914</p><p>(Pl. 35, Fig. 2)</p><p>-  Houéyogbé, forêt de Houéyogbé, 20 X 2014, U. V., dans excrément, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFF0EF97FFD36F8AA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFF0DF8FFFE7FF79F.text	857E879E8E26964BFF0DF8FFFE7FF79F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius anthracites (d'Orbigny 1904)	<div><p>*  Caccobius (s. l.) anthracites (d'Orbigny, 1904) (Pl. 35, Fig. 3)</p><p>- P. N. Pendjari, Batia, Camp Numi,  07 XI 2012, U. V., 2 ex.;  08 XI 2012, U. V., 1 ex.;  10 XI 2012, U. V., 1 ex.;  11 XI 2012, U. V., 3 ex.;  07 XI 2015, U. V., 1 ex.;  08 XI 2015, U. V., 1 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous excrément humain.</p><p>- P. N. duW, chutes deKoudou, Point Triple,sous excrément humain et sous crottins d’éléphants, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFF0DF8FFFE7FF79F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFC01FF0FFC40FD28.text	857E879E8E26964BFC01FF0FFC40FD28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius auberti d'Orbigny 1902	<div><p>*  Caccobius (s. l.) auberti d'Orbigny, 1902 (Pl. 35, Fig. 4)</p><p>- Allada, Donou, 10 VI 2016, sous bouse de zébu, 1 ex.</p><p>- Savalou, forêt de Logozohé, 06 VIII 2007, sous bouse de zébu, 1 ex.; 04 IX 2007, sous bouse de zébu, 3 ex.</p><p>- Bassila, Tchétou, 17 VI 2016, sous bouse de zébu, 3 ex.</p><p>-  Pénéssoulou, forêtde Saramanga, 24V 2013,U. V., 2 ex.; 25 V 2013, sous bouse de zébu, 1 ex.</p><p>- P.N.Pendjari,Batia, 27 VII 2009, piège au sol, 1 ex.</p><p>- P.N.Pendjari,versTanougou, VI 2005, bouses de zébus, quelques exemplaires.</p><p>- P.N.duW,réservedelaDjona,AlphaKouara, IX 2003, sous bouses de zébus.</p><p>-   Tanguiéta, Atakora, IX 2003, sur fleur fanée  !</p><p>- P.N. duW,pisteKérémou-Koudou, IX 2003,sous bouses de zébus, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFC01FF0FFC40FD28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFC6EFD7FFAF5FC8F.text	857E879E8E26964BFC6EFD7FFAF5FC8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius bama Walter 2014	<div><p>*  Caccobius (s. l.) bama Walter, 2014</p><p>(Pl. 36, Fig. 1)</p><p>-  P. N. du W, Karimama, 28 IX 2015, U. V., 1 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFC6EFD7FFAF5FC8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFC19FC98FB5FFBBA.text	857E879E8E26964BFC19FC98FB5FFBBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius cavatus d'Orbigny 1908	<div><p>Caccobius (s. l.) cavatus d'Orbigny, 1908 (Pl. 36, Fig. 2 &amp; Pl. 76, Fig. 12)</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 2 ex.</p><p>- Bohicon, quartier Soglogon, 25 VI  2018, U.V., 2 ex.</p><p>- [Walter, 2014: Tanguiéta (Atakora)]</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFC19FC98FB5FFBBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFC0AFBD0FA05FB57.text	857E879E8E26964BFC0AFBD0FA05FB57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius dorsalis von Harold 1867	<div><p>*  Caccobius (s. l.) dorsalis von Harold, 1867</p><p>(Pl. 36, Fig. 3)</p><p>-  Houéyogbé, forêt de Houéyogbé, 26 VI 2016, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 06 VI  2016, U.V., 6 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 4 ex.</p><p>- P.N.duW,Mamassi-Gourma, 29 IX 2015,lumière, 3 ex.; 30 IX 2015,lumière, 28 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFC0AFBD0FA05FB57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFC0AFAC0FC25F93A.text	857E879E8E26964BFC0AFAC0FC25F93A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius ferrugineus Fahraeus 1857	<div><p>Caccobius (s. l.) ferrugineus Fahraeus, 1857 (Pl. 36, Fig. 4)</p><p>-  Houéyogbé, forêt de Houéyogbé, 01 VI 2013, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon,  27IV 2013,U. V., 6 ex.; 11V  2013,U.V., 2 ex.;  21XI 2017,U. V., 1 ex.;  29 XI 2017,U. V., 1 ex.; 17V  2018,U.V., 2 ex.; 06 VIII  2018,U.V., 1 ex.</p><p>- Za-Kpota, Allahé, 30 V  2018, U.V., 3 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 3 ex.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 75 ex.</p><p>- P.N.duW,TapoaDjerma, IX 2003, piège lumineux, quelques exemplaires.</p><p>- [Orbigny, 1902: Ouidah. Orbigny, 1913: cercle de Djougou-Kouandé. Walter,2019: Bénin].</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFC0AFAC0FC25F93A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFC0AF950FC7DF8AF.text	857E879E8E26964BFC0AF950FC7DF8AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius ivorensis Cambefort 1984	<div><p>*  Caccobius (s. l.) ivorensis Cambefort 1984</p><p>(Pl. 37, Fig. 1 &amp; Pl. 77, Fig. 1-2)</p><p>- P. N. du W, chutes de Koudou, IX 2003, sous excrément humain, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFC0AF950FC7DF8AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFCC0F8F8FAEEF82F.text	857E879E8E26964BFCC0F8F8FAEEF82F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius mirabilepunctatus Cambefort 1971	<div><p>*  Caccobius (s. l.) mirabilepunctatus Cambefort, 1971 (Pl. 37, Fig. 2)</p><p>- P. N. de la Pendjari, Batia, 24 VIII 2010, lumière, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFCC0F8F8FAEEF82F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFC14F87AFC24F797.text	857E879E8E26964BFC14F87AFC24F797.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius morettoi Walter 2014	<div><p>Caccobius (s. l.) morettoi Walter, 2014</p><p>(Pl. 70, Fig. 3)</p><p>- [ P. N. duW, falaise de Tansarga (côté Burkina Faso), IX 2003, piège lumineux, quelques exemplaires] .</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFC14F87AFC24F797	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFF4CFDF0FE33FB97.text	857E879E8E26964BFF4CFDF0FE33FB97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Milichus apicalis (Fahraeus 1857)	<div><p>*  Milichus apicalis (Fahraeus, 1857) (Pl. 34, Fig. 2-3)</p><p>-   Athiémé, rive du Mono, 12 II 2017, U. V., 1 ex.</p><p>- Houéyogbé, forêt de Houéyogbé, 08 V  2013,U.V., 1 ex.; 25 VI  2016,U.V., 2 ex.</p><p>- Lokossa, Djetokpa, 03 IV 2006,sous bouse zébu, 1 ex.; 12VIII 2007,sousbouse de zébu, 1 ex.;  28 XII 2011, U. V., 2 ex.</p><p>- Attogon, forêt de Niaouli,  01 IV 2013, U. V., 1 ex.;  29 IV 2013, U. V., 1 ex.;  30 IV 2013, U. V., 4 ex.; 02 V  2013, U.V., 2 ex.; 03 V  2013, U.V., 1 ex.;  02 II 2017, U. V., 1 ex.;  12 II 2017, U. V., 2 ex.;  02 II 2017, U. V., 2 ex.</p><p>-  Zogbodomey, forêt de la Lama, 02 VI 2018, U. V., 2 ex.</p><p>- Bohicon, quartier Soglogon, 03 VI  2018, U.V., 1 ex.</p><p>-   Kpoba, bord du Mono, 02 I 2017, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 10 XII 2014, U. V., 1 ex.</p><p>-  Djougou, forêt de Sérou, 01 X 2019, U. V., 2 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFF4CFDF0FE33FB97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFF4EFB82FD98FAFA.text	857E879E8E26964BFF4EFB82FD98FAFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Milichus durandi Cambefort 1996	<div><p>*  Milichus durandi Cambefort, 1996</p><p>(Pl. 34, Fig. 4)</p><p>-  Pénéssoulou, forêt de Pénélan, 03 VII 2013, U. V., 1 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 9 ex.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 4 ex.</p><p>- P.N.duW,TaopaDjerma, IX 2003, piège lumineux, quelques exemplaires.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFF4EFB82FD98FAFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFF42FAAFFD6CF928.text	857E879E8E26964BFF42FAAFFD6CF928.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Milichus serratus d'Orbigny 1907	<div><p>*  Milichus serratus d'Orbigny, 1907 (Pl. 35, Fig. 1)</p><p>- Lokossa, 24 III 2006, sous bouse zébu, 3 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 15 VI 2016, lumière, 1 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 03 XI 2015, U. V., 1 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P. N. duW, réserve de la Djona,Alpha Kouara, IX 2003, sous bouses de zébus, quelques exemplaires.</p><p>-   Tanguiéta, Atakora, IX 2003, dans fleur fanée et sous excrément humain  .</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFF42FAAFFD6CF928	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFF7DFF0AFEDBFE87.text	857E879E8E26964BFF7DFF0AFEDBFE87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Stiptopodius doriae von Harold 1871	<div><p>*  Stiptopodius doriae von Harold, 1871 (Pl. 33, Fig. 4 &amp; Pl. 76, Fig. 11)</p><p>- P. N. du W, réservedelaDjona:Alpha Kouara, IX 2003, piège lumineux, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFF7DFF0AFEDBFE87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E26964BFF6BFE92FD9DFDA7.text	857E879E8E26964BFF6BFE92FD9DFDA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Stiptopodius gaillardi (Boucomont 1923)	<div><p>*  Stiptopodius gaillardi (Boucomont, 1923)</p><p>(Pl. 34, Fig. 1)</p><p>- P. N. du W, réservede laDjona:Alpha Kouara, IX 2003, piège lumineux, quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003, piège lumineux, quelques exemplaires.</p><p>- P.N.duW,MamassiGourma, 30 IX 2015, lumière, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E26964BFF6BFE92FD9DFDA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E27964AFF07FF0AFDB4FE98.text	857E879E8E27964AFF07FF0AFDB4FE98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius pentagonus d'Orbigny 1908	<div><p>*  Caccobius (s. l.) pentagonus d'Orbigny, 1908 (Pl. 37, Fig. 3)</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 11 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E27964AFF07FF0AFDB4FE98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E27964AFF34FE8FFEA1FCAF.text	857E879E8E27964AFF34FE8FFEA1FCAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius punctatissimus von Harold 1867	<div><p>Caccobius (s. l.) punctatissimus von Harold, 1867</p><p>(Pl. 37, Fig. 4)</p><p>-  Bohicon, quartier Soglogon, 27 IV 2013, U. V., 1 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 25 V  2013, U.V., 1 ex.; 27 V  2013, U.V., 1 ex.; 15 VI  2016, U.V., 1 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P.N.Pendjari,hôtel Pendjari, VI 2005, piège lumineux, bouses de zébus et sous excrément humain, plusieurs exemplaires.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 59 ex.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 39 ex.</p><p>- P.N.duW,TapoaDjerma, IX 2003, sous bouses de buffles, quelques exemplaires.</p><p>- [Orbigny, 1902: Abeokouta; Ouidah. ORBIGNY, 1913: cercle de Djougou-Kouandé. WALTER, 2019: Bénin].</p></div>	https://treatment.plazi.org/id/857E879E8E27964AFF34FE8FFEA1FCAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E27964AFF37FCF8FD10FC2F.text	857E879E8E27964AFF37FCF8FD10FC2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius signatipennis von Harold 1867	<div><p>*  Caccobius (s. l.) signatipennis von Harold, 1867 (Pl. 38, Fig. 1 &amp; Pl. 77, Fig. 3)</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous excrément humain, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E27964AFF37FCF8FD10FC2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E27964AFF60FC7AFBD0FEAF.text	857E879E8E27964AFF60FC7AFBD0FEAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius villiersi Cambefort 1975	<div><p>Caccobius (s. l.) villiersi Cambefort, 1975</p><p>(Pl. 38, Fig. 2 &amp; Pl. 77, Fig. 4)</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous excrément humain, quelques exemplaires.</p><p>- P. N.duW,TapoaDjerma, IX 2003, piège lumineux et excrément humain, quelques exemplaires.</p><p>- [Walter, 2019: Bénin].</p></div>	https://treatment.plazi.org/id/857E879E8E27964AFF60FC7AFBD0FEAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E27964AFCD8FEF8FC2EFE17.text	857E879E8E27964AFCD8FEF8FC2EFE17.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Cleptocaccobius balthasarianus (Cambefort 1971)	<div><p>*  Cleptocaccobius balthasarianus (Cambefort, 1971) (Pl. 38, Fig. 3 &amp; Pl. 77, Fig. 5)</p><p>- P. N. du W, piste Keremou-Koudou, 02 VI 2004, sous bouses de zébus, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E27964AFCD8FEF8FC2EFE17	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E27964AFC27FE02FA05FD90.text	857E879E8E27964AFC27FE02FA05FD90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Cleptocaccobius convexifrons (Raffray 1877)	<div><p>*  Cleptocaccobius convexifrons (Raffray, 1877)</p><p>(Pl. 38, Fig. 4 &amp; Pl. 77, Fig. 6)</p><p>- P.N.Pendjari,versTanougou, VI 2005,sous bouses de zébus,quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E27964AFC27FE02FA05FD90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E27964AFC32FD82FC3DFD52.text	857E879E8E27964AFC32FD82FC3DFD52.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Cleptocaccobius dorbignyi Cambefort 1984	<div><p>*  Cleptocaccobius dorbignyi Cambefort, 1984 (Pl. 39, Fig. 1 &amp; Pl. 77, Fig. 7)</p><p>- P. N. Pendjari, mare Bali, 7 VII 2012, sous bouse buffle, 1 ex.</p><p>- P.N.duW,réservedelaDjona,AlphaKouara, IX 2003, sous bouses de zébus, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E27964AFC32FD82FC3DFD52	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E27964AFC2DFCC8FA72FBBF.text	857E879E8E27964AFC2DFCC8FA72FBBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Cleptocaccobius signaticollis (d'Orbigny 1902)	<div><p>*  Cleptocaccobius signaticollis (d'Orbigny, 1902)</p><p>(Pl. 39, Fig. 2)</p><p>- P.N. Pendjari,vers Tonougou, VI 2005,sous excrément de Babouins, quelques exemplaires.</p><p>- P.N.duW,réservedelaDjona,AlphaKouara, IX 2003, sous bouses de zébus, quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003,sous excrément humain,quelques exemplaires.</p><p>- P. N. Pendjari, Batia, mare Bali, 07 XI 2012, sous bouse buffle, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E27964AFC2DFCC8FA72FBBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964DFF69F957FEB8F942.text	857E879E8E20964DFF69F957FEB8F942.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Afrostrandius adspersus (d'Orbigny 1908)	<div><p>Afrostrandius adspersus (d’Orbigny, 1908)</p><p>(Pl. 41, Fig. 4 &amp; Pl. 77, Fig. 11)</p><p>- [Moretto, 2009: Dahomey].</p></div>	https://treatment.plazi.org/id/857E879E8E20964DFF69F957FEB8F942	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964DFF08F8D7FBE2FB77.text	857E879E8E20964DFF08F8D7FBE2FB77.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Bicornonthophagus cornifrons (Thomson 1858)	<div><p>Bicornonthophagus cornifrons (Thomson, 1858) (Pl. 42, Fig. 1-2 &amp; Pl. 77, Fig. 12)</p><p>-  Ouidah, forêt de Pahou, 18 X 2014, U. V., 1 ex.</p><p>-  Zinvié, vallée du Sitatunga, 19 VII 2016, U. V., 1 ex.</p><p>- Houéyogbé, forêt de Houéyogbé,  02 IV 2013, U. V., 10 ex.; 01 VI  2013, U.V., 5 ex.;  24 XI 2015, U. V., 1 ex.; 25 VI  2016, U.V., 2 ex.</p><p>- Athiémé, bord du Mono, 31 V  2013, U.V., 4 ex.;  10 II 2017, U. V., 10 ex.</p><p>- Tokay,Lobogo, 14 VI  2013, U.V., 4 ex.</p><p>- Lokossa, 01 III 2006, fauchoir, 1 ex.; 18 III  2006, sous bouse zébu, 1 ex.; 12 VIII</p><p>2007, sous bouse zébu, 1 ex.</p><p>- Lokossa, Agonvé, 27 XII 2016, 1 ex.</p><p>- Attogon, forêt de Niaouli, 18 VI  2016, U.V., 1 ex.;  02 II 2017, U. V., 9 ex.</p><p>-   Ouédémé-Adja, bord du Mono, 30 XII 2016, U. V., 1 ex.</p><p>-   Kpoba, bord du Mono, 02 I 2017, U. V., 2 ex.</p><p>- Bohicon, 15 V  2018, U.V., 1 ex.</p><p>-  Pobé, forêt de Pobé, 27 X 2014, U. V., 1 ex.;  20 XI 2015, U. V., 1 ex.</p><p>- Zogbodomey, forêt de Lokoli, 05 V  2012, U.V., 1 ex.;  30 I 2019, U. V., 1 ex.</p><p>-  Zogbodomey, Domé, 13 XII 2014, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V., 1 ex.</p><p>- Savalou, Kpataba, montagne, 17 VI  2016, U.V., 1 ex.</p><p>- Pénéssoulou, centre ville, stade de foot,  13 X 2012, U. V., 5 ex.;  18 X 2012, U. V., 4 ex.; 12 V  2013,U.V., 3 ex.; 22 V  2013, U.V., 28 ex.; 27 V  2013,U.V., 8 ex.; 28 V  2013, U.V., 7 ex.; 17 VI  2013, U.V., 9 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 23 V  2013, U.V., 3 ex.; 25 V  2013, U.V., 2 ex.; 27V  2013,U.V., 2 ex.; 28 V  2013, U.V., 2 ex.; 02 VII  2013, U.V., 1 ex.; 03 VII  2013, U.V., 1 ex.; 16 VI  2016,U.V., 1 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 49 ex.</p><p>- P.N.Pendjari,Batia, 24VII 2009,piègeausol, 3 ex.; 27VII 2009,piège ausol, 1 ex.; 29 VII 2009,piège au sol, 7 ex.; 24 VIII2010,lumière, 8 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, piège au sol, 8 ex.</p><p>- P.N.Pendjari,hôtel Pendjari, VI 2006, piège lumineux, quelques exemplaires; 26 VIII 2010, sous bouse de buffle, 86 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 20 ex.</p><p>- P.N.duW,MamassiGourma, 30 IX 2015, lumière, 175 ex.</p><p>- P.N.duW,réservedelaDjona:AlphaKouara, IX 2003, sous bouses de zébus, quelques exemplaires.</p><p>- P.N.duW,pointtriple, IX 2003, sous excrément humain, quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003, piège lumineux, quelques exemplaires.</p><p>- [Orbigny, 1913: Dahomey].</p></div>	https://treatment.plazi.org/id/857E879E8E20964DFF08F8D7FBE2FB77	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964DFC2DFB20FA06FA6F.text	857E879E8E20964DFC2DFB20FA06FA6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Bicornonthophagus decorsei (d'Orbigny 1908)	<div><p>*  Bicornonthophagus decorsei (d'Orbigny, 1908) (Pl. 42, Fig. 3-4 &amp; Pl. 78, Fig. 1-2)</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 1 ex.</p><p>- P.N.duW,MamassiGourma, 30 IX 2015, lumière, 1 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2006, sous excrément humain, quelques exemplaires;</p><p>- P.N.duW, Tapoa Djerma, IX 2003, sous crottins d'éléphants,quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E20964DFC2DFB20FA06FA6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964DFC21FA3AFB6BF997.text	857E879E8E20964DFC21FA3AFB6BF997.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Bicornonthophagus mirifrons (d'Orbigny 1905)	<div><p>Bicornonthophagus mirifrons (d'Orbigny, 1905)</p><p>- [Orbigny, 1913: Nigeria: Benin (sic)].</p></div>	https://treatment.plazi.org/id/857E879E8E20964DFC21FA3AFB6BF997	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964DFF1AFF0AFDDAFE42.text	857E879E8E20964DFF1AFF0AFDDAFE42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Cleptocaccobius uniseries (d'Orbigny 1905)	<div><p>*  Cleptocaccobius uniseries (d’Orbigny, 1905) (Pl. 39, Fig. 3)</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu, 2 ex.</p><p>- P.N.Pendjari,Batia, 27 VII 2009, piège au sol, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005,sous crottins d’éléphants,quelques exemplaires;</p><p>- P. N.duW, réservede laDjona,AlphaKouara,sous bouses de zébus, quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003,sous bouses de zébus,quelques exemplaires.</p><p>-  P. N. du W, Karimama, 28 IX 2015, U. V., 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E20964DFF1AFF0AFDDAFE42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964DFF65FDD8FEB5FC58.text	857E879E8E20964DFF65FDD8FEB5FC58.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Diastellopalpus tridens (Fabricius 1781)	<div><p>Diastellopalpus tridens (Fabricius, 1781)</p><p>(Pl. 39, Fig. 4 &amp; Pl. 40, Fig. 1-2)</p><p>- Attogon, forêt de Niaouli, 27 IV 2012,sous excrément, 1 ex., 24 X 2019, 1 ex.</p><p>-   Dan, bord du Zou, 17 V 2013, 2 ex.</p><p>- Bassila, Tchétou, 17 VI 2016, sous bouse de zébu, 1 ex.</p><p>- Pénéssoulou, forêtde Pénélan, 15 XI 2012, sous excrément humain, 10 ex.; 22 V 2013,sous excrément humain, 2 ex.; 26 V 2013, sous excrément humain, 8 ex.; 16 VI 2016, sous excrément humain, 5 ex.</p><p>- P. N. du W, bordde l’Alibori, petite forêt galerie, IX 2003, sous excrément humain, quelques exemplaires.</p><p>- [Orbigny, 1913: Torricada (repris dans Josso &amp; Prévost, 2000)].</p><p>Remarque: on trouve à Pénéssoulou la forme scotias Thomson, 1858 en mélange avec la forme typique. La forme niger Kolbe, 1887 est également présente au Bénin.</p></div>	https://treatment.plazi.org/id/857E879E8E20964DFF65FDD8FEB5FC58	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964DFF65FBCAFF68FB4F.text	857E879E8E20964DFF65FBCAFF68FB4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Proagoderus auratus (Fabricius 1801)	<div><p>*  Proagoderus auratus (Fabricius, 1801) (Pl. 40, Fig. 3-4 &amp; Pl. 77, Fig. 8-9)</p><p>- P. N. Pendjari, Batia, 24 VII 2009, piège au sol, 3 ex.; 18-25 VIII 2010, piège au sol, 10 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous crottins d’éléphants, et sous excrément humain, quelques exemplaires.</p><p>- P. N. du W, Tapoa Djerma, IX 2003, sous excrément humain, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E20964DFF65FBCAFF68FB4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964DFF7DFADAFF6FFA40.text	857E879E8E20964DFF7DFADAFF6FFA40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Proagoderus patinatus Marcus 1917	<div><p>*  Proagoderus patinatus Marcus, 1917</p><p>(Pl. 41, Fig. 1-2)</p><p>- P. N. Pendjari, Batia, VI 2005, bouses de zébus, quelques exemplaires; 24 VII 2009, piège au sol, 6 ex.; 26 VIII 2010, piège au sol, 4 ex.; 27 VII 2009, piège au sol, 1 ex.; 18 VIII 2010, piège au sol, 3 ex.; 09 XI 2015, dans excrément, 1 ex.</p><p>- P. N. du W, chutes de Koudou, IX 2003, sous excrément humain, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E20964DFF7DFADAFF6FFA40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964DFF6EF9D2FE70F9C0.text	857E879E8E20964DFF6EF9D2FE70F9C0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Proagoderus rarus Guerin-Meneville 1829	<div><p>Proagoderus rarus Guérin-Méneville, 1829 (Pl. 41, Fig. 3 &amp; Pl. 77, Fig. 10)</p><p>- [Orbigny, 1913: Niger, Dahomey].</p></div>	https://treatment.plazi.org/id/857E879E8E20964DFF6EF9D2FE70F9C0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E20964FFC2EF980FE43FE67.text	857E879E8E20964FFC2EF980FE43FE67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Bicornonthophagus reticulatus (d'Orbigny 1902)	<div><p>Bicornonthophagus reticulatus (d'Orbigny, 1902) (Pl. 43, Fig. 1-2 &amp; Pl. 78, Fig. 3)</p><p>- Pénéssoulou, forêt de Pénélan, 23V  2013,U.V., 1 ex.; 28V  2013,U.V., 1 ex.; 16 VI  2016,U.V., 2 ex.</p><p>- Pénéssoulou, centre-ville, 17 VI 2013, lumière, 1 ex.</p><p>- P.N.Pendjari, Batia, 18 VIII2010,lumière, 4 ex.; 24 VIII2010,piège à crevettes, 2 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26 VIII 2010, sous bouse de buffle, 11 ex.</p><p>- P.N.Pendjari, hôtel Pendjari, VI 2005,sous crottins d’éléphants,quelques exemplaires.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 3 ex.</p><p>- P.N.duW,réservedelaDjona,AlphaKouara,sous bouses de zébus, quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003, sous bouses de zébus, quelques exemplaires.</p><p>- [Hautier et al., 2003: Ina]</p><p>Digitontophagus eucata Génier, 2017</p><p>(Pl. 43, Fig. 3 &amp; Pl. 78, Fig. 4)</p><p>- P. N. Pendjari, Batia, 24 VII 2009, 1 ex. (Paratype).</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, piège lumineux, quelques exemplaires.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 14 ex.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 33 ex.</p><p>- P.N.duW,PointTriple, IX2003,piègelumineuxetbousesdezébus,quelquesexemplaires.</p><p>- [Genier &amp; Moretto, 2017: Batia].</p></div>	https://treatment.plazi.org/id/857E879E8E20964FFC2EF980FE43FE67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E22964FFF7BFE32FE48FC78.text	857E879E8E22964FFF7BFE32FE48FC78.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Digitonthophagus falciger Genier 2017	<div><p>Digitonthophagus falciger Génier, 2017</p><p>(Pl. 43, Fig. 4 &amp; Pl. 78, Fig. 5)</p><p>- Lokossa, 2008, sous bouse de zébu, 1 ex.</p><p>-  Attogon, forêt de Niaouli, 22 XI 2014, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon,  28 IV 2013, U. V., 7 ex.; 06 VI  2016, U.V., 4 ex.;  16 XI 2017, U. V., 1 ex.; 15 V  2018, U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 23 V  2013, U.V., 1 ex.; 27 V  2013, U.V., 1 ex.</p><p>-  Pénéssoulou, centre-ville, 22 V 2013, U. V., 2 ex.; 17 VI 2013, lumière, 1 ex.</p><p>-  P.N.Pendjari, 10 XI 2012, U. V., 1 ex.</p><p>-  P.N.Pendjari, Tanguita, Batia, 08 XI 2015, U. V., 1 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 4 ex.</p><p>- P.N.duW,MamassiGourma, 30 IX 2015, lumière, 3 ex.</p><p>- [Genier &amp; Moretto, 2017: Agoué].</p></div>	https://treatment.plazi.org/id/857E879E8E22964FFF7BFE32FE48FC78	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E229641FF78FC2AFD12F90A.text	857E879E8E229641FF78FC2AFD12F90A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Digitonthophagus fimator Genier 2017	<div><p>Digitonthophagus fimator Génier, 2017 (Pl. 44, Fig. 1 &amp; Pl. 78, Fig. 6)</p><p>- Grand Popo, 22 II 2008, sous bouse zébu, 1 ex. -   Houéyogbé, forêt de Houéyogbé ,  02 IV 2013,U. V., 1 ex.;  23 IV 2013, U. V., 1 ex.; 25 VI  2016,U.V., 2 ex.; 12 VIII2007,sous bouse zébu, 2 ex.</p><p>- Lokossa, Djetokpa, 12 VIII 2007, boeuf, 1 ex.</p><p>- Lokossa, 2008, sous excrément de zébu, 1 ex.</p><p>-  Lokossa, Houédémé, 16 X 2017, U. V., 1 ex.</p><p>- Attogon, forêt de Niaouli,  18 IV 2012, U. V., 1 ex.;  29 IV 2013, U. V., 1 ex.;  22 XI 2014, U. V., 1 ex.; 14 VI  2016, U.V., 1 ex.; 18 VI  2016, U.V., 2 ex.;  03 II 2017, U. V., 1 ex.;  02 II 2017, U. V., 1 ex.</p><p>- Zogbodomey, forêt de la Lama,  26 IV 2013, U. V., 4 ex.; 15 V  2013, U.V., 1 ex.</p><p>- Bohicon, quartier Soglogon,  20 IV 2012, U. V., 1 ex.;  25 IV 2013, U. V., 1 ex.;  26 IV 2013, U. V., 1 ex.;  27 IV 2013, U. V., 4 ex.; 11 V  2013, U.V., 2 ex.;  15 XI 2014,U. V., 1 ex.;  18XI 2014,U. V., 1 ex.; 06 VI  2016,U.V., 3 ex.; 15 V  2018, U.V., 5 ex.</p><p>-   Dan, bord du Zou, 17 V  2013, U. V., 1 ex.</p><p>- Savalou, Kpataba montagne, 27 VI  2016, U.V., 2 ex.</p><p>-  Bassila, Manigri, stade de foot, 25 I 2017, U. V., 1 ex.</p><p>- Bassila, Tchétou, 17 VI 2016, sous bouse de zébu, 1 ex.</p><p>-  Bassila, Manigri, 15 I 2017, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 29 X 2011, lumière, 31 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 25 V 2013, U. V., 3 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 24 VIII 2010, sous excrément de zébu, 1 ex.; 12 V  2013, U.V., 2 ex.; 23 V  2013, U.V., 6 ex.; 27 V  2013, U.V., 51 ex.; 28 V  2013, U.V., 5 ex.; 02 VII  2013, U.V., 2 ex.; 03VII  2013, U.V., 1 ex.; 25 VII  2013, U.V., 1 ex.; 15 VI  2016, U.V., 1 ex.; 16 VI  2016, U.V., 4 ex.</p><p>- Pénéssoulou, centre-ville, 22 V 2013, sous excrément, 7 ex.; 17 VI 2013, sous excrément, 4 ex.</p><p>- P.N.Pendjari,Batia, 27 IX 2009, 1 ex. - P. N. Pendjari, Batia, Camp Numi, 24 VIII 2010, lumière, 2 ex.;  08 X 2013, U. V., 1 ex.;  08 XI 2015, U. V., 1 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 15 VII 2009,sous bouse éléphant, 3 ex.; 26 VII 2009, sous bouse éléphant, 3 ex. (paratypes).</p><p>- [  Genier &amp; Moretto, 2017: Batia; forêt de Niaouli, Attogon, 1 V 2013, 5 paratypes; Batia, 27 VII 2009, 1 paratype; Alafiarou, forêt de Wari-Maro; Pénéssoulou, forêt de Pénélan, 23 V 2013, 21 paratypes; Saramanga, 24 V 2013, 1 paratype;  Forêt de Houéyogbé, 8 V 2013, 4 paratypes; Lokossa, 24 III 2006 et 12 VIII 2007, 14 paratypes; Environs de Porto Novo; Bohicon, quartier Soglogon, 14 et 18 IV 2012, 10 paratypes]  .</p><p>*  Euonthophagus complexe carbonarius (Pl. 44, Fig. 2-3)</p><p>- Houéyogbé, forêt de Houéyogbé,  02 IV 2013, U. V., 1 ex.;  23 XI 2015, U. V., 2 ex.;  28 XII 2016, U. V., 2 ex.</p><p>- Athiémé, bord du Mono,  29 XII 2016, U. V., 3 ex.;  02 I 2017, U. V., 3 ex.;  10 II 2017, U. V., 12 ex.</p><p>- Tokay,Lobogo, 14 VI  2013, U.V., 1 ex.</p><p>-   Kpoba, bord du Mono, 02 I 2017, U. V., 2 ex.</p><p>- Zogbodomey, forêt de Lokoli, 05 V  2013, U.V., 1 ex.;  11 XII 2014,U. V., 2 ex.</p><p>- Bohicon, quartierSoglogon,  14 X 2017, U. V., 1 ex.;  29XI 2017, U. V., 1 ex.;  08 II 2018, U. V., 1 ex.</p><p>-  Bohicon, Pogba, 02 I 2017, U. V., 1 ex.</p><p>- Savalou, Kpataba montagne, 17 VI  2016, U.V., 7 ex.;  13 XII 2016, U. V., 5 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 29 XI 2011, lumière, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 13 X 2012, sous bouse de zébu, 1 ex.</p><p>- Pénéssoulou, centre-ville, stade de foot,  13 X 2012, U. V., 10 ex.;  18 X 2012, U. V., 3 ex.; 24 V  2013, U.V., 9 ex.; 22 V  2013, U.V., 41 ex.; 17 VI  2013, U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 27 V  2013, U.V., 1 ex.; 02 VII  2013, U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénéssoulou, 12 V 2013, sous excrément, 2 ex.</p><p>- Pénéssoulou, zone humide, sortie du village, 23 II 2020, 1 ex.,sous bouse de zébu.</p><p>-  Djougou, forêtde Sérou, 10 XI 2015, U. V., 1 ex.; Kouandé, antenne ORTB,  06 XI 2015, U. V., 1 ex.</p><p>- P.N.Pendjari,Batia,CampNumi,  10XI 2012,U. V., 1 ex.;  03XI2015,U. V., 7 ex.;  07 XI 2015,U. V., 1 ex.;  08 XI 2015, U. V., 3 ex.;  26 I 2017,U. V., 2 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005,sousexcrémenthumain,quelquesexemplaires; 26 VIII 2010,sous bouse de buffle, 1 ex.</p><p>- P.N.Pendjari,Batia, 29VII 2009,piège au sol, 5 ex.; 29IX2009,piègeausol, 1 ex.; 18VIII 2010,lumière, 1 ex.</p><p>-  P.N.Pendjari, Batia, CampNumi, 08 XI 2015,U. V., 4 ex.;  09 XI 2015,U. V., 3 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, sous bouse éléphant, 4 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 7 ex.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 23 ex.</p><p>- P.N.duW,réservedelaDjona,AlphaKouara, IX 2003, sous bouses de zébus, quelques exemplaires.</p><p>- P.N.duW,pointtriple, IX 2003, sous excrément humain,quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003, piège lumineux, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E229641FF78FC2AFD12F90A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2C9641FF34F900FF7FF818.text	857E879E8E2C9641FF34F900FF7FF818.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furconthophagus monforti Josso & Prevost 2006	<div><p>Furconthophagus monforti Josso &amp; Prévost, 2006</p><p>(Pl. 44, Fig. 4; Pl. 45, Fig. 1 &amp; Pl. 78, Fig. 7-8)</p><p>- [P. N. duW, 7km de Koudouendirection deKondio, IX 2003, sousexcrément de Mustelidés, quelques exemplaires; P. N. du W, falaise de Tansarga, IX 2003, piège lumineux,quelques exemplaires].</p><p>- [Josso &amp; Prévost, 2006: Tanguiéta, P. N. de la Pendjari, Tanougou; hôtel Pendjari].</p></div>	https://treatment.plazi.org/id/857E879E8E2C9641FF34F900FF7FF818	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2C9641FC0DFDE3FB1EFBE7.text	857E879E8E2C9641FC0DFDE3FB1EFBE7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furconthophagus ulula (Balthasar 1966)	<div><p>*  Furconthophagus ulula (Balthasar, 1966) (Pl. 45, Fig. 4 &amp; Pl. 46, Fig. 1)</p><p>-   Dan, bord du Zou, 10 XII 2014, U. V., 1 ex.</p><p>-  Djidja, forêt d’Atchérigbé, 14 II 2017, U. V., 1 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 3 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 22V  2013,U.V., 2 ex.; 23V  2013,U.V., 2 ex.; 25V  2013,U.V., 2 ex.; 28V  2013,U.V., 1 ex.; 04 VII  2013,U.V., 1 ex.; 15VI  2016,U.V., 1 ex.</p><p>- Pénéssoulou, stade de foot, 22 V  2013, U.V., 3 ex.; 17 VI  2013, U.V., 1 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, piège lumineux et sous excrément humain, quelques exemplaires.</p><p>- P.N.duW,pistedeKérémou-Koudou, IX 2003,sous bouses de zébus, quelques exemplaires.</p><p>- P. N. du W, chutes de Koudou, IX 2003, sous excrément humain, quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003, piège lumineux, quelques exemplaires.</p><p>-   Tanguiéta, Atakora, IX 2003, dans fleur fanée  .</p></div>	https://treatment.plazi.org/id/857E879E8E2C9641FC0DFDE3FB1EFBE7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2C9641FF06F80AFC39FDB0.text	857E879E8E2C9641FF06F80AFC39FDB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furcontophagus flaviclava (d'Orbigny 1902)	<div><p>*  Furcontophagus flaviclava (d'Orbigny, 1902) (Pl. 45, Fig. 2-3 &amp; Pl. 78, Fig. 9)</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, sous bouse de zébu, 5 ex.; 18 VI 2009, lumière, 5 ex.</p><p>- Lokossa, 01 III 2006, fauchage, 2 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 30 V 2012, sous bouse zébu, 1 ex.</p><p>-  Kouandé, antenne ORTB, 05 XI 2015,  U. V., 1 ex.</p><p>- P.N.Pendjari,Batia, 24 VIII 2010, lumière, 1 ex.; 26 VIII 2010,sous bouse de zébu sèche, 2 ex.</p><p>- P.N.duW, réserve de la Djona, mare 24, Alpha Kouara, IX 2003,sous excrément humain et sous excrément de babouins, quelques exemplaires.</p><p>- P.N.duW,chutesdeKoudou, IX 2003,sous poisson pourri,quelques exemplaires.</p><p>-   Tanguiéta, Atakora, IX 2003, sous fleur fanée et sous excrément de babouins, quelques exemplaires  .</p></div>	https://treatment.plazi.org/id/857E879E8E2C9641FF06F80AFC39FDB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2C9640FC23FBB2FD10FCC7.text	857E879E8E2C9640FC23FBB2FD10FCC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furconthophagus variegatus (Fabricius 1798)	<div><p>*  Furconthophagus variegatus (Fabricius, 1798)</p><p>(Pl. 46, Fig. 2-3 &amp; Pl. 78, Fig. 10)</p><p>- Ouidah, forêt de Pahou, 09 V  2013, U.V., 3 ex.;  21 XII 2014, U. V., 1 ex.;  22 XI 2015, U. V., 1 ex.;  05 I 2017, U. V., 2 ex.</p><p>- Houéyogbé, forêt de Houéyogbé,  02 IV 2013, U. V., 1 ex.; 08 V  2013, U.V., 1 ex.; 01 VI  2013, U.V., 2 ex.;  01 XI 2013, U. V., 1 ex.;  21 X 2014, U. V., 1 ex.;  23 XI 2015, U. V., 2 ex.;  24 XI 2015, U. V., 1 ex.; 16 VI  2016, U.V., 1 ex.; 25 VI  2016, U.V., 2 ex.;  28 XII 2016, U. V., 1 ex.</p><p>-   Athiémé, bord du Mono, 03 I 2017, U. V., 5 ex.</p><p>- Tokay, Lobogo, 14 VI  2013, U.V., 1 ex.</p><p>- Zinvié, vallée du Sitatunga, 18 V  2016, U.V., 1 ex.; 19 V  2016, U.V., 4 ex.; 26 V  2016,U.V., 8 ex.;  13 II 2019, U. V., 1 ex.</p><p>- Lokossa, 01 III 2006, 3 ex.</p><p>- Lokossa, quartier Agonvé, 24 III 2006, sous excrément humain, 2 ex.;  27 XII 2016, U. V., 1 ex.</p><p>- Attogon, forêt de Niaouli,  11 X 2014, U. V., 1 ex.;  18 XI 2015,U. V., 1 ex.; 25V  2016,U.V., 2 ex.</p><p>-  Attogon, Niaouli, 27 XII 2018, U. V., 1 ex.</p><p>-  Pobé, forêt de Pobé, 01 XII 2014, U. V., 1 ex.;  21 XII 2014, U. V., 1 ex.</p><p>-  Kotokpa, 29 XI 2017, U. V., 1 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 11 XI 2014,U. V., 1 ex.</p><p>-   Zogbodomey, Domé village, 13 XII 2014, U. V., 17 ex.</p><p>- Bohicon, quartier Soglogon, 15VI 2009,lumière, 3 ex.;  27IV 2013,U. V., 1 ex.;  28 IV 2013,U. V., 5 ex.; 11V  2013,U.V., 10 ex.; 16V  2013,U.V., 14 ex.;  19IX2014,U. V., 1 ex.;  19X 2014, U. V., 1 ex.;  11XI 2014,U. V., 6 ex.;  14XI2014,U. V., 4 ex.;  15 XI 2014, U. V., 2 ex.;  18 XI 2014,U. V., 3 ex.;  19XI2014,U. V., 12 ex.;  20XI 2014,U. V., 1 ex.;  21 XI 2014,U. V., 2 ex.;  22XI 2014,U. V., 9 ex.;  23 XI 2014,U. V., 4 ex.;  24XI 2014,U. V., 6 ex.;  26XI 2014,U. V., 5 ex.; 06VI  2016,U.V., 7 ex.;  19XI2016,U. V., 10 ex.;  23X 2017, U. V., 1 ex.;  08XI2017,U. V., 1 ex.;  10XI 2017,U. V., 1 ex.;  11XI2017,U. V., 3 ex.;  12 XI 2017,U. V., 1 ex.;  21XI2017,U. V., 2 ex.;  23XI2017,U. V., 5 ex.;  25XI 2017,U. V., 1 ex.;  12XII 2017,U. V., 4 ex.;  11 I 2018,U. V., 2 ex.; 15V  2018,U.V., 1 ex.; 16V  2018,U.V., 9 ex.; 17V  2018,U.V., 4 ex.; 03 VI  2018,U.V., 1 ex.;  01 X 2019,U. V., 1 ex.</p><p>- Savalou, Kpataba montagne, 17 VI  2016, U.V., 21 ex.</p><p>- Djougou, forêt de Sérou, 09 V  2013, U.V., 3 ex.;  23 I 2017, U. V., 8 ex.;  01 X 2019, U. V., 2 ex.;  20 X 2019, U. V., 4 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 21 X 2011, lumière, 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 24 V 2013, sous bouse de zébu, 1 ex.</p><p>- Pénéssoulou, stade de foot,  13 X 2012, U. V., 1 ex.;  16X 2012, U. V., 4 ex.;  18X 2012, U. V., 1 ex.; 22 V  2013, U.V., 43 ex.; 17 VI  2013, U.V., 1 ex.</p><p>- Pénéssoulou, forêtde Pénélan, 25 V  2013, U.V., 1 ex.; 28 V  2013, U.V., 1 ex.; 04 VII  2013, U.V., 1 ex.; 15 VI  2016, U.V., 2 ex.; 16 VI  2016, U.V., 2 ex.</p><p>-  Pénéssoulou, forêt de Gatakpal, 23 II 2020, U. V., 1 ex.</p><p>- Kouandé, antenne  ORTB, 04 XI 2015, U.  V., 1 ex.; 05 XI 2015, U.  V., 5 ex.; 06 XI 2015, U.  V., 6 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, piège lumineux, nombreux exemplaires.</p><p>- P.N.Pendjari, forêt de Bondjagou, VI 2005,piège lumineux,nombreux exemplaires.</p><p>- P.N.duW,Karimama, IX2015,lumière, 30 ex.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 4 ex.</p><p>- P.N.duW, piste de Kérémou-Koudou, IX 2003,piège lumineux,quelques exemplaires.</p><p>- P.N.duW,chutesdeKoudou, IX 2003, piège lumineux, quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003, piège lumineux, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E2C9640FC23FBB2FD10FCC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2D9640FF08FC52FDB9FB8A.text	857E879E8E2D9640FF08FC52FDB9FB8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gonionthophagus rufonotatus (d'Orbigny 1902)	<div><p>Gonionthophagus rufonotatus (d'Orbigny, 1902) (Pl. 46, Fig.4)</p><p>- [Dierkens, 2023: Savalou, forêt de Logozohé; Pénéssoulou, forêt de Goutansoukpa; Pénéssoulou, forêt de Saramanga; Pénéssoulou, forêt de Pénélan; Kouandé, antenne ORTB; Monts-Kouffé].</p></div>	https://treatment.plazi.org/id/857E879E8E2D9640FF08FC52FDB9FB8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2D9640FC08FF0FFAFEFDB7.text	857E879E8E2D9640FC08FF0FFAFEFDB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gonionthophagus stehliki (Balthasar 1966)	<div><p>Gonionthophagus stehliki (Balthasar, 1966) (Pl. 47, Fig.1)</p><p>Onthophagus clementianus Cambefort, 1971</p><p>-  P. N. Pendjari, Camp Numi, 09 XI 2015, U. V., 1 ex.</p><p>- P.N.Pendjari, hôtelPendjari, VI 2005, sous excrément humain et excrément de babouins, quelques exemplaires.</p><p>- P. N. du W, Tapoa Djerma, IX 2003, sous bouses de zébus, quelques exemplaires.</p><p>- [Dierkens, 2023: Kouandé, antenne ORTB; Batia, village; Batia, parc de la Pendjari; Monts Kouffè; Pobé, Ouémé].</p></div>	https://treatment.plazi.org/id/857E879E8E2D9640FC08FF0FFAFEFDB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2D9643FC6CFDFFFE21FC8F.text	857E879E8E2D9643FC6CFDFFFE21FC8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gonocyphus obliquus (Olivier 1789)	<div><p>Gonocyphus obliquus (Olivier, 1789)</p><p>(Pl. 47, Fig. 2-3)</p><p>-  Ouidah, forêt de Pahou, 06 I 2017, U. V., 1 ex.</p><p>- Zinvié, vallée du Sitatunga, 19 V  2016, U.V., 3 ex.; 26 V  2016, U.V., 2 ex.</p><p>- Attogon, forêt de Niaouli,  29 IV 2013,U. V., 2 ex.;  30 IV 2013, U. V., 2 ex.;  18 XI 2015, U. V., 2 ex.; 07 VI  2016,U.V., 3 ex.; 18 VI  2016,U.V., 2 ex.;  03 II 2017,U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 15VI  2009,U.V., 4 ex.;  20 IV 2012,U. V., 2 ex.;  28 IV 2013,U. V., 1 ex.; 11V  2013,U.V., 1 ex.; 16V  2013,U.V., 1 ex.; 06VI  2016,U.V., 2 ex.;  01 XII 2016,U. V., 1 ex.; 03 VI  2018, U.V., 2 ex.</p><p>- Dan, bord du Zou, 04 VI  2013, U.V., 3 ex.; 17 VII  2013, U.V., 1 ex.;  10 XII 2014, U. V., 8 ex.</p><p>-  Pobé, forêt de Pobé, 13 II 2017,U. V., 1 ex.</p><p>-  Djidja, forêt d’Atchérigbé, 14 II 2017, U. V., 1 ex.</p><p>- Savalou, Kpataba, montagne, 17 VI  2016, U.V., 17 ex.;  14 XII 2016, U. V., 4 ex.</p><p>- Bassila, Tchétou, 17 VI 2016, sous bouse de zébu, 5 ex. - Pénéssoulou, forêt de Goutansoukpa, 19 X 2011, lumière, 1 ex.; 21 X 2011, lumière, 10 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 22 V  2013, U.V., 1 ex.; 23 V  2013, U.V., 3 ex.; 25 V  2013, U.V., 1 ex.; 27 V  2013, U.V., 1 ex.; 28 V  2013,U.V., 3 ex.; 02 VII  2013, U.V., 5 ex.; 03 VII  2013, U.V., 1 ex.; 04 VII  2013, U.V., 3 ex.; 16 VI  2016,U.V., 5 ex.;  08 IX 2013, U. V., 1 ex.; 16 VI  2016, U.V., 1 ex.</p><p>-  Pénéssoulou, village, stade de foot, 13 X 2012, U. V., 1 ex.</p><p>- Pénéssoulou, zone humide, 23 II 2020, 1 ex. sous bouse de zébu.</p><p>-  Djougou, forêt de Sérou, 10 XI 2015, U. V., 1 ex.</p><p>-  Kouandé, antenne ORTB, 06 XI 2015,  U. V., 1 ex.</p><p>- P.N.Pendjari,Batia, 29 VII 2009, piège au sol, 2 ex.</p><p>-  P.N.Pendjari, Batia, CampNumi, 09 XI 2012,U. V., 1 ex.;  09 XI 2015,U. V., 1 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, piège au sol, 6 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, piège lumineux, quelques exemplaires.</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 6 ex.</p><p>- P. N. du W, Mamassi-Gourma, 30 IX 2015, lumière, 6 ex.</p><p>- P. N. du W, réserve de la Djona, mare, 24, IX 2003, sous crottins d’éléphants, quelques exemplaires.</p><p>- [Orbigny, 1913: cercle de Djougou-Kouandé].</p></div>	https://treatment.plazi.org/id/857E879E8E2D9643FC6CFDFFFE21FC8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2E9643FF00FC98FC3EFCEA.text	857E879E8E2E9643FF00FC98FC3EFCEA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Hamontophagus bituberculatus (Olivier 1789)	<div><p>Hamontophagus bituberculatus (Olivier, 1789) (Pl. 47, Fig. 4)</p><p>- Ouidah, forêt de Pahou, 09 V  2013, U.V., 1 ex.;  22 XI 2015, U. V., 1 ex.</p><p>- Houéyogbé, forêt de Houéyogbé, 15 VI 2009, sous excrément humain, 1 ex.;  22 X 2012, U. V., 4 ex.;  02 IV 2013,U. V., 1 ex.;  15 X 2014,U. V., 4 ex.;  16 X 2014, U. V., 3 ex.;  20 X 2014, U. V., 1 ex.;  19 XI 2014,U. V., 1 ex.;  24XI 2015,U. V., 2 ex.; 25 VI  2016, U.V., 4 ex.;  28 XII 2016, U. V., 2 ex.</p><p>- Athiémé, bord du Mono,  29 XII 2016, U. V., 3 ex.;  03 I 2017, U. V., 5 ex.;  10 II 2017, U. V., 4 ex.</p><p>- Tokay,Lobogo, 14 VI  2013, U.V., 3 ex.</p><p>- Zinvié, vallée du Sitatunga, 18 V  2016,U.V., 2 ex.; 19 V  2016, U.V., 7 ex.; 26 V  2016, U.V., 3 ex.</p><p>-  Lokossa, quartier Agonvé, 17 XII 2016, U. V., 1 ex.</p><p>-  Lokossa, 27 XII 2016, U. V., 1 ex.</p><p>- Attogon,Ayou, 26 XII 2018, 10 ex.</p><p>- Attogon, forêt de Niaouli, 02 V  2013,U.V., 1 ex.;  31 X 2014,U. V., 1 ex.;  18 XI 2015, U. V., 5 ex.; 07 VI  2016, U.V., 1 ex.; 18 VI  2016, U.V., 1 ex.; X 2016, sous bouse zébu, 1 ex.;  02 II 2017, U. V., 5 ex.;  03 II 2017, U. V., 1 ex.</p><p>-   Kpoba, bord du Mono, 02 I 2017, U. V., 1 ex.</p><p>-  Pobé, forêt de Pobé, 26 X 2014, U. V., 3 ex.;  27 X 2014, U. V., 3 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 13 V 2013, U. V., 1 ex.</p><p>-  Zogbodomey, Domé, village, 13 XII 2014, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 15 VI 2009,lumière, 5 ex.; 18 V  2012, U.V., 1 ex.;  26 IV 2013, U. V., 1 ex.;  27 IV 2013, U. V., 2 ex.;  28 IV 2013, U. V., 4 ex.; 11 V  2013, U.V., 1 ex.; 16V  2013, U.V., 1 ex.;  19 X 2014,U. V., 1 ex.;  11 XI 2014, U. V., 1 ex.;  14 XI 2014, U. V., 2 ex.;  15 XI 2014, U. V., 3 ex.;  16 XI 2014, U. V., 2 ex.;  19 XI 2014, U. V., 3 ex.;  20 XI 2014, U. V., 1 ex.;  21 XI 2014, U. V., 5 ex.;  22 XI 2014, U. V., 1 ex.;  23 XI 2014, U. V., 1 ex.;  24 XI 2014, U. V., 4 ex.;  26 XI 2014, U. V., 1 ex.; 01 VI  2015,U.V., 5 ex.; 06VI  2016, U.V., 1 ex.; 03 VI  2018,U.V., 1 ex.;  09 X 2017,U. V., 3 ex.;  12 X 2017, U. V., 6 ex.;  13 X 2017, U. V., 1 ex.;  15 X 2017, U. V., 1 ex.;  20 X 2017, U. V., 2 ex.;  31 X 2017, U. V., 1 ex.;  U. V., 12 XI 2017, 3 ex.;  26 XI 2017, U. V., 1 ex.; 16 V  2018, U.V., 2 ex.</p><p>- Za-Kpota, Allahé, 23 V  2018, U.V., 1 ex.,</p><p>- Dan, bord du Zou,  13 IV 2012,U. V., 2 ex.; 04VI  2013, U.V., 3 ex.;  10 XII 2014, U. V., 2 ex.</p><p>-  Djidja, forêt d’Atchérigbé, 14 II 2017, U. V., 2 ex.</p><p>- Savalou, Kpataba, montagne, 17 VI  2016, U.V., 2 ex.;  13 XII 2016, U. V., 9 ex.;</p><p>-  Djougou, forêt de Sérou, 01 X 2019, U. V., 5 ex.</p><p>-  Bassila, Manigri, stade de foot, 25 I 2017, U. V., 2 ex.</p><p>- Bétérou, bord de l’Ouémé,  27 I 2017, U. V., 1 ex. -   Pénéssoulou, forêt de Pénélan , 15 V  2012, U.V., 2 ex.;  13 X 2012, U. V., 11 ex.; 23V  2013, U.V., 23 ex.; 27V  2013, U.V., 3 ex.; 28 V  2013,U.V., 6 ex.; 02 VII  2013, U.V., 4 ex.; 03 VII  2013, U.V., 6 ex.; 04 VII  2013, U.V., 1 ex.; 16 VI  2016, U.V., 5 ex.;  24 I 2017, U. V., 1 ex.;  27 XI 2019, U. V., 1 ex.</p><p>- Pénéssoulou, stade de foot,  13 X 2012, U. V., 4 ex.;  16X 2012,U. V., 5 ex.;  18X 2012, U. V., 4 ex.;  04 XI 2012, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénéssoulou, sous excrément, 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 21 X 2011, lumière, 2 ex.; 29 X 2011, lumière, 21 ex.</p><p>- Pénéssoulou, zone humide,sortie du village, 23 II 2020, 1 ex., sous bouse de zébu.</p><p>-  Djougou, forêt de Sérou, 10 XI 2015, U. V., 2 ex.</p><p>- Kouandé, antenne  ORTB, 05 XI 2015, U.  V., 1 ex.; 06 X 2019, U.  V., 1 ex.</p><p>- P.N.Pendjari,Batia, 7 VII 2009, 1 ex.</p><p>- P.N.Pendjari,Batia,Camp Numi,  11 XI 2012, U. V., 2 ex.;  07 XI 2015, U. V., 1 ex.;  08 XI 2015, U. V., 3 ex.;  26 I 2017, U. V., 1 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 4 ex.</p><p>- P.N.duW, Mamassi-Gourma, 30 IX 2015, lumière, 30 ex.</p><p>- [Orbigny, 1913: Dahomey. ROGGERO et al., 2017: Attogon (Nyaouli); Bohiars; Borgou Wari-Maro; Kotonou; Moyen Dahomey Plateau de Gabuanado. HAUTIER et al., 2003: Ina, sub  Onthophagus laceratus Gerstaecker, 1871].</p></div>	https://treatment.plazi.org/id/857E879E8E2E9643FF00FC98FC3EFCEA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2E9643FCC5FCBFFA08FC4F.text	857E879E8E2E9643FCC5FCBFFA08FC4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Hyalonthophagus nigroviolaceus (d'Orbigny 1902)	<div><p>*  Hyalonthophagus nigroviolaceus (d'Orbigny, 1902) (Pl. 48, Fig. 1-2 &amp; Pl. 78, Fig. 11-12)</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005,piège lumineux, quelques exemplaires.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2005, piège lumineux,quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E2E9643FCC5FCBFFA08FC4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2E9643FCDFFBDAFBBFFA87.text	857E879E8E2E9643FCDFFBDAFBBFFA87.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Hyalonthophagus pseudoalcyon (d'Orbigny 1913)	<div><p>*  Hyalonthophagus pseudoalcyon (d'Orbigny, 1913)</p><p>(Pl. 48, Fig. 3-4 &amp; Pl. 79, Fig. 1-3)</p><p>- P. N. Pendjari, Batia, 27 VII 2009, piège au sol, 2 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, piège lumineux, quelques exemplaires.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2005, piège lumineux,quelques exemplaires.</p><p>*  Micronthophagus depressipennis (Cambefort, 1975) (Pl. 49, Fig. 1)</p><p>- Bohicon, 08 VI  2018, U.V., 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E2E9643FCDFFBDAFBBFFA87	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2E9643FC33FA12FA97F980.text	857E879E8E2E9643FC33FA12FA97F980.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Micronthophagus nigriceps (Raffray 1877)	<div><p>*  Micronthophagus nigriceps (Raffray, 1877) (Pl. 49, Fig. 3)</p><p>- P. N. du W, Mamassi Gourma, 30 IX 2015, lumière, 3 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E2E9643FC33FA12FA97F980	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2E9643FC22FA92FA97FA00.text	857E879E8E2E9643FC22FA92FA97FA00.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Micronthophagus ochreatus (d'Orbigny 1897)	<div><p>*  Micronthophagus ochreatus (d’Orbigny, 1897)</p><p>(Pl. 49, Fig. 2)</p><p>- P. N. du W, Mamassi Gourma, 30 IX 2015, lumière, 3 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E2E9643FC22FA92FA97FA00	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2E9643FCE8F917FAB1F882.text	857E879E8E2E9643FCE8F917FAB1F882.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Parascatonomus (Morettoeus) chloroderus (d'Orbigny 1905)	<div><p>Parascatonomus (Morettoeus) chloroderus (d’Orbigny, 1905) (Pl. 50, Fig. 1 &amp; Pl. 79, Fig. 5)</p><p>- [Hautier et al., 2003: Ina] Présence possible, à confirmer.</p></div>	https://treatment.plazi.org/id/857E879E8E2E9643FCE8F917FAB1F882	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2E9643FCF3F897FA9BF7B2.text	857E879E8E2E9643FCF3F897FA9BF7B2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Parascatonomus (Morettoeus) imbellis (d'Orbigny 1905)	<div><p>Parascatonomus (Morettoeus) imbellis (d’Orbigny, 1905)</p><p>(Pl. 70, Fig. 4)</p><p>- [Hautier et al., 2003:Ina, sub  Onthophagus planaticeps].  Parascatonomus (Morettoeus) planaticeps (d’Orbigny, 1913) est une espèce décrite du Congo, non signaléed’Afrique de l’Ouest.Confusionprobable avec  Parascatonomus (Morettoeus) imbellis, dont la présence au Bénin est possible.</p></div>	https://treatment.plazi.org/id/857E879E8E2E9643FCF3F897FA9BF7B2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2E9643FCFAF997FAF3F902.text	857E879E8E2E9643FCFAF997FAF3F902.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Parascatonomus (Morettoeus) maculatus (Fabricius 1801)	<div><p>Parascatonomus (Morettoeus) maculatus (Fabricius, 1801)</p><p>(Pl. 49, Fig. 4 &amp; Pl. 79, Fig. 4)</p><p>- [Orbigny, 1913: Haut Dahomey, région de Kandy].</p></div>	https://treatment.plazi.org/id/857E879E8E2E9643FCFAF997FAF3F902	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2F9642FF09FF0AFDD0FE3F.text	857E879E8E2F9642FF09FF0AFDD0FE3F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus chrysoderus (d'Orbigny 1905)	<div><p>Tiaronthophagus chrysoderus (d’Orbigny, 1905) (Pl. 71, Fig. 1)</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, piège lumineux et excrément de Mustelidés, quelques exemplaires.</p><p>- P.N.duW,chutesdeKoudou, IX 2003,dans poisson pourri,quelques exemplaires.</p><p>- [Roggero et al., 2019: E. of Kandi, around Saa].</p></div>	https://treatment.plazi.org/id/857E879E8E2F9642FF09FF0AFDD0FE3F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2F9642FF1DFE67FE94FC42.text	857E879E8E2F9642FF1DFE67FE94FC42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus curtipilis (d'Orbigny 1905)	<div><p>Tiaronthophagus curtipilis (d'Orbigny, 1905)</p><p>(Pl. 50, Fig. 2-3 &amp; Pl. 79, Fig. 6)</p><p>Onthophagus altidorsis d'Orbigny, 1905</p><p>- Forêt de Houéyogbé, 22 VIII 2011, sous excrément, 1 ex.; 23 VIII 2011, sous excrément humain, 1 ex.;  15 X 2014, U. V., 1 ex.;  16 X 2014, U. V., 2 ex.;  24 XI 2015, U. V., 1 ex.</p><p>-   Athiémé, bord du Mono, 31 V  2013, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V., 3 ex.</p><p>- Savalou, Kpataba, 17 VI  2016, U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 28 V  2013, U.V., 1 ex.; 16 VI  2016, U.V., 1 ex.</p><p>- P. N. Pendjari, Batia, 24 VIII 2010, piège crevette, 1 ex.; 26 VIII 2010, sous bouse zébu, 1 ex.</p><p>- Zogbodomey, forêt de Lokoli, 11 XI 2014, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, sous excrément humain.</p><p>- P. N. du W, chutes de Koudou, IX 2003, sous excrément humain et sous poissons pourris, quelques exemplaires.</p><p>- [Hautier et al., 2003: Ina].</p></div>	https://treatment.plazi.org/id/857E879E8E2F9642FF1DFE67FE94FC42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2F9642FC39FEAAFA4CFCE2.text	857E879E8E2F9642FC39FEAAFA4CFCE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus flexicornis (d'Orbigny 1902)	<div><p>Tiaronthophagus flexicornis (d'Orbigny, 1902)</p><p>(Pl. 51, Fig. 3-4 &amp; Pl. 79, Fig. 8)</p><p>- Zogbodomey, forêt de la Lama, 03 VI  2013, U.V., 1 ex.; 06 VI  2013, U.V., 1 ex.; 15 V  2013; U.V., 1 ex.; 05 VI  2013, U.V., 1 ex.; 06 VI  2013, U.V., 1 ex.; 09 VII  2014, U.V., 1 ex.;  17 II 2019, U. V., 1 ex.</p><p>- Zogbodomey, forêt de Lokoli,  26 IV 2013, U. V., 3 ex.; 07 VI  2013, U.V., 1 ex.;  11 XI 2014, U. V., 1 ex.</p><p>- Dan, bord du Zou, 04 VI  2013, U.V., 1 ex.; 06 VI  2013, U.V., 6 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 23 V  2013, U.V., 4 ex.; 27 V  2013, U.V., 1 ex.; 28 V  2013, U.V., 2 ex.; 15 VI  2016, U.V., 1 ex.</p><p>- P.N.Pendjari,Batia, 24 VIII 2010, piège à crevettes, 1 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 09 XI 2012, U. V., 1 ex.; 26 VIII 2010, sous bouse de buffle, 2 ex.</p><p>- P. N. du W, chutes de Koudou, IX 2003, sous excrément humain, quelques exemplaires.</p><p>- [Roggero et al., 2019: P. N. Pendjari, Tanougou et hôtel Pendjari].</p></div>	https://treatment.plazi.org/id/857E879E8E2F9642FC39FEAAFA4CFCE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2F9642FF03FBD0FF3CFB47.text	857E879E8E2F9642FF03FBD0FF3CFB47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus lamtoensis (Cambefort 1984)	<div><p>Tiaronthophagus lamtoensis (Cambefort, 1984) (Pl. 50, Fig. 4)</p><p>-  Pénéssoulou, forêt de Pénélan, 03 VII 2013, U. V., 1 ex.</p><p>- [P. N. du W, 7 km de Koudou en direction de Kondio, IX 2003, sous excrément de Mustelidés, quelques exemplaires].</p><p>- [Roggero et al., 2019: E. of Kandi, 4 km W of Bensekou; Parc Pendjari, vers Tanougou].</p></div>	https://treatment.plazi.org/id/857E879E8E2F9642FF03FBD0FF3CFB47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2F9642FCD5FF0FFA9CFEFF.text	857E879E8E2F9642FCD5FF0FFA9CFEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus liberianus (van Lansberge 1883)	<div><p>Tiaronthophagus liberianus (van Lansberge, 1883) (Pl. 51, Fig. 1-2 &amp; Pl. 79, Fig. 7)</p><p>- Zogbodomey, forêt de la Lama, 08 IX 2010, sous excrément humain, 2 ex.</p><p>- [Hautier et al., 2003: Ina. ROGGERO et al., 2019: Lama].</p></div>	https://treatment.plazi.org/id/857E879E8E2F9642FCD5FF0FFA9CFEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2F9642FCDFFCB7FA89FC0F.text	857E879E8E2F9642FCDFFCB7FA89FC0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus pendjarius (Josso & Prevost 2006)	<div><p>Tiaronthophagus pendjarius (Josso &amp; Prévost, 2006)</p><p>- [Josso &amp; Prévost, 2006: P. N. Pendjari, hôtel Pendjari].</p></div>	https://treatment.plazi.org/id/857E879E8E2F9642FCDFFCB7FA89FC0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2F9642FC2EFC18FBAFFBC8.text	857E879E8E2F9642FC2EFC18FBAFFBC8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus rougonorum (Cambefort 1984)	<div><p>Tiaronthophagus rougonorum (Cambefort, 1984) (Pl. 71, Fig. 2)</p><p>- P. N. Pendjari, vers Tanougou, VI 2005, sous excrément de babouins, quelques exemplaires.</p><p>- [Roggero et al., 2019: Tanguieta].</p></div>	https://treatment.plazi.org/id/857E879E8E2F9642FC2EFC18FBAFFBC8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2F9645FC3AFB5AFF7EFE50.text	857E879E8E2F9645FC3AFB5AFF7EFE50.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus viridiaerus (d'Orbigny 1908)	<div><p>Tiaronthophagus viridiaerus (d’Orbigny, 1908) (Pl. 52, Fig. 1-3 &amp; Pl. 79, Fig. 9-10)</p><p>- Attogon, forêt de Niaouli, 25 V 2016, lumière, 2 ex.; 14 VI  2016, U.V., 1 ex.;  03 II 2017, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, lumière, 1 ex.; 03 VI  2018,U.V., 2 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 27 V 2013, U. V., 1 ex.</p><p>- P. N. Pendjari, versTanougou, VI 2005, sous excréments de Mustelidés et de babouins, quelques exemplaires.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 3 ex.</p><p>- P.N.duW,Mammasi-Gourma 30 IX 2015, lumière, 1 ex.</p><p>- Savalou, Kpataba, montagne, 17 VI  2016, U.V., 1 ex.</p><p>- [Roggero et al., 2019: Ndali, 3 km W of Sontou; Pénéssoulou, forêt de Pénélan].</p></div>	https://treatment.plazi.org/id/857E879E8E2F9645FC3AFB5AFF7EFE50	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FC0BF99AFB61F951.text	857E879E8E289645FC0BF99AFB61F951.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus antennalis Frey 1961	<div><p>*  Onthophagus (s. l.) antennalis Frey, 1961</p><p>(Pl. 56, Fig. 1)</p><p>- P. N. Pendjari, Batia, 20/ 26 VIII 2010, piège au sol, 1 ex.</p><p>Groupe 2</p></div>	https://treatment.plazi.org/id/857E879E8E289645FC0BF99AFB61F951	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FC3FF8C0FB38F810.text	857E879E8E289645FC3FF8C0FB38F810.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus borassi Cambefort 1984	<div><p>*  Onthophagus (s. l.) borassi Cambefort, 1984 (Pl. 56, Fig. 2)</p><p>- Lokossa, 01 III 2006, sous bouse zébu, 4 ex.; 24 III 2006, sous bouse de zébu, 3 ex.</p><p>- Allada, Donou, 10 VI 2016, sous bouse zébu, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E289645FC3FF8C0FB38F810	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FC26F807FA1BF792.text	857E879E8E289645FC26F807FA1BF792.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus cribellum d'Orbigny 1902	<div><p>*  Onthophagus (s. l.) cribellum d'Orbigny, 1902</p><p>(Pl. 56, Fig. 3)</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu, 3 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E289645FC26F807FA1BF792	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FF69FDC2FDF1FCB7.text	857E879E8E289645FF69FDC2FDF1FCB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Phalops barbicornis van Lansberge 1883	<div><p>*  Phalops barbicornis van Lansberge, 1883 (Pl. 52, Fig. 4 &amp; Pl. 53, Fig. 1)</p><p>- P.N.Pendjari,Batia, 24 VII 2009,piège au sol, 9 ex.; 27 VII 2009,piège au sol, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26 VIII 2010, sous excrément, 1 ex.</p><p>- P.N.duW,pisteKeremou–Koudou, IX 2003,sous bouses de zébus,quelques exemplaires.</p><p>- [Barbero et al., 2003: mentionné en tant que présence probable de la moitié nord du Bénin sur la cartographie de l’espèce].</p></div>	https://treatment.plazi.org/id/857E879E8E289645FF69FDC2FDF1FCB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FF65FCE2FF65FBA7.text	857E879E8E289645FF65FCE2FF65FBA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Phalops batesi subsp. batesi von Harold 1867	<div><p>*  Phalops batesi batesi von Harold, 1867</p><p>(Pl. 53, Fig. 2-3 &amp; Pl. 79, Fig. 11)</p><p>- P. N. Pendjari, Batia, 24 VII 2009, piége au sol, 9 ex.; 27 VII 2009, piège au sol, 10 ex.; 18 VIII  2010, U.V., 1 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous crottins d’éléphants, quelques exemplaires.</p><p>- P. N. duW, chutesde Koudou, IX 2003, sous crottins d’éléphants, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E289645FF65FCE2FF65FBA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FF45FBF0FE0BFB67.text	857E879E8E289645FF45FBF0FE0BFB67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Phalops fimbriatus (Klug 1835)	<div><p>*  Phalops fimbriatus (Klug, 1835)</p><p>(Pl. 53, Fig. 4 &amp; Pl. 79, Fig. 12)</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 3 ex.</p><p>- [Barbero et al., 2003: mentionné en tantque présence probable de la moitié nord du Bénin sur la cartographie de l’espèce].</p></div>	https://treatment.plazi.org/id/857E879E8E289645FF45FBF0FE0BFB67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FEA4FB32FEC2F9AF.text	857E879E8E289645FEA4FB32FEC2F9AF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Phalops iphis (Olivier 1789)	<div><p>Phalops iphis (Olivier, 1789)</p><p>(Pl. 54, Fig. 1)</p><p>- P.N.Pendjari,forêtdeBondjagou,surbouseéléphant, 26 VII 2009, 1 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous crottins d’éléphants, quelques exemplaires.</p><p>- P. N. du W, chutes de Koudou, IX 2003, sous crottins d’éléphants, quelques exemplaires.</p><p>- [Orbigny, 1913: Kétou. BARBERO et al., 2003; Dahomey, Kétou; Parakou Ht Dahomey].</p></div>	https://treatment.plazi.org/id/857E879E8E289645FEA4FB32FEC2F9AF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FF7EF9F8FD82F89A.text	857E879E8E289645FF7EF9F8FD82F89A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Phalops vanellus van Lansberge 1883	<div><p>*  Phalops vanellus van Lansberge, 1883 (Pl. 54, Fig. 2-3)</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu, 4 ex.</p><p>- P.N.Pendjari,Batia, 24 VII 2009,piège au sol, 16 ex.; 18 VIII2010,lumière, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005,sous crottins d’éléphants,quelques exemplaires.</p><p>- [Barbero et al., 2003: mentionné au niveau de la cartographie de l’espèce comme probablement présent dans la moitié nord du Bénin].</p></div>	https://treatment.plazi.org/id/857E879E8E289645FF7EF9F8FD82F89A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FCD4FAF8FA72FA2F.text	857E879E8E289645FCD4FAF8FA72FA2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Trichonthophagus triacanthus (Castelnau 1840)	<div><p>*  Trichonthophagus triacanthus (Castelnau, 1840)</p><p>(Pl. 55, Fig. 2-3 &amp; Pl. 80, Fig. 1)</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse zébu, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E289645FCD4FAF8FA72FA2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FC2CFA78FBC2F98A.text	857E879E8E289645FC2CFA78FBC2F98A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Trichontophagus gonzalohalffteri Moretto 2023	<div><p>Trichontophagus gonzalohalffteri Moretto, 2023 (Pl. 55, Fig. 4 &amp; Pl. 80, Fig. 2)</p><p>- [Moretto, 2023: N’Dali, forêt classée de N’Dali, 11 XI 2015, sous bouse zébu, 1 paratype].</p></div>	https://treatment.plazi.org/id/857E879E8E289645FC2CFA78FBC2F98A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FF65F8F0FBE5FBAA.text	857E879E8E289645FF65F8F0FBE5FBAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Trichontophagus juvencus (Klug 1835)	<div><p>Trichontophagus juvencus (Klug, 1835)</p><p>(Pl. 54, Fig. 4)</p><p>-  Athiémé, 03 I 2017, U. V., 1 ex.</p><p>- Lokossa, 24 III 2006, sous bouse zébu, 1 ex.</p><p>-  Houéyogbé, forêt de Houéyogbé, 23 XI 2015,U. V., 2 ex.;  24 XI 2015,U. V., 1 ex.</p><p>- Attogon, forêt de Niaouli, 07 VI  2016, U.V., 3 ex.; 14 VI  2016, U.V., 1 ex.</p><p>-  Sahé, bord du Couffo, 14 V 2013, U. V., 1 ex. -  Zogbodomey, forêt de Lokoli, 05 VI 2013, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 14 VI 2012, lumière, 1 ex.; 15 VI 2009, lumière, 2 ex.,  19 XI 2014, U. V., 2 ex.;  24 XI 2014, U. V., 3 ex.;  17 XII 2014, U. V., 8 ex.; 06 VI  2016, U.V., 15 ex.;  10 XI 2017, U. V., 1 ex.;  17 XI 2017, U. V., 1 ex.;  26 XI 2017,U. V., 1 ex.;  29 XI 2017, U. V., 1 ex.;  11 XII 2017, U. V., 1 ex.;  11 I 2018, U. V., 8 ex.; 14 VI  2018, U.V., 1 ex.</p><p>- Dan, bord du Zou, 17 V  2013, U.V., 1 ex.;  10 XII 2014, U. V., 2 ex.</p><p>-  Savalou, Kpataba, montagne, 14 XII 2016, U. V., 1 ex.</p><p>-  Savalou, Kpataba, Lycée technique, bas fond, 14 XII 2016, U. V., 2 ex.</p><p>-  Bétérou, bord de l’Ouémé, 27 I 2017, U. V., 2 ex.</p><p>- Pénéssoulou,forêt de Pénélan, 23 V  2013,U.V., 5 ex.; 25V  2013,U.V., 2 ex.; 27V  2013,U.V., 1 ex.; 28V  2013,U.V., 7 ex.; 15VI  2013,U.V., 2 ex.; 02VII  2013,U.V., 7 ex.; 03VII  2013,U.V., 2 ex.; 30V  2014,U.V., 6 ex.;  02XI 2014,U. V., 1 ex.;  30 XI 2014,U. V., 3 ex.;  01 XII 2014,U. V., 2 ex.;  02 XII 2014,U. V., 1 ex.; 16 VI  2016,U.V., 1 ex.</p><p>-  Kouandé, antenne ORTB, 05 XI 2015,  U. V., 2 ex.</p><p>- Barrage de Tchakalakou, 27 I 2017, sous bouse de zébu sèche, 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 02 XII 2014, U. V., 1 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 08 XI 2015, U. V., 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, piège lumineux, quelques exemplaires.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2005, piège lumineux,quelques exemplaires.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 26 ex.</p><p>- P. N. du W, réserve de la Djona, mare 24, Alpha Kouara, IX 2003, sous excrément humain et sous excrément de babouins, quelques exemplaires.</p><p>- P.N.duW,chutesdeKoudou, IX 2003,sous bouses de zébus,quelques exemplaires.</p><p>-   Tanguiéta, Atakora, IX 2003, sous excrément de babouins, quelques exemplaires  .</p><p>- [Orbigny, 1913: Dahomey].</p></div>	https://treatment.plazi.org/id/857E879E8E289645FF65F8F0FBE5FBAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E289645FCDFFBFFFA07FAA2.text	857E879E8E289645FCDFFBFFFA07FAA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Trichontophagus mediofuscatus (d'Orbigny 1908)	<div><p>*  Trichontophagus mediofuscatus (d'Orbigny, 1908) (Pl. 55, Fig. 1)</p><p>- Pénéssoulou, forêt de Saramanga, 24 V  2013,U.V., 3 ex.; 26 V  2013,U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 23 V  2013, U.V., 1 ex.; 27 V  2013, U.V., 1 ex.; 28 V  2013, U.V., 2 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, piège au sol, 2 ex.</p><p>- P.N.duW,pisteKérmou-Koudou, IX 2003,sousbousesdezébus,quelquesexemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E289645FCDFFBFFFA07FAA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E299644FF3EFF0AFDECFE02.text	857E879E8E299644FF3EFF0AFDECFE02.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus cupreovirens d'Orbigny 1904	<div><p>*  Onthophagus (s. l.) cupreovirens d'Orbigny, 1904 (Pl. 56, Fig. 4)</p><p>-  Houéyogbé, forêt de Houéyogbé, 23 XI 2015, U. V., 1 ex.</p><p>-  Attogon, forêt de Niaouli, 18 XI 2015, U. V., 1 ex.;  02 II 2017, U. V., 2 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 13 XI 2015, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 17 V  2013, U. V., 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 03 VII 2013, U. V., 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E299644FF3EFF0AFDECFE02	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E299644FF1CFE18FDC1FDAA.text	857E879E8E299644FF1CFE18FDC1FDAA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus ducorpsi d'Orbigny 1913	<div><p>Onthophagus (s. l.) ducorpsi d'Orbigny, 1913</p><p>- [Orbigny, 1913 (sup.): plateau de Zagnanado].</p></div>	https://treatment.plazi.org/id/857E879E8E299644FF1CFE18FDC1FDAA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E299644FF56FC60FDE6FBB0.text	857E879E8E299644FF56FC60FDE6FBB0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>*  Onthophagus (s. l.) sp. (sp. 1)</p><p>(Pl. 57, Fig. 1)</p><p>-  Pénéssoulou, forêt de Pénélan, 15 VI 2016, U. V., 2 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E299644FF56FC60FDE6FBB0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E299644FC48FF0FFB1DFEFF.text	857E879E8E299644FC48FF0FFB1DFEFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>*  Onthophagus (s. l.) sp. (sp. 2)</p><p>(Pl. 57, Fig. 2)</p><p>- Grand Popo, 22 II 2008, sous bouse de zébu, 1 ex.</p><p>- Lokossa, 18 III 2006, sous bouse de zébu, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E299644FC48FF0FFB1DFEFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E299644FC48FEAAFB61FD9A.text	857E879E8E299644FC48FEAAFB61FD9A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>*  Onthophagus (s. l.) sp. (sp. 3)</p><p>(Pl. 57, Fig. 3)</p><p>- Allada, Donou, 10 VI 2016, sous bouse de zébu, 1 ex.</p><p>- Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, piège excrément, 3 ex.</p><p>Groupe 7</p></div>	https://treatment.plazi.org/id/857E879E8E299644FC48FEAAFB61FD9A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E299644FF14FDF8FD9EFCB7.text	857E879E8E299644FF14FDF8FD9EFCB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus nodieri d'Orbigny 1908	<div><p>Onthophagus (s. l.) nodieri d'Orbigny, 1908 (Pl. 71, Fig. 3)</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous crottins d’éléphants, quelques exemplaires.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2005,sous crottins d’éléphants,quelques exemplaires.</p><p>- [Orbigny, 1913: Dahomey], BRANCO, 1991: Parahou].</p></div>	https://treatment.plazi.org/id/857E879E8E299644FF14FDF8FD9EFCB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E299644FF09FCE0FD75FC37.text	857E879E8E299644FF09FCE0FD75FC37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus wensis Josso & Prevost 2006	<div><p>Onthophagus (s. l.) wensis Josso &amp; Prévost, 2006 (Pl. 71, Fig. 4)</p><p>- [Josso &amp; Prévost, 2006: parc du W, piste Keremou-Koudou].</p></div>	https://treatment.plazi.org/id/857E879E8E299644FF09FCE0FD75FC37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E299647FC02FD8AFA94FE77.text	857E879E8E299647FC02FD8AFA94FE77.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus vinctus Erichson 1843	<div><p>Onthophagus (s. l.) vinctus Erichson, 1843 (Pl. 57, Fig. 4)</p><p>- Ouidah, forêt de Pahou, 08V  2013,U.V., 29 ex.; 09V  2013,U.V., 35 ex.; 02 VII  2013, U.V., 1 ex.;  18 X 2014, U. V., 2 ex.;  15 XII 2014, U. V., 1 ex.;  21 XII 2014, U. V., 49 ex.;  21 XI 2015,U. V., 60 ex.;  22 XI 2015, U. V., 69 ex.;  25 XI 2015,U. V., 30 ex.;  05 I 2017,U. V., 23 ex.;  06 I 2017,U. V., 3 ex.</p><p>- Houéyogbé, forêt de Houéyogbé, 22 VIII  2011, U.V., 2 ex.;  22 X 2012, U. V., 6 ex.; 01 VI  2013, U.V., 1 ex.;  02 IV 2013, U. V., 1 ex.; 08 V  2013, U.V., 7 ex.; 01 VI  2013,U.V., 14 ex.; 02VI  2013,U.V., 12 ex.;  15X 2014,U. V., 9 ex.;  16 X 2014, U. V., 11 ex.;  23 XI 2015, U. V., 2 ex.;  24 XI 2015, U. V., 9 ex.;  25 XI 2015, U. V., 1 ex.; 06 VI  2016,U.V., 24 ex.; 25VI  2016,U.V., 4 ex.; 26 VI  2016, U.V., 3 ex.;  28 XII 2016, U. V., 1 ex.;  05I 2017,U. V., 202 ex.;  14 II 2017, U. V., 1 ex.; 26VI  2018,U.V., 3 ex.;  26 XII 2016, U. V., 1 ex.</p><p>- Athiémé, bord du Mono, 31 V  2013, U.V., 13 ex.;  10 XI 2016, U. V., 1 ex.;  29 XII 2016, U. V., 2 ex.;  03 I 2017, U. V., 8 ex.;  10 II 2017, U. V., 3 ex.;  12 II 2017, U. V. 1 ex.;  10 XI 2017,U. V., 1 ex.</p><p>- Tokay, Lobogo, 14 VI  2013, U.V., 5 ex.</p><p>- Zinvié, vallée du Sitatunga, 18V  2016,U.V., 18 ex.; 19V  2016,U.V., 45 ex.; 26V</p><p>2016,U.  V., 7 ex.; 12 II 2019, U.  V., 1 ex., 14 II 2019,U.  V., 1 ex.</p><p>- Lokossa, quartier Agonvé, 01 III  2006,U.V., 2 ex.: 24 III 2006, sous bouse de zébu, 2 ex.; 15 V  2016, U.V., 1 ex.;  27XII 2016,U. V., 1 ex.;  Ouédémé-Adja, bord du Mono,  30 XII 2016,U. V., 5 ex.;  16 X 2017,U. V., 1 ex.</p><p>- Attogon, forêt de Niaouli,  01 IV 2013, U. V., 3 ex.;  02 IV 2013, U. V., 23 ex.;  26 IV 2013, U. V., 1 ex.;  29 IV 2013, U. V., 43 ex.;  30 IV 2013, U. V., 8 ex.; 18 VI  2013, U.V., 13 ex.; 26 VI  2013, U.V., 1 ex.;  30 X 2014, U. V., 1 ex.;  18 XI 2014, U. V., 1 ex.;  14 XII 2014, U. V., 1 ex.;  03 II 2015, U. V., 9 ex.;  18 XI 2015, U. V., 12 ex.; 25 V  2016, U.V., 17 ex.; 26 V  2016, U.V., 4 ex.; 07 VI  2016, U.V., 78 ex.; 14 VI  2016, U.V., 37 ex.; 18 VI  2016, U.V., 14 ex.;  02 II 2017, U. V., 1 ex.;  03 II 2017, U. V., 40 ex.;  12 II 2017, U. V., 1 ex.;  01 X 2019, U. V., 1 ex.;  24 X 2019, U. V., 2 ex.</p><p>- Attogon, forêt de Ayou, 28 XII 2018, 2 ex.</p><p>-  Adidévo, Dévé, carrière, 07 II 2017, U. V., 15 ex.</p><p>- Pobé, forêt de Pobé, 18VI  2013,U.V., 9 ex.;  26X 2014,U. V., 8 ex.;  27 X 2014, U. V., 3 ex.;  17 XII 2014, U. V., 2 ex.;  21 XII 2014,U. V., 4 ex.;  19XI 2015,U. V., 1 ex.;  20 XI 2015,U. V., 1 ex.;  13 II 2017,U. V., 5 ex.</p><p>- Za-Kpota, Allahé, 05 VI  2018, U.V., 1 ex.</p><p>- Allahé, forêt d’Allahé, 31 V  2018, U.V., 9 ex.; 05 VI  2018, U.V., 7 ex.</p><p>-  Adidevo, Dévé, carrière, 07 II 2017, U. V., 1 ex.</p><p>-  Natitingou, cascade de la Kota, 21 I 2010, U. V., 14 ex.</p><p>- Zogbodomey,forêt de la Lama, 06 VI  2013,U.V., 4 ex.;  26XI 2015,U. V., 6 ex.; 02 VI 2018, U.V., 1 ex</p><p>-  Zogbodomey, forêt de Lokoli, 14 XI 2015, U. V., 2 ex.</p><p>-   Sahé, bord du Couffo, 14 V  2013, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 15VI 2009,lumière, 4 ex.;  20IV2012,U. V., 1 ex.;  27IV 2013,U. V., 1 ex.; 11V  2013,U.V., 21 ex.; 16V  2013,U.V., 5 ex.;  27 IV 2013, U. V., 4 ex.;  28 IV 2013, U. V., 15 ex.;  11 XI 2014, U. V., 4 ex.;  14 XI 2014,U. V., 5 ex.;  15 XI 2014,U. V., 8 ex.;  16XI 2014,U. V., 2 ex.;  17XI2014,U. V., 1 ex.;  18XI 2014,U. V., 3 ex.;  19XI 2014,U. V., 5 ex.;  20XI2014,U. V., 2 ex.;  21XI 2014,U. V., 2 ex.;  23 XI 2014,U. V., 5 ex.;  24XI 2014, U. V., 4 ex.;  26 XI 2014, U. V., 2 ex.; 04VI  2016, U.V., 1 ex.; 06 VI  2016,U.V., 39 ex.; 23V  2018, U.V., 2 ex.; 03VI  2018,U.V., 16 ex.; 08VI  2018,U.V., 2 ex.; 15VI  2018,U.V., 1 ex.; 25 VI  2018,U.V., 2 ex.</p><p>- Dan, bord du Zou,  03 I 2012, U. V., 1 ex.;  20 IV 2012, U. V., 1 ex.; 17 V  2013, U.V., 8 ex.; 04 VI  2013, U.V., 25 ex.; 16VI  2014,U.V., 1 ex.;  10XII 2014, U. V., 15 ex.;  09 X 2017, U. V., 7 ex.;  10 XI 2017, U. V., 2 ex.;  12 X 2017, U. V., 1 ex.;  15 X 2017, U. V., 2 ex.;  16 X 2017,U. V., 2 ex.;  06 XI 2017,U. V., 1 ex.;  12 XI 2017,U. V., 1 ex.;  22 XI 2017, U. V., 5 ex.;  23 XI 2017,U. V., 5 ex.;  25 XI 2017,U. V., 1 ex.;  26 XI 2017, U. V., 4 ex.;  30 XI 2017,U. V., 2 ex.;  12 XII 2017, U. V., 1 ex.;  11 I 2018, U. V., 2 ex.; 01 VI  2018, U.V., 2 ex.; 03 VI  2018, U.V., 3 ex.</p><p>-  Djidja, forêt d’Atchérigbé, 14 II 2017, U. V., 9 ex.;  17 II 2017,U. V., 7 ex.</p><p>- Savalou, Kpataba, montagne, 17 VI  2016, U.V., 100 ex.;  13 XII 2016, U. V., 6 ex.</p><p>-  Savalou, Kpataba, Lycée technique, bas fond, 14 XII 2016, U. V., 5 ex.</p><p>-  Djougou, forêt de Sérou, 01 X 2019, U. V., 3 ex.;  03 X 2019, U. V., 2 ex.</p><p>-  Bassila, Manigri, stade de foot, 25 I 2017, U. V., 4 ex.</p><p>-  Bétérou, bord de l’Ouémé, 27 I 2017, U. V., 17 ex.</p><p>- Pénéssoulou, forêt de Pénélan,  13X 2012,U. V., 2 ex.;  16X 2012,U. V., 10 ex.; 17 V  2013, U.V., 15 ex.; 22 V  2013, U.V., 5 ex.; 23 V  2013, U.V., 32 ex.; 24V  2013, U.V., 1 ex.; 25 V  2013, U.V., 13 ex.; 27 V  2013,U.V., 11 ex.; 28 V  2013,U.V., 16 ex.; 17VI  2013,U.V., 8 ex.; 02VII  2013,U.V., 9 ex.; 03VII  2013,U.V., 19 ex.; 04 VII  2013, U.V., 16 ex.;  01XII2014,U. V., 6 ex.;  02XII2014,U. V., 3 ex.;  03XII2014,U. V., 5 ex.; 15VI  2016, U.V., 13 ex.; 16 VI  2016,U.V., 11 ex.;  24 I 2017,U. V., 2 ex.;  27 XI 2019, U. V., 7 ex.</p><p>- Pénéssoulou, centre-ville, lumière, 1 ex.;  stade de foot, 16 X 2012, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 21 X 2011, lumière, 7 ex.; 29 X 2011, lumière, 1 ex.</p><p>-  Pénéssoulou, forêt de Katakpal, 23 XI 2020, U. V., 12 ex.</p><p>-  Parakou, forêt de Parakou, 21 XI 2015, U. V., 3 ex.</p><p>-  Djougou, forêt de Sérou, 10 XI 2015, U. V., 2 ex.;  23 I 2017, U. V., 2 ex.</p><p>- Kouandé,antenne  ORTB, 04XI 2015,U.  V., 1 ex.; 05 XI 2015,U.  V., 3 ex.; 06 XI 2015, U.  V., 8 ex.;</p><p>- P. N. Pendjari, Camp Numi, 29 VII 2009, piège au sol, 1 ex.; 24 VIII 2010,</p><p>U.  V., 1 ex.; 11XI 2012,U.  V., 1 ex.; 07 XI 2015,U.  V., 1 ex.; 08 XI 2015, U.  V., 6 ex.; 09 XI 2015,U.  V., 2 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, 26 VIII 2010, sous excrément, 1 ex.; P. N. Pendjari, forêt de Bondjagou, 26 VII 2009, sous bouse éléphant, 2 ex.</p><p>- P. N. du W, Karimama, 18 IX 2015, lumière, 1 ex.</p><p>- P. N. du W, Mamassi-Gourma, 30 IX 2015, lumière, 1 ex.</p><p>- [Tagliaferi et al., 2012: Djougou; Attogon; env. de Cotonou; Niaouli; Zou, village Akongbere near Save; Abomey].</p></div>	https://treatment.plazi.org/id/857E879E8E299647FC02FD8AFA94FE77	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2A9647FC26FA4AFC42FA40.text	857E879E8E2A9647FC26FA4AFC42FA40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus atridorsis d'Orbigny 1902	<div><p>*  Onthophagus (s. l.) atridorsis d'Orbigny, 1902 (Pl. 58, Fig. 3 &amp; Pl. 80, Fig. 3)</p><p>- P. N. duW, chutes de Koudou, IX 2003, sous excrément humain, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E2A9647FC26FA4AFC42FA40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2A9647FCD5F9D7FB1FF8D1.text	857E879E8E2A9647FCD5F9D7FB1FF8D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>*  Onthophagus (s. l.) complexe pullus Roth, 1851</p><p>(Pl. 58, Fig. 4)</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, lumière, 3 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse zébu sèche, 2 ex.</p><p>- P.N.Pendjari,Batia, 24VII 2009,piège au sol, 1 ex.; 27VII 2009,piège au sol, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI2005,sous excrément humain,quelques exemplaires.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 2 ex.</p><p>- P.N.duW,TapoaDjerma,sous excrément d’éléphants, quelques exemplaires.</p><p>Groupe 11</p></div>	https://treatment.plazi.org/id/857E879E8E2A9647FCD5F9D7FB1FF8D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2A9647FC37F840FB39F7A8.text	857E879E8E2A9647FC37F840FB39F7A8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus lioides d'Orbigny 1902	<div><p>*  Onthophagus (s. l.) lioides d'Orbigny, 1902 (Pl. 59, Fig. 1)</p><p>- Bohicon, quartier Soglogon, 15 VI 2009, lumière, 2 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E2A9647FC37F840FB39F7A8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2A9647FCFBFE20FB61FB8F.text	857E879E8E2A9647FCFBFE20FB61FB8F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus paravinctus Tagliaferri et Moretto 2012	<div><p>Onthophagus (s. l.) paravinctus Tagliaferri et Moretto, 2012 (Pl. 58, Fig. 1)</p><p>-  Djidja, forêt d’Atchérigbé, 14 II 2017, U. V., 8 ex.</p><p>- P.N.Pendjari,Batia,Camp Numi, 08 XI 2013, 26 ex.;  03 XI 2015, U. V., 12 ex.;  11 XII 2012, U. V., 1 ex.;  26 I 2017, U. V., 15 ex.</p><p>-  P. N. Pendjari, Batia, 26 I 2017, U. V., 1 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, sous excrément humain et pièges lumineux, quelques exemplaires; 26 VIII 2010, sous bouse de buffle, 1 ex.</p><p>- P.N.Pendjari,forêtdelaBondjagou, 26 VII 2009,sous bouse éléphant, 1 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 32 ex.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 14 ex.</p><p>- P. N. du W, réserve de la Djona, IX 2003, sous bouses de zébus, quelques exemplaires.</p><p>- P.N.duW, Tapoa Djerma,sous excrément d’éléphants,quelques exemplaires.</p><p>- [Tagliaferi et al., 2012: 15 km N Tanguieta; 15 km N Kouarfa; Koutiakou].</p><p>Groupe 9</p></div>	https://treatment.plazi.org/id/857E879E8E2A9647FCFBFE20FB61FB8F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2A9647FC09FB97FB18FADA.text	857E879E8E2A9647FC09FB97FB18FADA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus picatus d'Orbigny 1902	<div><p>Onthophagus (s. l.) picatus d'Orbigny, 1902</p><p>(Pl. 58, Fig. 2)</p><p>-  Houéyogbé, forêt de Houéyogbé, 02 IV 2013, U. V., 1 ex.;</p><p>-  Pobé, forêt de Pobé, 20 XI 2015, U. V., 1 ex.;</p><p>-  Pénéssoulou, forêt de Pénélan, 15 VI 2016, U. V., 1 ex.</p><p>- [Orbigny, 1913: cercle de Djougou-Kouandé].</p><p>Groupe 10</p></div>	https://treatment.plazi.org/id/857E879E8E2A9647FC09FB97FB18FADA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2B9646FF0FFD08FD52FC9F.text	857E879E8E2B9646FF0FFD08FD52FC9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus brevipennis d'Orbigny 1902	<div><p>Onthophagus (s. l.) brevipennis d'Orbigny, 1902 (Pl. 59, Fig. 4 &amp; Pl. 80, Fig. 4)</p><p>- [Hautier et al., 2003: Ina]. Présence possible, à confirmer.</p></div>	https://treatment.plazi.org/id/857E879E8E2B9646FF0FFD08FD52FC9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2B9646FC22FCDAFA68FC22.text	857E879E8E2B9646FC22FCDAFA68FC22.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus ecopas Josso et Prevost 2006	<div><p>Onthophagus (s. l.) ecopas Josso et Prévost, 2006 (Pl. 61, Fig. 1 &amp; Pl. 80, Fig. 6)</p><p>- P. N. du W, Mamassi Gourma, 30 IX 2015, 1 ex.</p><p>- [Josso &amp; Prévost, 2006: 15 km N de Kouarfa, Bouyagnindi village].</p></div>	https://treatment.plazi.org/id/857E879E8E2B9646FC22FCDAFA68FC22	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2B9646FC06FF32FB36FEE0.text	857E879E8E2B9646FC06FF32FB36FEE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus feai d'Orbigny 1905	<div><p>*  Onthophagus (s. l.) feai d'Orbigny, 1905 (Pl. 60, Fig. 3)</p><p>- Zogbodomey, forêt de la Lama, X 2010, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E2B9646FC06FF32FB36FEE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2B9646FFD1FC87FB18FF62.text	857E879E8E2B9646FFD1FC87FB18FF62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>Onthophagus (s. l.) complexe  cupreus von Harold, 1880</p><p>(Pl. 60, Fig. 1-2)</p><p>-  Pénéssoulou, forêt de Goutansoukpa, 30 V 2012, U. V., 1 ex.</p><p>- P.N.Pendjari,Batia, 18 VIII 2010, lumière, 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26 VIII 2010, dans excrément, 1 ex.</p><p>- [Orbigny, 1913: Ouidah].</p><p>Groupe 17</p></div>	https://treatment.plazi.org/id/857E879E8E2B9646FFD1FC87FB18FF62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2B9646FF18FF0AFD03FE57.text	857E879E8E2B9646FF18FF0AFD03FE57.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus notatus d'Orbigny 1902	<div><p>*  Onthophagus (s. l.) notatus d'Orbigny, 1902 (Pl. 59, Fig. 2)</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu, 1 ex.</p><p>- P.N.Pendjari,Batia, 27 VII 2009, piège au sol, 15 ex.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005,sous excrément humain,quelques exemplaires.</p><p>P. N. du W, Karimama, 28 IX 2015, lumière, 1 ex.</p><p>- P.N.duW,réservedelaDjona, IX 2003,sous bouses de zébus,quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma,sousexcrémentd’éléphants,quelquesexemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E2B9646FF18FF0AFD03FE57	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2B9646FC12FEB7FB18FD4A.text	857E879E8E2B9646FC12FEB7FB18FD4A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus sellatus Klug 1845	<div><p>Onthophagus (s. l.) sellatus Klug, 1845</p><p>(Pl. 60, Fig. 4 &amp; Pl. 80, Fig. 5)</p><p>-  Attogon, forêt de Niaouli, 14 XII 2014, U. V., 1 ex.</p><p>-  Pobé, forêt de Pobé, 27 X 2014, U. V., 1 ex.</p><p>-   Sahé, bord du Couffo, 14 V  2013, U. V., 2 ex.</p><p>-  Bohicon, 19 IX 2014, U. V., 1 ex.</p><p>-  Pénéssoulou, centre ville, stade de foot, 13 X 2012, U. V., 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 16 VI 2016, U. V., 1 ex.</p><p>- [Orbigny, 1913: Ouidah].</p><p>Groupe 18</p></div>	https://treatment.plazi.org/id/857E879E8E2B9646FC12FEB7FB18FD4A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2B9646FF14FDC2FE03FD19.text	857E879E8E2B9646FF14FDC2FE03FD19.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus tersipennis d'Orbigny 1902	<div><p>*  Onthophagus tersipennis d'Orbigny, 1902</p><p>(Pl. 59, Fig. 3)</p><p>- P. N. du W, Karimama, lumière, 28 IX 2015, 1 ex.</p><p>Groupe 16</p></div>	https://treatment.plazi.org/id/857E879E8E2B9646FF14FDC2FE03FD19	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E2B9679FC26FC78FF68FC6F.text	857E879E8E2B9679FC26FC78FF68FC6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus latiggiber d'Orbigny 1902	<div><p>*  Onthophagus (s. l.) latiggiber d'Orbigny, 1902</p><p>(Pl. 61, Fig. 2-3 &amp; Pl. 80, Fig. 7)</p><p>-  Ouidah, forêt de Pahou, 18 X 2014, U. V., 2 ex. - Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, piège à excrément, 20 ex.;  22 X 2012, U. V., 2 ex.;  22 XI 2015, U. V., 1 ex.</p><p>- Attogon, forêt de Niaouli,  02 IV 2013, U. V., 1 ex.; 03 V  2013,U.V., 1 ex.;  15 X 2014, U. V., 1 ex.; 25 V  2016, U.V., 3 ex.; 26 V  2016, U.V., 3 ex.; 07 VI  2016, U.V., 4 ex.; 14 VI  2016, U.V., 4 ex.; 18 VI  2016, U.V., 5 ex.;</p><p>- Pobé, forêtde Pobé, 18VI  2013,U.V., 1 ex.; 20 VI 2013,à vue, 1 ex.;  27 X 2014, U. V., 1 ex. 19 XI 2015,U. V., 3 ex.;  20 XI 2015,U. V., 5 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 13 XI 2015, U. V., 1 ex.;  27 XI 2015, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V., 1 ex.</p><p>- Za-Kpota, Allahé, 30 V  2018, U.V., 1 ex.</p><p>- Savalou, Kpataba montagne, 17 VI  2016, U.V., 1 ex.</p><p>-   Parakou, forêt de  Parakou Forêt de Parakou, 11 XI 2015, U. V., 1 ex.</p><p>-  Kouandé, antenne ORTB, 05 XI 2015,  U. V., 10 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 15 V  2012, U.V., 1 ex.;  11 X 2012, U. V., 2 ex.; 28 V  2013, U.V., 3 ex.; 03 VII  2013, U.V., 1 ex.; 15 VI  2016, U.V., 1 ex.</p><p>-   Pénéssoulou village, stade de foot, 04 XI 2012, U. V., 1 ex.</p><p>- P.N.Pendjari,Batia, 24 VIII 2010, piège à crevettes, 1 ex.</p><p>- P.N. Pendjari,Batia,CampNumi,  09 XI 2012, U. V., 2 ex.;  11 XI 2012, U. V., 1 ex.;  03 XI 2015, U. V., 1 ex.;  07 XI 2015, U. V., 1 ex.;  08 XI 2015, U. V., 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari,sousbousedebuffle, 2 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 3 ex.</p><p>- P.N.duW,MamassiGourma, 30 IX 2015, lumière, 1 ex</p><p>- P.N.duW, réserve de laDjona,mare 24, IX 2003, piège lumineux, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E2B9679FC26FC78FF68FC6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E149679FC2BFF32FB8EFE07.text	857E879E8E149679FC2BFF32FB8EFE07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus bimarginatus d'Orbigny 1902	<div><p>Onthophagus (s. l.) bimarginatus d'Orbigny, 1902 (Pl. 62, Fig. 2-3 &amp; Pl. 80, Fig. 9)</p><p>- Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, piège excrément, 1 ex.; 08 V  2013, U.V., 1 ex.;  15 X 2014, U. V., 1 ex.</p><p>- P.N.Pendjari,Batia, 25 VIII 2010, lumière, 1 ex.</p><p>- [Josso, 2022: Bohicon; Batia]</p></div>	https://treatment.plazi.org/id/857E879E8E149679FC2BFF32FB8EFE07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E149679FCDAFE12FBE8FDE8.text	857E879E8E149679FCDAFE12FBE8FDE8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus callosipennis Boucomont 1930	<div><p>*  Onthophagus (s. l.) callosipennis Boucomont, 1930</p><p>(Pl. 62, Fig. 4)</p><p>- Pénéssoulou, forêt de Pénélan, 23 V  2013, U.V., 1 ex.; 02 VII  2013, U.V., 1 ex.; 16 VI  2016, U.V., 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E149679FCDAFE12FBE8FDE8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E149679FC21FDBAFA52FD42.text	857E879E8E149679FC21FDBAFA52FD42.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus ghanensis Balthasar 1966	<div><p>*  Onthophagus (s. l.) ghanensis Balthasar, 1966 (Pl. 63, Fig. 1)</p><p>-  Kouandé, antenne ORTB, 05 XI 2015,  U. V., 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26 VIII 2010, sous bouse de buffle, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E149679FC21FDBAFA52FD42	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E149679FC23FCD8FBF1FC7A.text	857E879E8E149679FC23FCD8FBF1FC7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus grandifrons d'Orbigny 1907	<div><p>Onthophagus (s. l.) grandifrons d'Orbigny, 1907</p><p>(Pl. 63, Fig. 2)</p><p>Houéyogbé, 22 VIII 2011, Sous écrément humain, 1 ex.</p><p>- Lokossa, sous bouse de zébu, 18 III 2006, 1 ex.</p><p>- [Hautier et al., 2003: Ina].</p></div>	https://treatment.plazi.org/id/857E879E8E149679FC23FCD8FBF1FC7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E149679FF3AFC38FB18FF62.text	857E879E8E149679FF3AFC38FB18FF62.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus trinominatus Goidanich 1926	<div><p>*  Onthophagus (s. l.) trinominatus Goidanich, 1926 (Pl. 61, Fig. 4; Pl. 62, Fig. 1 &amp; Pl. 80, Fig. 8)</p><p>Onthophagus rostrifer d'Orbigny, 1904</p><p>-  Houéyogbé, forêt de Houéyogbé, 25 VI 2016, U. V., 1 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 13 V 2013, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 17 VII  2013, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 28 V  2013, U.V., 3 ex.; 03 VII  2013, U.V., 2 ex.;</p><p>- Pénéssoulou, stade de foot, 22 V  2013, U.V., 1 ex.; 17 VI  2013, U.V., 1 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 1 ex.</p><p>- P.N.duW,MamassiGourma, 30 IX 2015, lumière, 1 ex.</p><p>- P.N.duW,TapoaDjerma, IX 2003, piège lumineux, quelques exemplaires.</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, piège lumineux, quelques exemplaires.</p><p>Groupe 19</p></div>	https://treatment.plazi.org/id/857E879E8E149679FF3AFC38FB18FF62	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E149678FCDEFC2FFD9EFE28.text	857E879E8E149678FCDEFC2FFD9EFE28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus mankonoensis Balthasar 1966	<div><p>*  Onthophagus (s. l.) mankonoensis Balthasar, 1966 (Pl. 63, Fig. 3-4 &amp; Pl. 80, Fig. 10)</p><p>- Tokay, Lobogo, 15 VI  2013, U.V., 1 ex.</p><p>-  Houéyogbé, forêt de Houéyogbé, 25 VI 2016, U. V., 2 ex.</p><p>- Zogbodomey, forêt de Lokoli, 13 V  2013, U.V., 1 ex.; 05 VI  2013, U.V., 1 ex.</p><p>-  Bohicon, quartier Soglogon, 14 IV 2012, U. V., 2 ex.</p><p>-   Dan, bord du Zou, 04 VI  2013, U. V., 1 ex.</p><p>- Za-Kpota, Allahé, 31 V  2018, U.V., 1 ex.</p><p>-  Savalou, Kpataba, lycée technique, bas fond, 14 XII 2016, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 06 V  2013, U.V., 1 ex.; 23 V  2013, U.V., 1 ex.; 27 V  2013, U.V., 1 ex.; 02 VII  2013, U.V., 1 ex.; 03 VII  2013, U.V., 2 ex.; 16 VI  2016, U.V., 1 ex., sous excrément, 12 V  2013, U.V., 1 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 2 ex.</p><p>- Kouandé, Antenne ORTB,  U. V., 1 ex.</p><p>- P.N.Pendjari,Batia, 24 VIII 2010, piège à crevettes, 2 ex.;</p><p>- P.N.Pendjari,Batia,Camp Numi, 08 X 2013, U.V., 1. ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26 VIII 2010, sous bouse de buffle, 44 ex.</p><p>- P.N.duW,Karimama, 30 IX 2015, lumière, 1 ex.</p><p>- P.N.duW,MamassiGourma, 30 IX 2015, lumière, 2 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E149678FCDEFC2FFD9EFE28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E159678FBA1FB8FFB18FB67.text	857E879E8E159678FBA1FB8FFB18FB67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>Onthophagus (s. l.) sp.</p><p>(Pl. 64, Fig. 4)</p><p>- P.N.Pendjari,Batia, 24 VIII 2010, piège à crevettes, 1 ex.;</p><p>Groupe 20</p></div>	https://treatment.plazi.org/id/857E879E8E159678FBA1FB8FFB18FB67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E159678FF36FE7AFF68FDD2.text	857E879E8E159678FF36FE7AFF68FDD2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus mocquerysi d'Orbigny 1902	<div><p>*  Onthophagus (s. l.) mocquerysi d'Orbigny, 1902 (Pl. 72, Fig. 1)</p><p>- P. N. du W, réserve de la Djona, mare 24, IX 2003, piège lumineux, quelques exemplaires.</p><p>- P.N. du W,TapoaDjema,chutesKoudou, IX 2003, piège lumineux, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E159678FF36FE7AFF68FDD2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E159678FF0CFD48FEA7FCBA.text	857E879E8E159678FF0CFD48FEA7FCBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus obenbergeri Balthasar 1965	<div><p>Onthophagus (s. l.) obenbergeri Balthasar, 1965</p><p>(Pl. 64, Fig. 1 &amp; Pl. 80, Fig. 11)</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005</p><p>- [Moretto &amp; Josso, 2021: Bénin].</p></div>	https://treatment.plazi.org/id/857E879E8E159678FF0CFD48FEA7FCBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E159678FF25FCEFFA0EFD10.text	857E879E8E159678FF25FCEFFA0EFD10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus semivirescens d'Orbigny 1902	<div><p>*  Onthophagus (s. l.) semivirescens d'Orbigny, 1902 (Pl. 64, Fig. 2)</p><p>-  Ouidah, forêt de Pahou, 18 X 2014, U. V., 5 ex.;</p><p>- Tokay,Lobogo, 14 VI  2013, U.V., 1 ex.;</p><p>- Houéyogbé, forêt de Houéyogbé,  02 IV 2013, U. V., 1 ex.;  15 X 2014, U. V., 1 ex.;  16 X 2014, U. V., 1 ex.;  15 X 2015,U. V., 1 ex.; 25 VI  2016,U.V., 2 ex.; 26 VI  2018, U.V., 1 ex.;</p><p>- Lokossa, 24 III 2006, sous bouse de zébu, 2 ex.</p><p>-   Lokossa, bord du Mono, U. V., 1 ex.</p><p>- Attogon, forêtde Niaouli, 24 VI  2013, U.V., 1 ex.; 14 VI  2016, U.V., 2 ex.; 18 VI  2016, U.V., 1 ex.;  12 II 2017, U. V., 1 ex.</p><p>- Pobé, forêt de Pobé, 18 VI  2013, U.V., 2 ex.; 20 VI  2013, U.V., 21 ex.;  27 X 2014, U. V., 1 ex.;  19 XI 2015, U. V., 1 ex.;  20 XI 2015, U. V., 1 ex.;  13 II 2017, U. V., 3 ex.</p><p>- Zogbodomey, forêt de la Lama, 03 VI  2013,U.V., 2 ex.;  21 X 2014, U. V., 3 ex.</p><p>- Zogbodomey, forêt de Lokoli, 05 VI  2013, U.V., 5 ex.; 08 VI  2013, U.V., 1 ex.;  05 XI 2013, U. V., 1 ex.;  23 X 2014, U. V., 1 ex.;  13 XI 2015, U. V., 1 ex.;  15 XII 2016, U. V., 1 ex.</p><p>-  Zogbodomey, Domé, village, CEG, 12 XII 2014, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon,  20 IV 2012, U. V., 1 ex.;  20 X 2014, U. V., 2 ex. -  Dan, bord du Zou, 04 VI  2013, U.V., 2 ex.; 17 V  2013, U.V., 1 ex.</p><p>-  Savalou, Kpataba, lycée technique, bas fond, 14 XII 2016, U. V., 2 ex.;</p><p>-  Djougou, forêt de Sérou, 01 X 2019, U. V., 1 ex.</p><p>-  Bétérou, bord de l’Ouémé, 21 I 2017, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 15 V  2012, U.V., 1 ex.;  11 X 2012, U. V., 2 ex.; 23 V  2013, U.V., 1 ex.; 15 VI  2016, U.V., 2 ex.; 16 VI  2016, U.V., 1 ex.</p><p>- Pénéssoulou, stade de foot, 22 V  2013, U.V., 1 ex.</p><p>-  Kouandé, antenne ORTB, 05 XI 2015,  U. V., 1 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26  VIII 2010,sous bouse de zébu, 2 ex.; VI 2005, piège lumineux, quelques exemplaires.</p><p>- P.N.Pendjari,Batia,Camp Numi,  07 XI 2012, U. V., 1 ex.;  03XI 2015,U. V., 2 ex.;  08 XI 2015, U. V., 2 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2005,piège lumineux,et danschampignons pourris,quelques exemplaires.</p><p>- P.N.Pendjari,versTanougou, VI 2005, piège lumineux, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E159678FF25FCEFFA0EFD10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E159678FC2CFD02FB39FB98.text	857E879E8E159678FC2CFD02FB39FB98.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus viridicatus d'Orbigny 1902	<div><p>*  Onthophagus (s. l.) viridicatus d'Orbigny, 1902 (Pl. 64, Fig. 3)</p><p>- Attogon, forêt de Niaouli, 25 V  2016, U.V., 1 ex.; 14 VI  2016, U.V., 3 ex.; 18 VI  2016, U.V., 1 ex.;  03 II 2017, U. V., 1 ex.</p><p>- Bohicon, Quartier Soglogon, 15 VI 2009, lumière, 1 ex.</p><p>- Savalou, Kpataba, montagne, 17 VI  2016, U.V., 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 27 XI 2019, U. V., 1 ex.</p><p>-  P.N.Pendjari, Batia, Camp Numi, 03 XI 2015, U. V., 1 ex.</p><p>-  Ouidah, forêt de Pahou, 5 I 2017, U. V., 1 ex.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 2 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E159678FC2CFD02FB39FB98	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E15967BFCD0FAC8FD57FEE0.text	857E879E8E15967BFCD0FAC8FD57FEE0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus margaritifer d'Orbigny 1898	<div><p>*  Onthophagus (s. l.) margaritifer d’Orbigny, 1898</p><p>(Pl. 65, Fig. 1)</p><p>Onthophagus inermifrons d'Orbigny, 1902</p><p>- P. N. du W, Karimama, 28 IX 2015, lumière, 4 ex.</p><p>Groupe 22</p><p>*  Onthophagus (s. l.) circumdatus d'Orbigny, 1913 (Pl. 65, Fig. 2 &amp; Pl. 80, Fig. 12)</p><p>- Tanguiéta,chaîne de l’Atakora, IX 2003, dans fleur fané, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E15967BFCD0FAC8FD57FEE0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E16967BFF11FB87FEB8FA8A.text	857E879E8E16967BFF11FB87FEB8FA8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus fimetarius Roth 1851	<div><p>*  Onthophagus (s. l.) fimetarius Roth, 1851 (Pl. 65, Fig. 4)</p><p>-   Dan, bord du Zou, 10 XII 2014, U. V., 1 ex.</p><p>-  P. N. Pendjari, Batia, Camp Numi, 07 XI 2015, U. V., 1 ex.</p><p>- P.N.duW,chutesdeKoudou, IX 2003, piège lumineux, quelques exemplaires.</p><p>- P. N. du W, Tapoa Djerma, IX 2003, sous excrément humain et sous crottins d’éléphants, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E16967BFF11FB87FEB8FA8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E16967BFF68FA80FAF5FEF8.text	857E879E8E16967BFF68FA80FAF5FEF8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus lobi Cambefort 1984	<div><p>*  Onthophagus (s. l.) lobi Cambefort, 1984</p><p>(Pl. 66, Fig. 1-2 &amp; Pl. 81, Fig. 2)</p><p>- Pénéssoulou, forêt de Pénélan, 28 V  2013, U.V., 1 ex.; 03 VII  2013, U.V., 1 ex.; 15 VI  2016, U.V., 1 ex.; 16 VI  2016, U.V., 1 ex.</p><p>-  Pénéssoulou, forêt de Saramanga, 24 V 2013, U. V., 1 ex.; 25 V 2013, sous bouse sèche de zébu, 1 ex.</p><p>- P.N.duW,Karimama, IX 2015, lumière, 1 ex.</p><p>- P.N.duW,Mamassi-Gourma, 30 IX 2015, lumière, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E16967BFF68FA80FAF5FEF8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E16967BFC2DFEAAFB31FDE2.text	857E879E8E16967BFC2DFEAAFB31FDE2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus lutaticollis d'Orbigny 1907	<div><p>*  Onthophagus (s. l.) lutaticollis d'Orbigny, 1907 (Pl. 66, Fig. 3-4 &amp; Pl. 81, Fig. 3-5)</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu, 1 ex.</p><p>- P.N.Pendjari,Batia, 27 VII 2009, piège au sol, 3 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26 VIII 2010, sous bouse de zébu, 1 ex.</p><p>- P.N.duW,pisteKérémou-Koudou, IX 2003, sous bouses de zébus.</p><p>-   Tanguiéta, Atakora, IX 2003, sous fleur fanée, 1 ex.</p></div>	https://treatment.plazi.org/id/857E879E8E16967BFC2DFEAAFB31FDE2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E16967BFC28FDB8FA08FC5F.text	857E879E8E16967BFC28FDB8FA08FC5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus mucronifer d'Orbigny 1905	<div><p>*  Onthophagus (s. l.) mucronifer d'Orbigny, 1905</p><p>(Pl. 67, Fig. 1)</p><p>-  Houéyogbé, forêt de Houéyogbé, 02 VI 2013, U. V., 2 ex.</p><p>- Lokossa, 24 III 2006, sous bouse zébu, 2 ex.</p><p>- Bohicon, quartier Soglogon,  27 IV 2013, U. V., 3 ex.;  28 IV 2013, U. V., 1 ex.; 24 V  2018, U.V., 1 ex.; 15 VI  2018, U.V., 1 ex.</p><p>-   Dan, bord du Zou, 17 VII  2013, U. V., 1 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 28 V 2013, U. V., 1 ex.</p><p>- P.N.Pendjari,hôtel Pendjari, VI 2005, piège lumineux, quelques exemplaires; 26 VIII 2010, sous bouse de buffle, 1 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2005, piège lumineux,quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E16967BFC28FDB8FA08FC5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E16967BFC26FBC8FBC7FB18.text	857E879E8E16967BFC26FBC8FBC7FB18.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus proximus d'Orbigny 1908	<div><p>*  Onthophagus (s. l.) proximus d'Orbigny, 1908 (Pl. 67, Fig. 2-3 &amp; Pl. 81, Fig. 6-7)</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, piège lumineux, quelques exemplaires.</p><p>- P. N. duW, réserve de la Djona,Alpha Kouara, IX 2003, sous bouses de zébus, quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E16967BFC26FBC8FBC7FB18	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E16967BFF04FEB7FE03FB9F.text	857E879E8E16967BFF04FEB7FE03FB9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus tripartitus d'Orbigny 1902	<div><p>Onthophagus (s. l.) tripartitus d'Orbigny, 1902</p><p>(Pl. 65, Fig. 3 &amp; Pl. 81, Fig. 1)</p><p>-  Ouidah, forêt de Pahou, 05 I 2017, U. V., 1 ex.</p><p>- Tokay,Lobogo, 14 VI  2013, U.V., 1 ex.</p><p>- Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, piège excrément, 1 ex.;  22 X 2012, U. V., 1 ex.;</p><p>-   Kpoba, bord du Mono, 02 I 2017, U. V., 1 ex.</p><p>-  Pobé, forêt de Pobé, 18 VI 2013, U. V., 1 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 05 VI 2013, U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon,  27 IV 2013, U. V., 1 ex.;  28 IV 2013, U. V., 1 ex.;  11 I 2018,U. V., 1 ex.;  Dan, bord du Zou, 17 VI  2012, U.V., 1 ex.</p><p>- Savalou, Kpataba, montagne, 17 VI  2016, U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Pénélan, 28 V  2013, U.V., 1 ex.;  30 XI 2014, U. V., 1 ex.</p><p>- P.N.Pendjari,Batia, 18 VIII 2010, lumière, 4 ex.; 24 VIII 2010, lumière, 2 ex.</p><p>- P.N.Pendjari,hôtelPendjari, 26 VIII 2010, sous bouse de buffle, 8 ex.</p><p>- P.N.duW,Karimama, 28 IX 2010, lumière, 4 ex.</p><p>- P.N.duW,TapoaDjerma, IX 2003,sous excrément humain,quelques exemplaires.</p><p>- [Orbigny, 1913: Ouidah].</p><p>Groupe 23</p></div>	https://treatment.plazi.org/id/857E879E8E16967BFF04FEB7FE03FB9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E16967AFC26FB0FFE03FF61.text	857E879E8E16967AFC26FB0FFE03FF61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus sellatulus d'Orbigny 1902	<div><p>*  Onthophagus (s. l.) sellatulus d'Orbigny, 1902</p><p>(Pl. 67, Fig. 4 &amp; Pl. 68, Fig. 1)</p><p>-  Bohicon, quartier Soglogon, 19 X 2014, 1 ex. U. V .</p><p>-  Pobé, forêt de Pobé, 27 X 2014, 1 ex., U. V .</p><p>- P. N. duW, réserve de laDjona, mare24, IX 2003, sous excrément humain, quelques exemplaires.</p><p>- P.N.duW,chutesdeKoudou, IX 2003,sousexcrémenthumain,quelquesexemplaires.</p><p>- Tanguiéta,chaînede l’Atakora, IX 2003, sur fleur fanée, 1 ex.</p><p>Groupe 24</p></div>	https://treatment.plazi.org/id/857E879E8E16967AFC26FB0FFE03FF61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E17967AFF0AFF30FD33FE12.text	857E879E8E17967AFF0AFF30FD33FE12.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus bandamai Cambefort 1984	<div><p>*  Onthophagus (s. l.) bandamai Cambefort, 1984 (Pl. 68, Fig. 2 &amp; Pl. 81, Fig. 8)</p><p>- Pénéssoulou, Batia, 18-25 VIII 2008, piège au sol, 2 ex.</p><p>- Pénéssoulou, forêt de Goutansoukpa, 30 V 2012, lumière, 1 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2006,piège lumineux,quelques exemplaires.</p></div>	https://treatment.plazi.org/id/857E879E8E17967AFF0AFF30FD33FE12	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E17967AFF6DFE08FBE5FD6A.text	857E879E8E17967AFF6DFE08FBE5FD6A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus bidens (Olivier 1789)	<div><p>Onthophagus (s. l.) bidens (Olivier, 1789)</p><p>(Pl. 68, Fig. 3-4 &amp; Pl. 81, Fig. 9-10)</p><p>-  Ouidah, forêt de Pahou, 06 I 2017, U. V., 1 ex.</p><p>- Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, piège excrément, 7 ex.;  07 X 2012, U. V., 1 ex.;  23 XI 2015, U. V., 3 ex.</p><p>- Athiémé, bord du Mono, 31 V  2013, U.V., 4 ex.;  10 II 2017, U. V., 4 ex.</p><p>-  Zinvié, vallée du Sitatunga, 19 V 2016, U. V., 2 ex.</p><p>- Allada, Donou, 10 VI 2016, sous bouse zébu, 1 ex.</p><p>- Attogon, forêt de Niaouli,  18 XI 2015, U. V., 1 ex.; 26 V  2016, U.V., 1 ex.; 07 VI  2016, U.V., 1 ex.; 14 VI  2016, U.V., 1 ex.;  29 IV 2017, U. V., 1 ex.;  02 II 2017, U. V., 4 ex.,  03 II 2017, U. V., 1 ex.;  12 II 2017, U. V., 1 ex.</p><p>-  Adidévo, Dévé, carrière, 07 II 2017, U. V., 1 ex.</p><p>- Pobé, forêt de Pobé, 18 VI  2013, U.V., 1 ex.;  12 II 2017, U. V., 3 ex.;  13 II 2017, U. V., 3 ex.</p><p>-  Zogbodomey, forêt de Lokoli, 11 XI 2014,U. V., 1 ex.</p><p>- Bohicon, quartier Soglogon, 15 VI  2009, U.V., 3 ex.;  22 XI 2017, U. V., 1 ex.</p><p>-   Dan, bord du Zou, 17 VI  2013, U. V., 2 ex.</p><p>- Savalou, Kpataba, montagne, 17 VI  2016, U.V., 1 ex. -   Pénéssoulou, forêt de Pénélan , 25 V  2013, U.V., 1 ex.; 28 V  2013, U.V., 1 ex.; 15 VI  2016, U.V., 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu sèche, 34 ex.</p><p>- Pénéssoulou,forêt de Goutansoukpa, 21X 2011,lumière, 1 ex.; 30V  2012,U.V., 9 ex.</p><p>- Pénéssoulou, zone humide,sortie du village, 23 II 2020, 1 ex., sous bouse de zébu.</p><p>- P.N. Pendjari, Batia, 20 VIII 2010,lumière, 7 ex.; 20-26 X 2010, piège au sol, 1 ex.; 15 VI 2009, sous bouse de zébu, 1 ex.; 26 VIII 2016, sous bouse de zébu, 7 ex.</p><p>-  P.N.Pendjari, Batia, CampNumi, 11XI 2012,U. V., 1 ex.;  08 XII 2015,U. V., 1 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, 26 VII 2009, sous bouse éléphant, 1 ex.</p><p>- P. N. Pendjari, hôtel Pendjari, VI 2005, piège lumineux et excrément humain, quelques exemplaires.</p><p>- P.N.duW,Karimama, 28 IX 2015, lumière, 28 ex.</p><p>- P.N.duW,MamassiGourma, 30 IX 2015, lumière, 1 ex.</p><p>- P. N. du W,réservede laDjona,mare 24, IX 2003, sous crottins d’éléphants, quelques exemplaires.</p><p>- [Orbigny, 1913: Dahomey].</p></div>	https://treatment.plazi.org/id/857E879E8E17967AFF6DFE08FBE5FD6A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E17967AFC2AFD3FFBF4FC7A.text	857E879E8E17967AFC2AFD3FFBF4FC7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus cyanochlorus d'Orbigny 1902	<div><p>Onthophagus (s. l.) cyanochlorus d'Orbigny, 1902 (Pl. 72, Fig. 2)</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005, piège lumineux, quelques exemplaires.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2005, piège lumineux,quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003,sous crottins d'éléphants,quelques exemplaires.</p><p>- [Orbigny, 1913: Ouidah].</p></div>	https://treatment.plazi.org/id/857E879E8E17967AFC2AFD3FFBF4FC7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E17967DFC2AFC10FDC4FD6F.text	857E879E8E17967DFC2AFC10FDC4FD6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus loxodontae Cambefort 1984	<div><p>*  Onthophagus (s. l.) loxodontae Cambefort, 1984</p><p>(Pl. 69, Fig. 1 &amp; Pl. 81, Fig. 11)</p><p>- P.N.Pendjari,hôtelPendjari, VI 2005,sous excrément humain,quelques exemplaires.</p><p>Onthophagus (s. l.) mucronatus Thomson, 1858</p><p>(Pl. 69, Fig. 2-3 &amp; Pl. 81, Fig. 12)</p><p>- Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, piège excrément, 12 ex.</p><p>-  Attogon, forêt de Niaouli, 07 VI 2016, U. V., 17 ex.</p><p>-  Zogbodomey, forêt de la Lama, 12 XI 2015, U. V., 1 ex.</p><p>- Bohicon, 15 VI 2009, sous bouse de zébu, 3 ex.</p><p>-  Pénéssoulou, forêt de Pénélan, 16 VI 2016, U. V., 1 ex.</p><p>- Pénéssoulou, forêt de Saramanga, 25 V 2013, sous bouse de zébu, 3 ex.</p><p>-  Kouandé, antenne ORTB, 05 XI 2015,  U. V., 1 ex.</p><p>- P.N.Pendjari,Batia, 24 VIII 2010, lumière, 1 ex.; 26 VIII 2010, lumière, 1 ex.</p><p>- P.N.Pendjari,hôtel Pendjari, VI 2005,piège lumineux, quelques exemplaires; sous bouse de buffle, 26 VIII 2010, 7 ex.</p><p>- P.N.Pendjari,forêtdeBondjagou, VI 2005,piège lumineux,quelques exemplaires.</p><p>- P.N.duW,TapoaDjerma, IX 2003,sous crottins d'éléphants,quelques exemplaires.</p><p>- [Orbigny, 1913: plateau de Zagnanado; Ouidah].</p></div>	https://treatment.plazi.org/id/857E879E8E17967DFC2AFC10FDC4FD6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E10967DFF11FC1AFEC1FBFA.text	857E879E8E10967DFF11FC1AFEC1FBFA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus semichalcites d'Orbigny 1902	<div><p>Onthophagus semichalcites d'Orbigny, 1902</p><p>- [  Orbigny, 1902: Bénin].  Il s’agit en fait d’Onthophagus bidentatus (Drapiez, 1819), espèce américaine dont la localité type est erronée (Rossini et al., 2016)  .</p></div>	https://treatment.plazi.org/id/857E879E8E10967DFF11FC1AFEC1FBFA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E10967DFF0CFD3AFEB1FC97.text	857E879E8E10967DFF0CFD3AFEB1FC97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus seminitidus d'Orbigny 1908	<div><p>Onthophagus (s. l.) seminitidus d'Orbigny, 1908</p><p>- [Orbigny, 1908: Dahomey].</p></div>	https://treatment.plazi.org/id/857E879E8E10967DFF0CFD3AFEB1FC97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1D9670FF77F82FFADBF7A2.text	857E879E8E1D9670FF77F82FFADBF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Escarabaeus furcatus (de Castelnau 1840)	<div><p>1:  Escarabaeus furcatus (de Castelnau, 1840),   P. N. W, Mamassi-Gourma, Karimama, 30 IX 2015, 15 mm. 2:  Escarabaeus furcatus (de Castelnau, 1840), [ Réserve de Fathala (Sénégal), 12 VII 2010, 19 mm]  .</p></div>	https://treatment.plazi.org/id/857E879E8E1D9670FF77F82FFADBF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1D9670FF77F7EAFB18F781.text	857E879E8E1D9670FF77F7EAFB18F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Kheper festivus subsp. festivus (von Harold 1868)	<div><p>3:  Kheper festivus festivus (von Harold, 1868), Attogon, forêt de Niaouli, 18 XI 2015, 25 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E1D9670FF77F7EAFB18F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1D9673FF77F78AFC5EF877.text	857E879E8E1D9673FF77F78AFC5EF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Kheper subaeneus (von Harold 1869)	<div><p>4:  Kheper subaeneus (von Harold, 1869), Houéyogbé, 24 V 2009, 30 mm.</p><p>Planche 2.</p></div>	https://treatment.plazi.org/id/857E879E8E1D9673FF77F78AFC5EF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1E9673FF51F82DFB13F842.text	857E879E8E1E9673FF51F82DFB13F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Parateuchus palemo (Olivier 1789)	<div><p>1:  Parateuchus palemo (Olivier, 1789),  [Yamoussoukro (Côte d'Ivoire), IX 1997, 10,8 mm] .</p></div>	https://treatment.plazi.org/id/857E879E8E1E9673FF51F82DFB13F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1E9673FF51F7CDFAD7F781.text	857E879E8E1E9673FF51F7CDFAD7F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Scarabaeus goryi (de Castelnau 1840)	<div><p>2:  Scarabaeus goryi (de Castelnau, 1840),   ♂, [P. N. W, Tapoa Djerma (Burkina Faso), 9/ 11 VI 2004, 33 mm]. 3:  Scarabaeus goryi (de Castelnau, 1840), ♀, [ Réserve de Fathala (Sénégal), 24 VI 2009, 33 mm]  .</p></div>	https://treatment.plazi.org/id/857E879E8E1E9673FF51F7CDFAD7F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1E9672FF51F78BFC88F877.text	857E879E8E1E9672FF51F78BFC88F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Scarabaeus isidis (de Castelnau 1840)	<div><p>4:  Scarabaeus isidis (de Castelnau, 1840), Mamassi-Gourma, 29 IX 2015, 28 mm.</p><p>Planche 3.</p></div>	https://treatment.plazi.org/id/857E879E8E1E9672FF51F78BFC88F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1F9672FF42F82DFB4AF781.text	857E879E8E1F9672FF42F82DFB4AF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Chalconotus aurescens (Bates 1888)	<div><p>1:  Chalconotus aurescens (Bates, 1888), ♂, P. N. W, Mamassi-Gourma, Karimama, 30 IX 2015, 33 mm. 2:  Chalconotus convexus (Boheman, 1857), P. N. W, Mamassi-Gourma, Karimama, 30 IX 2015, 26 mm. 3:  Chalconotus cupreus (Fabricius, 1775), ♂, Attogon, forêt de Niaouli, X 2016, 25 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E1F9672FF42F82DFB4AF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1F9672FF42F78BFB8BF7E1.text	857E879E8E1F9672FF42F78BFB8BF7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Chalconotus suturalis (Janssens 1938)	<div><p>4:  Chalconotus suturalis (Janssens, 1938),   Bord du Zou,  Dan, 04 VI 2013, 22 mm  .</p></div>	https://treatment.plazi.org/id/857E879E8E1F9672FF42F78BFB8BF7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E189675FF57F82FFAFAF842.text	857E879E8E189675FF57F82FFAFAF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Allogymnopleurus aeneus (von Harold 1875)	<div><p>1:  Allogymnopleurus aeneus (von Harold, 1875), ♂, Batia, Parc de la Penjari, 18 VIII 2010, 19 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E189675FF57F82FFAFAF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E189674FF57F7CFFC8AF877.text	857E879E8E189674FF57F7CFFC8AF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Allogymnopleurus olivieri (de Castelnau 1840)	<div><p>2:  Allogymnopleurus olivieri (de Castelnau, 1840), ♂, Batia, Parc de la Penjari, 03 VII 2005, 12,3 mm. 3:  Allogymnopleurus umbrinus (Gerstaecker, 1871), ♂, Batia, Parc de la Penjari, 05 VII 2005, 10,3 mm. 4:  Allogymnopleurus youngai Endrödi, 1973, ♂, Batia, Parc de la Penjari, 18 VIII 2010, 12 mm.</p><p>Planche 5.</p></div>	https://treatment.plazi.org/id/857E879E8E189674FF57F7CFFC8AF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E199674FF42F82FFB10F842.text	857E879E8E199674FF42F82FFB10F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Garreta laetus subsp. laetus (Hope 1842)	<div><p>1:  Garreta laetus laetus (Hope, 1842), ♂, Pénéssoulou, forêt de Pénélan, 16 VI 2016, 18 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E199674FF42F82FFB10F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E199674FF42F7CFFB82F7A2.text	857E879E8E199674FF42F7CFFB82F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Garreta nitens (Olivier 1789)	<div><p>2:  Garreta nitens (Olivier, 1789), ♂, Batia, Parc de la Penjari, 24 VII 2009, 17 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E199674FF42F7CFFB82F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E199677FF42F7EAFCABF877.text	857E879E8E199677FF42F7EAFCABF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gymnopleurus coerulescens (Olivier 1789)	<div><p>3:  Gymnopleurus coerulescens (Olivier, 1789), ♀, Pénéssoulou, forêt de Saramanga, 25 V 2013, 10 mm. 4:  Gymnopleurus fulgidus (Olivier, 1789), Batia, ♂, Parc de la Penjari, 8 XI 2012, 10 mm.</p><p>Planche 6.</p></div>	https://treatment.plazi.org/id/857E879E8E199677FF42F7EAFCABF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1A9677FF7AF82FFB26F842.text	857E879E8E1A9677FF7AF82FFB26F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gymnopleurus nyankpalaensis Endrodi 1976	<div><p>1:  Gymnopleurus nyankpalaensis Endrödi, 1976, ♀, Batia, Parc de la Penjari, 18 VIII 2010, 14 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E1A9677FF7AF82FFB26F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1A9677FF7AF7CFFBB4F7A2.text	857E879E8E1A9677FF7AF7CFFBB4F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gymnopleurus profanus (Fabricius 1792)	<div><p>2:  Gymnopleurus profanus (Fabricius, 1792), Attogon, forêt de Niaouli, 2014, 13 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E1A9677FF7AF7CFFBB4F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1A9677FF7AF7EAFB6CF781.text	857E879E8E1A9677FF7AF7EAFB6CF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gymnopleurus puncticollis Gillet 1909	<div><p>3:  Gymnopleurus puncticollis Gillet, 1909, ♀, Batia, Parc de la Penjari, 27 VII 2009, 12 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E1A9677FF7AF7EAFB6CF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1A9676FF7AF78BFC8AF877.text	857E879E8E1A9676FF7AF78BFC8AF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Neosisyphus) armatus Gory 1833	<div><p>4:  
Sisyphus (Neosisyphus) armatus 
Gory, 1833, ♂, [ Réserve de Fathala (Sénégal), 14 VIII 2010, 7,5 mm].</p><p>Planche 7.</p></div>	https://treatment.plazi.org/id/857E879E8E1A9676FF7AF78BFC8AF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1B9676FF0FF82FFB30F842.text	857E879E8E1B9676FF0FF82FFB30F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Neosisyphus) baoule (Cambefort 1984)	<div><p>1:  Sisyphus (Neosisyphus) baoule (Cambefort, 1984), ♂, Batia, Parc de la Penjari, 07 XI 2015, 10 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E1B9676FF0FF82FFB30F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1B9676FF0FF7CFFB61F781.text	857E879E8E1B9676FF0FF7CFFB61F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Neosisyphus) gladiator Arrow 1927	<div><p>2:  
Sisyphus (Neosisyphus) gladiator 
Arrow, 1927, ♀, [Côte d'Ivoire,  Parc National la Marahoué, IX 2000, 11,3 mm]. 3:  Sisyphus (Neosisyphus) gladiator Arrow, 1927, ♂, Batia, Parc de la Penjari, 05 VI 2005, 9 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E1B9676FF0FF7CFFB61F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E1B9669FF0FF78BFC8BF877.text	857E879E8E1B9669FF0FF78BFC8BF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Neosisyphus) paschalidisae (Cambefort 1984)	<div><p>4:  Sisyphus (Neosisyphus) paschalidisae (Cambefort, 1984), ♂, Batia, Parc de la Penjari, 20 VIII 2010, 10,5 mm,.</p><p>Planche 8.</p></div>	https://treatment.plazi.org/id/857E879E8E1B9669FF0FF78BFC8BF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E049669FF01F82FFB65F842.text	857E879E8E049669FF01F82FFB65F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Neosisyphus) youngai Endrodi 1973	<div><p>1:  
Sisyphus (Neosisyphus) youngai 
Endrödi, 1973, ♂, [ Tou (Côte d'Ivoire), 01 III 2017, 8,1 mm].</p></div>	https://treatment.plazi.org/id/857E879E8E049669FF01F82FFB65F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E049669FF01F7CFFB65F7A2.text	857E879E8E049669FF01F7CFFB65F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Neosisyphus) youngai Endrodi 1973	<div><p>2:  
Sisyphus (Neosisyphus) youngai 
Endrödi, 1973, ♀, [ Tou (Côte d'Ivoire), 01 III 2017, 7,4 mm].</p></div>	https://treatment.plazi.org/id/857E879E8E049669FF01F7CFFB65F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E049668FF01F7EAFCE2F877.text	857E879E8E049668FF01F7EAFCE2F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Sisyphus) africanus subsp. africanus Montreuil 2017	<div><p>3:  Sisyphus (Sisyphus) africanus africanus Montreuil, 2017, Pénéssoulou, forêt de Saramanga, 25 V 2013, 4,5 mm. 4:  Sisyphus (Sisyphus) arboreus Walter, 1982, Attogon, forêt de Niaouli, 31 X 2014, 5 mm.</p><p>2</p><p>Planche 9.</p></div>	https://treatment.plazi.org/id/857E879E8E049668FF01F7EAFCE2F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E059668FF79F7CFFBD7F781.text	857E879E8E059668FF79F7CFFBD7F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Catharsius achates (Olivier 1789)	<div><p>2:  Catharsius achates (Olivier, 1789), ♂, P. N. du W, Mamassi-Gourma, Karimama, 29 IX 2015, 32 mm. 3:  Catharsius eteocles (de Castelnau, 1840), ♂, Bohicon, 28 IV 2013, 26 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E059668FF79F7CFFBD7F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E059668FF79F82FFBDDF842.text	857E879E8E059668FF79F82FFBDDF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Sisyphus (Parasisyphus) Barbero, Palestrini & Zunino 1991	<div><p>1:  Sisyphus (Parasisyphus) complexe  goryi, Lokossa, 01 III 2006, 4,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E059668FF79F82FFBDDF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E05966BFF79F78BFC87F877.text	857E879E8E05966BFF79F78BFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Catharsius guineensis Ferreira 1963	<div><p>4:  Catharsius guineensis Ferreira, 1963, ♂, Pénéssoulou, forêt de Saramanga, 25 V 2013, 19 mm.</p><p>Planche 10.</p></div>	https://treatment.plazi.org/id/857E879E8E05966BFF79F78BFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E06966BFF42F7CFFAFAF781.text	857E879E8E06966BFF42F7CFFAFAF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Catharsius	<div><p>2:  Catharsius complexe polynices Kolbe, 1893, ♀, Pénéssoulou, forêt de Pénélan, VII-IX 2013, 26 mm. 3:  Catharsius complexe polynices Kolbe, 1893, ♂, Pénéssoulou, forêt de Pénélan, 15 V 2013, 30 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E06966BFF42F7CFFAFAF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E06966BFF42F82FFC32F842.text	857E879E8E06966BFF42F82FFC32F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Catharsius guineensis Ferreira 1963	<div><p>1:  Catharsius guineensis Ferreira, 1963, ♀, Bohicon, 6 VI 2016, 19 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E06966BFF42F82FFC32F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E06966AFF42F78BFC87F877.text	857E879E8E06966AFF42F78BFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Catharsius pseudolycaon Ferreira 1960	<div><p>4:  Catharsius pseudolycaon Ferreira, 1960, ♂, Kouandé, 05 XI 2015, 23 mm.</p><p>Planche 11.</p></div>	https://treatment.plazi.org/id/857E879E8E06966AFF42F78BFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E07966AFF47F7CFFB68F781.text	857E879E8E07966AFF47F7CFFB68F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Catharsius	<div><p>2:  Catharsius complexe sesostris Waterhouse, 1891, ♂, Attogon, forêt de Niaouli, 02 IV 2013, 21 mm. 3:  Catharsius complexe sesostris Waterhouse, 1891, ♀, forêt de Ndali, 11 XI 2015, 17 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E07966AFF47F7CFFB68F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E07966AFF47F82FFBF4F842.text	857E879E8E07966AFF47F82FFBF4F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Catharsius pseudolycaon Ferreira 1960	<div><p>1:  Catharsius pseudolycaon Ferreira, 1960, ♀, Kouandé, 05 XI 2015, 19 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E07966AFF47F82FFBF4F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E07966DFF47F78AFC80F877.text	857E879E8E07966DFF47F78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris angustus Nguyen-Phung et Cambefort 1986	<div><p>4:  
Copris angustus 
Nguyen-Phung et Cambefort, 1986, ♂, [Comoé (Côte d’Ivoire), 05 2004, 21,2 mm].</p><p>Planche 12.</p></div>	https://treatment.plazi.org/id/857E879E8E07966DFF47F78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E00966DFF52F82FFBF8F7A2.text	857E879E8E00966DFF52F82FFBF8F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris angustus Nguyen-Phung et Cambefort 1986	<div><p>1:  
Copris angustus 
Nguyen-Phung et Cambefort, 1986, ♀, [Comoé (Côte d’Ivoire), 05 2004, 19,7 mm]. 2:  Copris carmelita (Fabricius, 1801), ♀,  Goutansoupka, 21 X 2011, 14 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E00966DFF52F82FFBF8F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E00966DFF52F7EAFBF3F781.text	857E879E8E00966DFF52F7EAFBF3F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris coriarius Gillet 1907	<div><p>3:  Copris coriarius Gillet, 1907, ♂, forêt de Pénéssoulou, 12 V 2013, 14 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E00966DFF52F7EAFBF3F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E00966CFF52F78AFCAEF877.text	857E879E8E00966CFF52F78AFCAEF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris coriarius Gillet 1907	<div><p>4:  Copris coriarius Gillet, 1907, ♀, Batia, Parc de la Pendjari, 07 XI 2012, 14 mm.</p><p>Planche 13.</p></div>	https://treatment.plazi.org/id/857E879E8E00966CFF52F78AFCAEF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E01966CFE97F82FFBE0F842.text	857E879E8E01966CFE97F82FFBE0F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris gazellarum Gillet 1918	<div><p>1:  Copris gazellarum Gillet, 1918, ♀, Pénéssoulou, 17 VI 2013, 15 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E01966CFE97F82FFBE0F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E01966CFE97F7CFFB00F7A2.text	857E879E8E01966CFE97F7CFFB00F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris interioris Kolbe 1897	<div><p>2:  Copris interioris Kolbe, 1897, ♂, Pénéssoulou, forêt de Pénélan, 27 V 2013, 18 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E01966CFE97F7CFFB00F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E01966CFE97F7EAFB32F781.text	857E879E8E01966CFE97F7EAFB32F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris interioris Kolbe 1897	<div><p>3:  Copris interioris Kolbe, 1897, ♀, Pénéssoulou, forêt de Pénélan, 02 VII 2013, 17 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E01966CFE97F7EAFB32F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E01966FFE97F78AFC87F877.text	857E879E8E01966FFE97F78AFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris laius subsp. laius von Harold 1868	<div><p>4:  Copris laius laius von Harold, 1868, ♂, Pénéssoulou, forêt de Pénélan, 28 V 2013, 23 mm.</p><p>Planche 14.</p></div>	https://treatment.plazi.org/id/857E879E8E01966FFE97F78AFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E02966FFF5AF82FFAD0F842.text	857E879E8E02966FFF5AF82FFAD0F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris laius subsp. laius von Harold 1868	<div><p>1:  Copris laius laius von Harold, 1868, ♀, Pénéssoulou, forêt de Goutansoukpa, VIII 2011, 23 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E02966FFF5AF82FFAD0F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E02966FFF5AF7CFFB85F781.text	857E879E8E02966FFF5AF7CFFB85F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris megaceratoides Waterhouse 1891	<div><p>2:  Copris megaceratoides Waterhouse, 1891, ♂, Pénéssoulou, forêt de Saramanga, 24 VI 2013, 23 mm. 3:  Copris megaceratoides Waterhouse, 1891, ♀, forêt de Dan, 04 VI 2013, 23 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E02966FFF5AF7CFFB85F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E02966EFF5AF78AFC8BF877.text	857E879E8E02966EFF5AF78AFC8BF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris moffartsi Gillet 1907	<div><p>4:  Copris moffartsi Gillet, 1907, ♂, Pénéssoulou, forêt de Pénélan, VII-IX 2013, 18 mm.</p><p>Planche 15.</p></div>	https://treatment.plazi.org/id/857E879E8E02966EFF5AF78AFC8BF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E03966EFEBAF82FFB7DF842.text	857E879E8E03966EFEBAF82FFB7DF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris moffartsi Gillet 1907	<div><p>1:  Copris moffartsi Gillet, 1907, ♀, Pénéssoulou, forêt de Pénélan, 24 I 2017, 19 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E03966EFEBAF82FFB7DF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E03966EFEBAF7CFFC3CF7A2.text	857E879E8E03966EFEBAF7CFFC3CF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris orion subsp. orion Klug 1835	<div><p>2:  Copris orion orion Klug, 1835, ♂, Bohicon, 12 VI 2009, 13 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E03966EFEBAF7CFFC3CF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E03966EFEBAF7EAFB60F781.text	857E879E8E03966EFEBAF7EAFB60F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris orion subsp. orion Klug 1835	<div><p>3:  Copris orion orion Klug, 1835, ♀, Pénéssoulou, forêt de Pénélan, 27 V 2013, 12 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E03966EFEBAF7EAFB60F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E039661FEBAF78AFC82F877.text	857E879E8E039661FEBAF78AFC82F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris orphanus Guerin-Meneville 1847	<div><p>4:  Copris orphanus Guérin-Méneville, 1847, Pénéssoulou, forêt de Pénélan, 10 XI 2014, 18 mm.</p><p>Planche 16.</p></div>	https://treatment.plazi.org/id/857E879E8E039661FEBAF78AFC82F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0C9661FF7FF82FFC52F842.text	857E879E8E0C9661FF7FF82FFC52F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris sinon Olivier 1789	<div><p>1:  Copris sinon Olivier, 1789, ♂, Karimama, 28 IX 2015, 11 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0C9661FF7FF82FFC52F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0C9661FF7FF7CFFB8EF7A2.text	857E879E8E0C9661FF7FF7CFFB8EF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Heliocopris antenor (Olivier 1789)	<div><p>2:  Heliocopris antenor (Olivier, 1789),   ♀, [ Nianing (Sénégal), 15 IX 2009, 41 mm]  .</p></div>	https://treatment.plazi.org/id/857E879E8E0C9661FF7FF7CFFB8EF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0C9660FF7FF7EAFC87F877.text	857E879E8E0C9660FF7FF7EAFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Heliocopris dilloni Guerin-Meneville 1847	<div><p>3:  
Heliocopris 
cf. dilloni Guérin-Méneville, 1847, ♂, [ Tapoa Djerma (Burkina Faso), 09-11 VI 2004, 51 mm]. 4:  Heliocopris myrmidon Kolbe, 1893, ♂, Pénéssoulou, forêt de Pénélan, 15 V 2013, 36 mm.</p><p>Planche 17.</p></div>	https://treatment.plazi.org/id/857E879E8E0C9660FF7FF7EAFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0D9660FEBAF82FFB82F7A2.text	857E879E8E0D9660FEBAF82FFB82F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Litocopris punctiventris Waterhouse 1891	<div><p>1:  Litocopris punctiventris Waterhouse, 1891, Pénéssoulou, forêt de Pénélan, 16 VI 2015, 11 mm. 2:  Metacatharsius abortivus Fairmaire, 1891, ♀, Bohicon, 28 IV 2013, 5,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0D9660FEBAF82FFB82F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0D9660FEBAF7EAFBC6F781.text	857E879E8E0D9660FEBAF7EAFBC6F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Metacatharsius Paulian 1939	<div><p>3:  Metacatharsius complexe inermis sp. 1, ♀, Lokossa, 2008, 11,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0D9660FEBAF7EAFBC6F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0D9663FEBAF78AFC84F877.text	857E879E8E0D9663FEBAF78AFC84F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Metacatharsius Paulian 1939	<div><p>4:  Metacatharsius complexe inermis sp. 2, ♂, Karimama, 28 IX 2015, 14 mm.</p><p>Planche 18.</p></div>	https://treatment.plazi.org/id/857E879E8E0D9663FEBAF78AFC84F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0E9663FEBAF82FFB31F7A2.text	857E879E8E0E9663FEBAF82FFB31F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Metacatharsius peleus (Olivier 1789)	<div><p>1:  Metacatharsius peleus (Olivier, 1789), ♀, P. N. du W, Mamassi-Gourma, 30 IX 2015, 15 mm. 2:  Metacatharsius renaudi Frey, 1960, ♂, P. N. du W, réserve de la Djona, IX 2003, 4,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0E9663FEBAF82FFB31F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0E9663FEBAF7EAFB32F781.text	857E879E8E0E9663FEBAF7EAFB32F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Metacatharsius renaudi Frey 1960	<div><p>3:  Metacatharsius renaudi Frey, 1960, ♀, P. N. du W, réserve de la Djona, IX 2003, 5,3 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0E9663FEBAF7EAFB32F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0E9663FEBAF78AFAD0F7E1.text	857E879E8E0E9663FEBAF78AFAD0F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Metacatharsius tschadensis Balthasar 1961	<div><p>4:  Metacatharsius tschadensis Balthasar, 1961 ?, Zinvié, vallée du Sitatunga, 19 V 2016, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0E9663FEBAF78AFAD0F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0F9662FE82F82FFB03F781.text	857E879E8E0F9662FE82F82FFB03F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria coprinarum Cambefort 1982	<div><p>1:  Pedaria coprinarum Cambefort, 1982, ♂, Bohicon, quartier Soglogon, 25 VI 2018, 6 mm. 2:  Pedaria criberrima Waterhouse, 1890, ♀, Bohicon, Quartier Soglogon, 28 IV 2013, 7 mm. 3:  Pedaria durandi Paulian, Cambefort &amp; Mauchamp, 1982, ♀, Bohicon, 06 X 2009, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0F9662FE82F82FFB03F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0F9665FE82F78AFCBAF871.text	857E879E8E0F9665FE82F78AFCBAF871.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria estellae Josso & Prevost 2006	<div><p>4:  Pedaria estellae Josso &amp; Prévost, 2006, ♂, Parc de la Pendjari, 01-03 VI 2005, 7,2 mm.</p><p>Planche 20.</p></div>	https://treatment.plazi.org/id/857E879E8E0F9665FE82F78AFCBAF871	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E089665FF67F810FB55F84F.text	857E879E8E089665FF67F810FB55F84F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria fernandezi Cambefort 1982	<div><p>1:  Pedaria fernandezi Cambefort, 1982, Savalou, forêt de Logozohé, 06 VIII 2007, 6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E089665FF67F810FB55F84F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E089665FF67F830FB9DF782.text	857E879E8E089665FF67F830FB9DF782.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pedaria humana Cambefort 1982	<div><p>2:  
Pedaria humana 
Cambefort, 1982, [Côte d'Ivoire, Parc Naturel de la Comoe  Karpin, 04-08 V 1981, 8,1 mm]. 3:  Pedaria renwarti Cambefort, 1982, Bohicon, quartier Soglogon, 03 VI 2018, 6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E089665FF67F830FB9DF782	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E089664FF67F78FFC80F877.text	857E879E8E089664FF67F78FFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Heteronitis tridens (de Castelnau 1840)	<div><p>4:  Heteronitis tridens (de Castelnau, 1840),  ♀, [P. N. W, Tapoa Djerma (Burkina Faso), 09-11 VI 2004, 28 mm] .</p><p>Planche 21.</p></div>	https://treatment.plazi.org/id/857E879E8E089664FF67F78FFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E099664FF79F82FFB5CF7A2.text	857E879E8E099664FF79F82FFB5CF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Heteronitis tridens (de Castelnau 1840)	<div><p>1:  Heteronitis tridens (de Castelnau, 1840), ♂, Parc de la Penjari, forêt de Bondjagou, 6-7 VI 2005, 28,3 mm. 2:  Onitis aeneus van Lansberge, 1875, ♂, Batia, Parc de la Penjari, 26 VIII 2010, 20 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E099664FF79F82FFB5CF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E099664FF79F7EAFB5DF781.text	857E879E8E099664FF79F7EAFB5DF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis aeneus van Lansberge 1875	<div><p>3:  Onitis aeneus van Lansberge, 1875, ♀, Batia, Parc de la Penjari, 26 VIII 2010, 20 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E099664FF79F7EAFB5DF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E099667FF79F78AFC87F877.text	857E879E8E099667FF79F78AFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis affinis Felsche 1911	<div><p>4:  Onitis affinis Felsche, 1911, ♂, Parc de la Penjari, forêt de Bondjagou, 25 VII 2009, 22 mm.</p><p>Planche 22.</p></div>	https://treatment.plazi.org/id/857E879E8E099667FF79F78AFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0A9667FEB2F82FFBA8F842.text	857E879E8E0A9667FEB2F82FFBA8F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis alexis subsp. alexis Klug 1835	<div><p>1:  Onitis alexis alexis Klug, 1835, ♂, Ouidah, forêt de Pahou, 05 I 2017, 17 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0A9667FEB2F82FFBA8F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0A9667FEB2F7CFFB78F7A2.text	857E879E8E0A9667FEB2F7CFFB78F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis alexis subsp. alexis Klug 1835	<div><p>2:  Onitis alexis alexis Klug, 1835, ♀, Bohicon, quartier Soglogon, 15 VI 2009, 18 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0A9667FEB2F7CFFB78F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0A9666FEB2F7EAFC82F877.text	857E879E8E0A9666FEB2F7EAFC82F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis bocandei Gillet 1911	<div><p>3:  Onitis bocandei Gillet, 1911, ♂, Parc de la Penjari, forêt de Bondjagou, 26 VII 2009, 23 mm. 4:  Onitis camerunus Gillet, 1933, ♂, Pénéssoulou, forêt de Saramanga, 24 V 2013, 30 mm.</p><p>Planche 23.</p></div>	https://treatment.plazi.org/id/857E879E8E0A9666FEB2F7EAFC82F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0B9666FEBAF82FFB07F842.text	857E879E8E0B9666FEBAF82FFB07F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis camerunus Gillet 1933	<div><p>1:  Onitis camerunus Gillet, 1933, ♀, Pénéssoulou, forêt de Saramanga, 24 V 2013, 26 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0B9666FEBAF82FFB07F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0B9666FEBAF7CFFB78F7A2.text	857E879E8E0B9666FEBAF7CFFB78F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis cupreus de Castelnau 1840	<div><p>2:  Onitis cupreus de Castelnau, 1840, ♂, Batia, Parc de la Penjari, 03 XI 2015, 16 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0B9666FEBAF7CFFB78F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0B9666FEBAF7EAFB61F781.text	857E879E8E0B9666FEBAF7EAFB61F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis cupreus de Castelnau 1840	<div><p>3:  Onitis cupreus de Castelnau, ♀, 1840, Batia, Parc de la Penjari, 26 VIII 2010, 15 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E0B9666FEBAF7EAFB61F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E0B9619FEBAF78AFC80F877.text	857E879E8E0B9619FEBAF78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis gilleti Boucomont 1933	<div><p>4:  Onitis gilleti Boucomont, 1933, ♂, Parc de la Penjari, forêt de Bondjagou, 26 VII 2009, 22 mm.</p><p>Planche 24.</p></div>	https://treatment.plazi.org/id/857E879E8E0B9619FEBAF78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E749619FEA9F82FFB80F842.text	857E879E8E749619FEA9F82FFB80F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis guineensis Gillet 1918	<div><p>1:  Onitis guineensis Gillet, 1918, ♂, Parc de la Pendjari, 01-03 VI 2005, 22 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E749619FEA9F82FFB80F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E749619FEA9F7CFFB14F781.text	857E879E8E749619FEA9F7CFFB14F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis guineensis Gillet 1918	<div><p>2:  Onitis guineensis Gillet, 1918, ♀, Parc de la Penjari, forêt de Bondjagou, 6-7 VI 2005, 20,5 mm. 3:  Onitis nigeriensis Ferreira, 1977, ♂, Pénéssoulou, forêt de Pénélan, 17 VI 2013, 20 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E749619FEA9F7CFFB14F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E749618FEA9F78AFC80F877.text	857E879E8E749618FEA9F78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis nigeriensis Ferreira 1977	<div><p>4:  Onitis nigeriensis Ferreira, 1977, ♀, Pénéssoulou, forêt de Saramanga, 24 V 2013, 18 mm.</p><p>Planche 25.</p></div>	https://treatment.plazi.org/id/857E879E8E749618FEA9F78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E759618FF57F82FFB16F842.text	857E879E8E759618FF57F82FFB16F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis reichei subsp. reichei van Lansberge 1875	<div><p>1:  
Onitis reichei reichei 
van Lansberge, ♂, 1875,  Kpoba, bord du Mono, 02 I 2017, 18 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E759618FF57F82FFB16F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E75961BFF57F7CFFC87F862.text	857E879E8E75961BFF57F7CFFC87F862.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis reichei subsp. reichei van Lansberge 1875	<div><p>2:  Onitis reichei reichei van Lansberge, ♀, 1875, Pénéssoulou, forêt de Pénélan, 15 VI 2016, 18 mm. 3:  Onitis sibutensis Janssens, 1943, ♂, Parc de la Penjari, forêt de Bondjagou, 26 VII 2009, 19 mm. 4:  Onitis vanderkelleni van Lansberge, 1886, ♂, Houéyogbé, forêt de Houéyogbé, III 2012, 19 mm.</p><p>Planche 26.</p></div>	https://treatment.plazi.org/id/857E879E8E75961BFF57F7CFFC87F862	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E76961BFF11F822FB32F7B9.text	857E879E8E76961BFF11F822FB32F7B9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis violaceus van Lansberge 1875	<div><p>1:  Onitis violaceus van Lansberge, 1875, Parc de la Penjari, forêt de Bondjagou, 25 VII 2009, 18 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E76961BFF11F822FB32F7B9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E76961BFF11F7C2FBAEF799.text	857E879E8E76961BFF11F7C2FBAEF799.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pleuronitis boucomonti Janssens 1937	<div><p>2:  Pleuronitis boucomonti Janssens, 1937, Batia, Parc de la Penjari, 24 VII 2009, 14 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E76961BFF11F7C2FBAEF799	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E76961AFF11F7E0FC80F877.text	857E879E8E76961AFF11F7E0FC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Clypeodrepanus strigatus (Janssens 1953)	<div><p>3:  Clypeodrepanus strigatus (Janssens, 1953),   [République Centrafricaine, P. N. Ndoki Lac, 28 0 I-2012, 3,8 mm]. 4:  Cyptochirus distinctus (Janssens, 1953), Batia, Parc de la Penjari, 27 VII 2009, 11 mm  .</p><p>Planche 27.</p></div>	https://treatment.plazi.org/id/857E879E8E76961AFF11F7E0FC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E77961AFF49F82FFC27F842.text	857E879E8E77961AFF49F82FFC27F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Eodrepanus bechynei (Janssens 1953)	<div><p>1:  Eodrepanus bechynei (Janssens, 1953), ♀, Bohicon, 15 VI 2009, 4 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E77961AFF49F82FFC27F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E77961AFF49F7CFFB5BF781.text	857E879E8E77961AFF49F7CFFB5BF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Euoniticellus capnus Cambefort 1996	<div><p>2:  Euoniticellus capnus Cambefort, 1996, ♂, Parc de la Penjari, forêt de Bondjagou, 26 VII 2009, 5,5 mm. 3:  Euoniticellus intermedius (Reiche, 1849), ♂, Lokossa, Djotokpa, 03 IV 2006, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E77961AFF49F7CFFB5BF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E77961DFF49F78AFCB3F877.text	857E879E8E77961DFF49F78AFCB3F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Euoniticellus kawanus Janssens 1939	<div><p>4:  Euoniticellus kawanus Janssens, 1939, ♀, Parc de la Penjari, forêt de Bondjagou, 26 VII 2009, 8 mm.</p><p>Planche 28.</p></div>	https://treatment.plazi.org/id/857E879E8E77961DFF49F78AFCB3F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E70961DFF61F82FFB18F842.text	857E879E8E70961DFF61F82FFB18F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Euoniticellus parvus (Kraatz 1895)	<div><p>1:  Euoniticellus parvus (Kraatz, 1895), ♀, Pénéssoulou, forêt de Saramanga, 24 V 2013, 4 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E70961DFF61F82FFB18F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E70961DFF61F7CFFB9FF781.text	857E879E8E70961DFF61F7CFFB9FF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Euoniticellus triangulatus (von Harold 1873)	<div><p>2:  Euoniticellus triangulatus (von Harold, 1873),   ♂, [Zambie (E. Mongu), piste de Ndanda, 10 XI 2008, 8,3 mm]. 3:  Ixodina bos (Endrödi, 1976), Savalou, forêt de Logozohé, 06 VIII 2007, 3,5 mm  .</p></div>	https://treatment.plazi.org/id/857E879E8E70961DFF61F7CFFB9FF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E70961CFF61F78AFC80F877.text	857E879E8E70961CFF61F78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Ixodina endroedyi (Endrodi 1976)	<div><p>4:  Ixodina endroedyi (Endrödi, 1976), Pénéssoulou, forêt de Saramanga, 25 V 2013, 3,5 mm.</p><p>Planche 29.</p></div>	https://treatment.plazi.org/id/857E879E8E70961CFF61F78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E71961CFF4AF82FFB71F842.text	857E879E8E71961CFF4AF82FFB71F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Ixodina szunyoghyi subsp. occidentalis (Endrodi 1976)	<div><p>1:  Ixodina szunyoghyi occidentalis (Endrödi, 1976), Bassila, Tchétou, 17 XI 2016, 3,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E71961CFF4AF82FFB71F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E71961FFF4AF7CFFC80F877.text	857E879E8E71961FFF4AF7CFFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Latodrepanus simonisi (Barbero 2009)	<div><p>2:  Latodrepanus simonisi (Barbero et al., 2009), ♂, Pénéssoulou, forêt de Saramanga, 25 V 2013, 4,5 mm. 3:  Liatongus militaris (de Castelnau, 1840), ♀, Alphahouara, Réserve de la Djona, 23-28 IX 2003, 9,3 mm. 4:  Oniticellus formosus Chevrolat, 1830, Batia, Parc de la Penjari, 09 XI 2015, 10 mm.</p><p>Planche 30.</p></div>	https://treatment.plazi.org/id/857E879E8E71961FFF4AF7CFFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E72961FFF72F82FFB62F7A2.text	857E879E8E72961FFF72F82FFB62F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tibiodrepanus sulcicollis (de Castelnau 1840)	<div><p>1:  Tibiodrepanus sulcicollis (de Castelnau, 1840),   ♂, [Zambie,  Province de Ikelenge, 08 XII-2012, 4,6 mm]. 2:  Tibiodrepanus sulcicollis (de Castelnau, 1840), ♀, Lokossa, Djetopka, 12 VIII 2007, 5 mm  .</p></div>	https://treatment.plazi.org/id/857E879E8E72961FFF72F82FFB62F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E72961FFF72F7EAFB4AF781.text	857E879E8E72961FFF72F7EAFB4AF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiniocellus praetermissus Branco 2010	<div><p>3:  Tiniocellus praetermissus Branco, 2010, Batia, Parc de la Penjari, 11 XI 2012, 6,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E72961FFF72F7EAFB4AF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E72961EFF72F78AFC9BF877.text	857E879E8E72961EFF72F78AFC9BF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiniocellus setifer (Kraatz 1895)	<div><p>4:  Tiniocellus setifer (Kraatz, 1895), Pénéssoulou, forêt de Pénélan, 03 XII 2014, 6 mm.</p><p>Planche 31.</p></div>	https://treatment.plazi.org/id/857E879E8E72961EFF72F78AFC9BF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E73961EFF7FF82FFB35F7A2.text	857E879E8E73961EFF7FF82FFB35F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Alloscelus paradoxus Boucomont 1923	<div><p>1:  Alloscelus paradoxus Boucomont, 1923 sensu lato, Houéyogbé, forêt de Houéyogbé, 15 X 2014, 8 mm. 2:  Cambefortius bidentiger (d’Orbigny, 1905), Pénéssoulou, forêt de Pénélan, 16 VI 2016, 5,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E73961EFF7FF82FFB35F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E739611FF7FF78AFCB4F877.text	857E879E8E739611FF7FF78AFCB4F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pinacotarsus dohrni von Harold 1875	<div><p>4:  Pinacotarsus dohrni von Harold, 1875, Pénéssoulou, forêt de Pénélan, 23 V 2013, 8,5 mm.</p><p>Planche 32.</p></div>	https://treatment.plazi.org/id/857E879E8E739611FF7FF78AFCB4F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7C9611FF7FF82FFADAF842.text	857E879E8E7C9611FF7FF82FFADAF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pseudosaproecius comatus (d'Orbigny 1902)	<div><p>1:  Pseudosaproecius comatus (d’Orbigny, 1902), ♂, Pénéssoulou, forêt de Pénélan, 28 V 2013, 6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7C9611FF7FF82FFADAF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7C9611FF7FF7CFFB28F781.text	857E879E8E7C9611FF7FF7CFFB28F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pseudosaproecius cylindroides (d'Orbigny 1908)	<div><p>2:  Pseudosaproecius cf cylindroides (d’Orbigny, 1908), ♀, Houéyogbé, forêt de Houéyogbé, 01 VI 2013, 6 mm. 3:  Pseudosaproecius ocellatus (Frey, 1958), ♀, Zogbodomey, forêt de la Lama, 06 VI 2013, 4 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7C9611FF7FF7CFFB28F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7C9610FF7FF78AFC87F877.text	857E879E8E7C9610FF7FF78AFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pseudosaproecius portentosus (Felsche 1907)	<div><p>4:  Pseudosaproecius portentosus (Felsche, 1907), ♂, Pénéssoulou, forêt de Pénélan, 23 V 2013, 10 mm.</p><p>Planche 33.</p></div>	https://treatment.plazi.org/id/857E879E8E7C9610FF7FF78AFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7D9610FF4FF82FFB2BF842.text	857E879E8E7D9610FF4FF82FFB2BF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pseudosaproecius portentosus (Felsche 1907)	<div><p>1:  Pseudosaproecius portentosus (Felsche, 1907), ♀, Batia, Parc de la Penjari, 07 XI 2015, 8 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7D9610FF4FF82FFB2BF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7D9613FF4FF7CFFC87F877.text	857E879E8E7D9613FF4FF7CFFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Pseudosaproecius Balthasar 1941	<div><p>2:  Pseudosaproecius  complexe validicornis (Quedenfeldt, 1884), ♂, Dan, bord du Zou, 04 VI 2013, 5 mm. 3:  Pseudosaproecius complexe validicornis (Quedenfeldt, 1884), ♀,  Savalou, Kpataba, 14 XII 2016, 5 mm. 4:  Stiptopodius doriae von Harold, 1871, [Niger, Kouré, 19-21 IX 2003, 4,9 mm]  .</p><p>Planche 34.</p></div>	https://treatment.plazi.org/id/857E879E8E7D9613FF4FF7CFFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7E9613FE9AF7CFFBACF7A2.text	857E879E8E7E9613FE9AF7CFFBACF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Milichus apicalis (Fahraeus 1857)	<div><p>2:  Milichus apicalis (Fahraeus, 1857), Lokossa, Djetokpa, 28 XII 2011, 6,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7E9613FE9AF7CFFBACF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7E9613FE9AF7EAFBBDF781.text	857E879E8E7E9613FE9AF7EAFBBDF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Milichus apicalis (Fahraeus 1857)	<div><p>3:  Milichus apicalis (Fahraeus, 1857), Lokossa, Djetokpa, 12 VIII 2007, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7E9613FE9AF7EAFBBDF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7E9613FE9AF82FFB65F842.text	857E879E8E7E9613FE9AF82FFB65F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Stiptopodius gaillardi (Boucomont 1923)	<div><p>1:  Stiptopodius gaillardi (Boucomont, 1923),   [Niger,  Kouré, 19-21 IX 2003, 6,8 mm]  .</p></div>	https://treatment.plazi.org/id/857E879E8E7E9613FE9AF82FFB65F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7E9612FE9AF78AFC80F877.text	857E879E8E7E9612FE9AF78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Milichus durandi Cambefort 1996	<div><p>4:  Milichus durandi Cambefort, 1996, Pénéssoulou, forêt de Pénélan, 03 VII 2013, 6,5 mm.</p><p>Planche 35.</p></div>	https://treatment.plazi.org/id/857E879E8E7E9612FE9AF78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7F9612FF52F7CFFB2CF781.text	857E879E8E7F9612FF52F7CFFB2CF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobiomorphus megaponerae Brauns 1914	<div><p>2:  Caccobiomorphus megaponerae Brauns, 1914, Houéyogbé, forêt de Houéyogbé, 20 X 2014, 5 mm. 3:  Caccobius (s. l.) anthracites (d'Orbigny, 1904), Batia, Parc de la Penjari, 07 XI 2015, 11 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7F9612FF52F7CFFB2CF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7F9612FF52F82FFBA4F842.text	857E879E8E7F9612FF52F82FFBA4F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Milichus serratus d'Orbigny 1907	<div><p>1:  Milichus serratus d'Orbigny, 1907, Batia, Parc de la Penjari, 03 XI 2015, 7,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7F9612FF52F82FFBA4F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7F9615FF52F78AFCB0F877.text	857E879E8E7F9615FF52F78AFCB0F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius auberti d'Orbigny 1902	<div><p>4:  Caccobius (s. l.) auberti d'Orbigny, 1902, ♀, Batia, Parc de la Penjari, 27 VII 2009, 3,5 mm.</p><p>Planche 36.</p></div>	https://treatment.plazi.org/id/857E879E8E7F9615FF52F78AFCB0F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E789615FE8AF82FFB28F842.text	857E879E8E789615FE8AF82FFB28F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius bama Walter 2014	<div><p>1:  Caccobius (s. l.) bama Walter, 2014, ♀, Pénéssoulou, forêt de Saramanga, 24 V 2013, 3 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E789615FE8AF82FFB28F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E789615FE8AF7CFFB3EF7A2.text	857E879E8E789615FE8AF7CFFB3EF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius cavatus d'Orbigny 1908	<div><p>2:  Caccobius (s. l.) cavatus d'Orbigny, 1908, ♂, Karimama, P. N. du W, 28 IX 2015, 3 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E789615FE8AF7CFFB3EF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E789614FE8AF7EAFC80F877.text	857E879E8E789614FE8AF7EAFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius dorsalis von Harold 1867	<div><p>3:  Caccobius (s. l.) dorsalis von Harold, 1867, Bohicon, quartier Soglogon, 06 VI 2016, 4,5 mm. 4:  Caccobius (s. l.) ferrugineus Fahraeus, 1857, Bohicon, quartier Soglogon, 27 IV 2013, 4,5 mm.</p><p>Planche 37.</p></div>	https://treatment.plazi.org/id/857E879E8E789614FE8AF7EAFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E799614FF62F82FFAD2F842.text	857E879E8E799614FF62F82FFAD2F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius ivorensis Cambefort 1984	<div><p>1:  Caccobius (s. l.) ivorensis Cambefort 1984, ♀, P. N. du W, réserve de la Djona, 28 IX 2003, 3,7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E799614FF62F82FFAD2F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E799614FF62F7CFFAC7F7A2.text	857E879E8E799614FF62F7CFFAC7F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius mirabilepunctatus Cambefort 1971	<div><p>2:  Caccobius (s. l.) mirabilepunctatus Cambefort, 1971, Batia, P. N. de la Pendjari, 24 VIII 2010, 5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E799614FF62F7CFFAC7F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E799617FF62F7EAFC40F877.text	857E879E8E799617FF62F7EAFC40F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius pentagonus d'Orbigny 1908	<div><p>3:  Caccobius (s. l.) pentagonus d'Orbigny, 1908, Karimama, Mamassi-Gourma, P. N. du W, 30 IX 2015, 3 mm. 4:  Caccobius (s. l.) punctatissimus von Harold, 1867, Karimama, P.N.duW, 28IX 2015, 4,5 mm.</p><p>Planche 38.</p></div>	https://treatment.plazi.org/id/857E879E8E799617FF62F7EAFC40F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7A9617FF47F82FFB3DF842.text	857E879E8E7A9617FF47F82FFB3DF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius signatipennis von Harold 1867	<div><p>1:  Caccobius (s. l.) signatipennis von Harold, 1867, Parc de la Penjari, 01-03 VI 2005, 4,4 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7A9617FF47F82FFB3DF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7A9617FF47F7CFFB45F7A2.text	857E879E8E7A9617FF47F7CFFB45F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius villiersi Cambefort 1975	<div><p>2:  Caccobius (s. l.) villiersi Cambefort, 1975, Parc de la Penjari, 01-03 VI 2005, 3,7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7A9617FF47F7CFFB45F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7A9616FF47F7EAFCA2F877.text	857E879E8E7A9616FF47F7EAFCA2F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Cleptocaccobius balthasarianus (Cambefort 1971)	<div><p>3:  Cleptocaccobius balthasarianus (Cambefort, 1971), ♂, P. N. du W, piste Keremou-Koudou, 02 VI 2004, 2,7 mm. 4:  Cleptocaccobius convexifrons (Raffray, 1877), ♂, Parc de la Penjari, vers Tanoungou, 04 VI 2005, 3,2 mm.</p><p>Planche 39.</p></div>	https://treatment.plazi.org/id/857E879E8E7A9616FF47F7EAFCA2F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7B9616FF52F82FFADFF842.text	857E879E8E7B9616FF52F82FFADFF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Cleptocaccobius dorbignyi Cambefort 1984	<div><p>1:  Cleptocaccobius dorbignyi Cambefort, 1984, P. N. du W, réserve de la Djona, IX 2003, 3,3 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7B9616FF52F82FFADFF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7B9616FF52F7CFFA97F781.text	857E879E8E7B9616FF52F7CFFA97F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Cleptocaccobius signaticollis (d'Orbigny 1902)	<div><p>2:  Cleptocaccobius signaticollis (d'Orbigny, 1902), ♂, Batia, P. N. Pendjari, mare Bali, 07 XI 2012, 2,8 mm. 3:  Cleptocaccobius uniseries (d’Orbigny, 1905), ♂, Pénéssoulou, forêt de Saramanga, 25 V 2013, 3 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E7B9616FF52F7CFFA97F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E7B9609FF52F78AFCA3F877.text	857E879E8E7B9609FF52F78AFCA3F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Diastellopalpus tridens (Fabricius 1781)	<div><p>4:  Diastellopalpus tridens (Fabricius, 1781), ♀, Pénéssoulou, forêt de Pénélan, 16 VI 2016, 16 mm.</p><p>Planche 40.</p></div>	https://treatment.plazi.org/id/857E879E8E7B9609FF52F78AFCA3F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E649609FEA9F82FFBA3F842.text	857E879E8E649609FEA9F82FFBA3F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Diastellopalpus tridens (Fabricius 1781)	<div><p>1:  Diastellopalpus tridens (Fabricius, 1781),   ♂,  Dan, bord du Zou, 17 V 2013, 17 mm  .</p></div>	https://treatment.plazi.org/id/857E879E8E649609FEA9F82FFBA3F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E649609FEA9F7CFFB2CF781.text	857E879E8E649609FEA9F7CFFB2CF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Diastellopalpus tridens (Fabricius 1781)	<div><p>2:  Diastellopalpus tridens (Fabricius, 1781), ♂, forme noire, Attogon, forêt de Niaouli, 24 X 2019, 15 mm. 3:  Proagoderus auratus (Fabricius, 1801), ♂, Batia, P. N. de la Pendjari, 18 VIII 2010, 15 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E649609FEA9F7CFFB2CF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E649608FEA9F78AFC87F877.text	857E879E8E649608FEA9F78AFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Proagoderus auratus (Fabricius 1801)	<div><p>4:  Proagoderus auratus (Fabricius, 1801), ♀, Batia, P. N. de la Pendjari, 18 VIII 2010, 13 mm.</p><p>Planche 41.</p></div>	https://treatment.plazi.org/id/857E879E8E649608FEA9F78AFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E659608FEAFF82FFB13F842.text	857E879E8E659608FEAFF82FFB13F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Proagoderus patinatus Marcus 1917	<div><p>1:  Proagoderus patinatus Marcus, 1917, ♂, Batia, Parc de la Penjari, 26 VIII 2010, 17 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E659608FEAFF82FFB13F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E659608FEAFF7CFFB6BF7A2.text	857E879E8E659608FEAFF7CFFB6BF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Proagoderus patinatus Marcus 1917	<div><p>2:  Proagoderus patinatus Marcus, 1917, ♀, Batia, Parc de la Penjari, 09 XI 2015, 15 mm</p></div>	https://treatment.plazi.org/id/857E879E8E659608FEAFF7CFFB6BF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E659608FEAFF7EAFB3FF7E1.text	857E879E8E659608FEAFF7EAFB3FF7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Proagoderus rarus Guerin-Meneville 1829	<div><p>3:  
Proagoderus rarus 
Guérin-Méneville, 1829, ♂, [R. D. C.,  Katanga, Kapanga, IX 1973, 15,8 mm]. 4:  Afrostrandius adspersus (d’Orbigny, 1908), ♂, [Côte d'Ivoire, Zamou, IV 1998, 12,2 mm].</p></div>	https://treatment.plazi.org/id/857E879E8E659608FEAFF7EAFB3FF7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E66960BFF0FF82FFAE7F842.text	857E879E8E66960BFF0FF82FFAE7F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Bicornonthophagus cornifrons (Thomson 1858)	<div><p>1:  Bicornonthophagus cornifrons (Thomson, 1858), ♂, Houéyogbé, forêt de Houéyogbé, 02 IV 2013, 7,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E66960BFF0FF82FFAE7F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E66960BFF0FF7CFFAFBF7A2.text	857E879E8E66960BFF0FF7CFFAFBF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Bicornonthophagus cornifrons (Thomson 1858)	<div><p>2:  Bicornonthophagus cornifrons (Thomson, 1858), ♀, Houéyogbé, forêt de Houéyogbé, 02 IV 2013, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E66960BFF0FF7CFFAFBF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E66960BFF0FF7EAFAA1F7E1.text	857E879E8E66960BFF0FF7EAFAA1F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Bicornonthophagus decorsei (d'Orbigny 1908)	<div><p>3:  Bicornonthophagus decorsei (d'Orbigny, 1908),   ♂, [P. N. W, Tapoa Djerma (Burkina Faso), 01 VI 2004, 7,3 mm]. 4:  Bicornonthophagus decorsei (d'Orbigny, 1908), ♀, [P. N. W, Tapoa Djerma (Burkina Faso), 01 VI 2004, 6,9 mm]  .</p></div>	https://treatment.plazi.org/id/857E879E8E66960BFF0FF7EAFAA1F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E67960AFF77F82FFB50F781.text	857E879E8E67960AFF77F82FFB50F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Bicornonthophagus reticulatus (d'Orbigny 1902)	<div><p>1:  Bicornonthophagus reticulatus (d'Orbigny, 1902), ♀, Pénéssoulou, forêt de Pénélan, 16 VI 2016, 7 mm. 2:  Bicornonthophagus reticulatus (d'Orbigny, 1902), ♂, Batia, Parc de la Penjari, 18 VIII 2010, 8 mm.</p><p>3: Digitontophagus eucata Génier, 2017, ♂, Karimama, P. N. du W, 28 IX 2015, 12 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E67960AFF77F82FFB50F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E67960AFF77F78AFAD3F7E1.text	857E879E8E67960AFF77F78AFAD3F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Digitonthophagus falciger Genier 2017	<div><p>4:  Digitonthophagus falciger Génier, 2017, ♂, Batia, Tanguita, P. N. Pendjari, 08 XI 2015, 10,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E67960AFF77F78AFAD3F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E60960DFF37F82FFB95F842.text	857E879E8E60960DFF37F82FFB95F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Digitonthophagus fimator Genier 2017	<div><p>1:  Digitonthophagus fimator Génier, 2017, ♂, Lokossa, Djetokpa, 12 VIII 2007, 10,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E60960DFF37F82FFB95F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E60960DFF37F7CFFBA7F7A2.text	857E879E8E60960DFF37F7CFFBA7F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Euonthophagus Balthasar 1959	<div><p>2:  Euonthophagus complexe carbonarius, ♀ Batia, Parc de la Penjari,, 29 VII 2009, 8 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E60960DFF37F7CFFBA7F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E60960DFF37F7EAFC07F781.text	857E879E8E60960DFF37F7EAFC07F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Euonthophagus Balthasar 1959	<div><p>3:  Euonthophagus complexe carbonarius, ♂, Pénéssoulou, 22 V 2013, 9 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E60960DFF37F7EAFC07F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E60960CFF37F78AFC80F877.text	857E879E8E60960CFF37F78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furconthophagus monforti Josso & Prevost 2006	<div><p>4:  Furconthophagus monforti Josso &amp; Prévost, 2006, ♂, holotype, Tanougou, Parc de la Pendjari, 04 VI 2006, 4,5 mm.</p><p>Planche 45.</p></div>	https://treatment.plazi.org/id/857E879E8E60960CFF37F78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E61960CFF0FF82FFB64F7A2.text	857E879E8E61960CFF0FF82FFB64F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furconthophagus monforti Josso & Prevost 2006	<div><p>1:  Furconthophagus monforti Josso &amp; Prévost, 2006, ♀, paratype, Tanougou, Parc de la Pendjari, 04 VI 2006, 4,3 mm. 2:  Furcontophagus flaviclava (d'Orbigny, 1902), ♂, Batia, Parc de la Penjari, 26 VIII 2010, 5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E61960CFF0FF82FFB64F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E61960CFF0FF78AFB31F7E1.text	857E879E8E61960CFF0FF78AFB31F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furconthophagus ulula (Balthasar 1966)	<div><p>4:  Furconthophagus ulula (Balthasar, 1966), ♂, Pénéssoulou, forêt de Saramanga, 24 V 2013, 4,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E61960CFF0FF78AFB31F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E61960CFF0FF7EAFB72F781.text	857E879E8E61960CFF0FF7EAFB72F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furcontophagus flaviclava (d'Orbigny 1902)	<div><p>3:  Furcontophagus flaviclava (d'Orbigny, 1902),  ♂, Kouandé, antenne ORTB, 05 XI 2015, 4,5 mm .</p></div>	https://treatment.plazi.org/id/857E879E8E61960CFF0FF7EAFB72F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E62960FFF3FF82FFB73F842.text	857E879E8E62960FFF3FF82FFB73F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furconthophagus ulula (Balthasar 1966)	<div><p>1:  Furconthophagus ulula (Balthasar, 1966), ♀, Pénéssoulou, forêt de Pénélan, 15 VI 2016, 4 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E62960FFF3FF82FFB73F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E62960FFF3FF7CFFB7CF781.text	857E879E8E62960FFF3FF7CFFB7CF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Furconthophagus variegatus (Fabricius 1798)	<div><p>2:  Furconthophagus variegatus (Fabricius, 1798), ♂, Houéyogbé, forêt de Houéyogbé, 16 VI 2016, 4,5 mm. 3:  Furconthophagus variegatus (Fabricius, 1798), ♀, Zinvié, vallée du Sitatunga, 19 V 2016, 5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E62960FFF3FF7CFFB7CF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E62960FFF3FF78AFB04F7E1.text	857E879E8E62960FFF3FF78AFB04F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gonionthophagus rufonotatus (d'Orbigny 1902)	<div><p>4:  Gonionthophagus rufonotatus (d'Orbigny, 1902),  ♂, Kouandé, antenne ORTB, 05 XI 2015, 7,5 mm .</p></div>	https://treatment.plazi.org/id/857E879E8E62960FFF3FF78AFB04F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E63960EFF57F82FFBBAF7A2.text	857E879E8E63960EFF57F82FFBBAF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gonionthophagus stehliki (Balthasar 1966)	<div><p>1:  Gonionthophagus stehliki (Balthasar, 1966),   ♂, P. N. Pendjari, Camp Numi, 09 XI 2015, 6,5 mm. 2:  Gonocyphus obliquus (Olivier, 1789), ♂,  Dan, bord du Zou, 17 VII 2013, 16 mm  .</p></div>	https://treatment.plazi.org/id/857E879E8E63960EFF57F82FFBBAF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E63960EFF57F7EAFBB9F781.text	857E879E8E63960EFF57F7EAFBB9F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Gonocyphus obliquus (Olivier 1789)	<div><p>3:  Gonocyphus obliquus (Olivier, 1789), ♀, Savalou, Kpataba, 14 XII 2016, 14 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E63960EFF57F7EAFBB9F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E63960EFF57F78AFAF2F7E1.text	857E879E8E63960EFF57F78AFAF2F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Hamontophagus bituberculatus (Olivier 1789)	<div><p>4:  Hamontophagus bituberculatus (Olivier, 1789), ♂, Bohicon, quartier Soglogon, 28 IV 2013, 8,5mm.</p></div>	https://treatment.plazi.org/id/857E879E8E63960EFF57F78AFAF2F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6C9601FF61F82FFAEFF842.text	857E879E8E6C9601FF61F82FFAEFF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Hyalonthophagus nigroviolaceus (d'Orbigny 1902)	<div><p>1:  Hyalonthophagus nigroviolaceus (d'Orbigny, 1902), ♂, Batia, Parc de la Pendjari, 05 VI 2005, 6,6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6C9601FF61F82FFAEFF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6C9601FF61F7CFFB10F7A2.text	857E879E8E6C9601FF61F7CFFB10F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Hyalonthophagus nigroviolaceus (d'Orbigny 1902)	<div><p>2:  Hyalonthophagus nigroviolaceus (d'Orbigny, 1902), ♀, Parc de la Pendjari, VI 2005, 6,1 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6C9601FF61F7CFFB10F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6C9601FF61F7EAFAEDF7E1.text	857E879E8E6C9601FF61F7EAFAEDF7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Hyalonthophagus pseudoalcyon (d'Orbigny 1913)	<div><p>3:  Hyalonthophagus pseudoalcyon (d'Orbigny, 1913),   ♂, [ Réserve de Fathala (Sénégal), 12 VII 2010, 10 mm]. 4:  Hyalonthophagus pseudoalcyon (d'Orbigny, 1913), ♀, Batia, Parc de la Pendjari, 27 VII 2009, 8,5 mm  .</p></div>	https://treatment.plazi.org/id/857E879E8E6C9601FF61F7EAFAEDF7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6D9600FF72F82FFBAEF842.text	857E879E8E6D9600FF72F82FFBAEF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Micronthophagus depressipennis (Cambefort 1975)	<div><p>1:  Micronthophagus depressipennis (Cambefort, 1975), ♂, Bohicon, 8 VI 2018, 5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6D9600FF72F82FFBAEF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6D9600FF72F7EAFADBF781.text	857E879E8E6D9600FF72F7EAFADBF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Micronthophagus nigriceps (Raffray 1877)	<div><p>3:  Micronthophagus nigriceps (Raffray, 1877), ♀, Mamassi Gourma, P. N. du W, 30 IX 2015, 6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6D9600FF72F7EAFADBF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6D9600FF72F7CFFAF6F7A2.text	857E879E8E6D9600FF72F7CFFAF6F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Micronthophagus ochreatus (d'Orbigny 1897)	<div><p>2:  Micronthophagus ochreatus (d’Orbigny, 1897), ♀, Mamassi Gourma, P. N. du W, 30 IX 2015, 4 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6D9600FF72F7CFFAF6F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6D9603FF72F78AFC80F877.text	857E879E8E6D9603FF72F78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Parascatonomus (Morettoeus) maculatus (Fabricius 1801)	<div><p>4:  Parascatonomus (Morettoeus) maculatus (Fabricius, 1801),  Est de Kandi Araoun SAA, VI 2001, 10 mm .</p><p>Planche 50.</p></div>	https://treatment.plazi.org/id/857E879E8E6D9603FF72F78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6E9603FF02F82FFADAF842.text	857E879E8E6E9603FF02F82FFADAF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Parascatonomus (Morettoeus) chloroderus (d'Orbigny 1905)	<div><p>1:  Parascatonomus (Morettoeus) chloroderus (d’Orbigny, 1905), Bembereke, Gando, VII 2001, 8,6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6E9603FF02F82FFADAF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6E9603FF02F7CFFBA1F7A2.text	857E879E8E6E9603FF02F7CFFBA1F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus curtipilis d'Orbigny 1905	<div><p>2:  Tiaronthophagus curtipilis d'Orbigny, 1905, ♀, forêt de Houéyogbé, 22 VIII 2011, 5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6E9603FF02F7CFFBA1F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6E9602FF02F7EAFCEAF877.text	857E879E8E6E9602FF02F7EAFCEAF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus curtipilis d'Orbigny 1905	<div><p>3:  
Tiaronthophagus curtipilis 
d'Orbigny, 1905, ♂, [P. N. du W, chutes de Koudou (Burkina Faso), VI 2004, 6,8 mm]. 4:  Tiaronthophagus lamtoensis (Cambefort, 1984), ♀, Pénéssoulou, forêt de Pénélan, 03 VII 2013, 7 mm.</p><p>Planche 51.</p></div>	https://treatment.plazi.org/id/857E879E8E6E9602FF02F7EAFCEAF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6F9602FF07F78AFAD6F7E1.text	857E879E8E6F9602FF07F78AFAD6F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus flexicornis (d'Orbigny 1902)	<div><p>4:  Tiaronthophagus flexicornis (d'Orbigny, 1902), ♀, Pénéssoulou, forêt de Pénélan, 23 V 2013, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6F9602FF07F78AFAD6F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6F9602FF07F82FFAD6F781.text	857E879E8E6F9602FF07F82FFAD6F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus liberianus (van Lansberge 1883)	<div><p>1:  Tiaronthophagus liberianus (van Lansberge, 1883), ♂, Zogbodomey, forêt de la Lama, 08 IX 2010, 9 mm. 2:  Tiaronthophagus liberianus (van Lansberge, 1883), ♀, Zogbodomey, forêt de la Lama, 08 IX 2010, 7 mm. 3:  Tiaronthophagus flexicornis (d'Orbigny, 1902), ♂, Pénéssoulou, forêt de Pénélan, 28 V 2013, 8 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6F9602FF07F82FFAD6F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E689605FF62F82FFB54F842.text	857E879E8E689605FF62F82FFB54F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus viridiaerus (d'Orbigny 1908)	<div><p>1:  Tiaronthophagus viridiaerus (d’Orbigny, 1908), ♂, Savalou, Kpataba, 17 VI 2016, 6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E689605FF62F82FFB54F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E689605FF62F7CFFB25F7A2.text	857E879E8E689605FF62F7CFFB25F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus viridiaerus (d'Orbigny 1908)	<div><p>2:  Tiaronthophagus viridiaerus (d’Orbigny, 1908), ♂, Pénéssoulou, forêt de Pénélan, 27 V 2013, 6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E689605FF62F7CFFB25F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E689605FF62F7EAFB50F7E1.text	857E879E8E689605FF62F7EAFB50F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus viridiaerus (d'Orbigny 1908)	<div><p>3:  Tiaronthophagus viridiaerus (d’Orbigny, 1908), ♂, Tanoungou, P. N. de la Pendjari, VI 2005, 6,1 mm. 4:  Phalops barbicornis van Lansberge, 1883, ♂, Batia, P. N. Pendjari, 24 VII 2009, 10 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E689605FF62F7EAFB50F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E699604FE81F82FFB0BF781.text	857E879E8E699604FE81F82FFB0BF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Phalops barbicornis van Lansberge 1883	<div><p>1:  Phalops barbicornis van Lansberge, 1883, ♀, Batia, P. N. Pendjari, 24 VII 2009, 10 mm. 2:  Phalops batesi batesi von Harold, 1867, ♂, Batia, P. N. Pendjari, 27 VII 2009, 11,5 mm. 3:  Phalops batesi batesi von Harold, 1867, ♀, Batia, P. N. Pendjari, 24 VII 2009, 12 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E699604FE81F82FFB0BF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E699607FE81F78AFB7DF842.text	857E879E8E699607FE81F78AFB7DF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Phalops fimbriatus (Klug 1835)	<div><p>4:  Phalops fimbriatus (Klug, 1835),   ♂, [ Réserve de Fathala (Sénégal), 24 VI 2009, 11 mm]. 1:  Phalops iphis (Olivier, 1789), ♂, P.N.Pendjari,forêtdeBondjagou, 26VII 2009, 9 mm  .</p></div>	https://treatment.plazi.org/id/857E879E8E699607FE81F78AFB7DF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6A9607FEA2F7CFFBDDF7E1.text	857E879E8E6A9607FEA2F7CFFBDDF7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Phalops vanellus van Lansberge 1883	<div><p>2:  Phalops vanellus van Lansberge, ♂, 1883, Pénéssoulou, forêt de Saramanga, 25 V 2013, 10 mm. 3:  Phalops vanellus van Lansberge, 1883, ♀, Pénéssoulou, forêt de Saramanga, 25 V 2013, 12 mm. 4:  Trichontophagus juvencus (Klug, 1835), Lokossa, 24 III 2006, 6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6A9607FEA2F7CFFBDDF7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6B9606FF71F7CFFA84F7A2.text	857E879E8E6B9606FF71F7CFFA84F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Trichonthophagus triacanthus (Castelnau 1840)	<div><p>2:  Trichonthophagus triacanthus (Castelnau, 1840),   ♂, [ Réserve de Fathala (Sénégal), 14 VII 2010, 8 mm]  .</p></div>	https://treatment.plazi.org/id/857E879E8E6B9606FF71F7CFFA84F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6B9606FF71F7EAFA82F781.text	857E879E8E6B9606FF71F7EAFA82F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Trichonthophagus triacanthus (Castelnau 1840)	<div><p>3:  Trichonthophagus triacanthus (Castelnau, 1840), ♀, Pénéssoulou, forêt de Saramanga, 25 V 2013, 8 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6B9606FF71F7EAFA82F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6B9606FF71F82FFAF2F842.text	857E879E8E6B9606FF71F82FFAF2F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Trichontophagus mediofuscatus (d'Orbigny 1908)	<div><p>1:  Trichontophagus mediofuscatus (d'Orbigny, 1908), ♀, Pénéssoulou, forêt de Pénélan, 23 V 2013, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E6B9606FF71F82FFAF2F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E6B9639FF71F78AFC80F877.text	857E879E8E6B9639FF71F78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Trichontophagus gonzalohalffteri Moretto 2023	<div><p>4:  Trichontophagus gonzalohalffteri Moretto, 2023, ♀, paratype, N’Dali, forêt classée de N’Dali, 11 XI 2015, 7 mm.</p><p>Planche 56.</p></div>	https://treatment.plazi.org/id/857E879E8E6B9639FF71F78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E549639FF5FF82FFB5DF842.text	857E879E8E549639FF5FF82FFB5DF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus antennalis Frey 1961	<div><p>1:  Onthophagus (s. l.) antennalis Frey, 1961, Batia, P. N. Pendjari, VIII 2010, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E549639FF5FF82FFB5DF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E549639FF5FF7CFFB8AF7A2.text	857E879E8E549639FF5FF7CFFB8AF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus borassi Cambefort 1984	<div><p>2:  Onthophagus (s. l.) borassi Cambefort, 1984, Allada, Donou, 10 VI 2016, 4 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E549639FF5FF7CFFB8AF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E549638FF5FF7EAFC96F862.text	857E879E8E549638FF5FF7EAFC96F862.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus cribellum d'Orbigny 1902	<div><p>3:  
Onthophagus (s. l.) cribellum 
d'Orbigny, 1902, Pénéssoulou, forêt de Saramanga, 25 V 2013, 5 mm. 4:  Onthophagus (s. l.) cupreovirens d'Orbigny, 1904, ♂,  Dan, bord du Zou, 17 V 2013, 5 mm.</p><p>Planche 57.</p></div>	https://treatment.plazi.org/id/857E879E8E549638FF5FF7EAFC96F862	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E559638FEB1F822FC22F799.text	857E879E8E559638FEB1F822FC22F799.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>1:  Onthophagus (s. l.) sp. (sp. 1, gr. 2), Pénéssoulou, forêt de Pénélan, 15 VI 2016, 5 mm. 2:  Onthophagus (s. l.) sp. (sp. 2, gr. 2), Lokossa, 18 III 2006, 3,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E559638FEB1F822FC22F799	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E55963BFEB1F7E0FC80F877.text	857E879E8E55963BFEB1F7E0FC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>3:  Onthophagus (s. l.) sp. (sp. 3, gr. 2), Houéyogbé, forêt de Houéyogbé, 22 VIII 2011, 3 mm. 4:  Onthophagus (s. l.) vinctus Erichson, 1843, ♀, Savalou, Kpataba, 13 XII 2016, 6,5 mm.</p><p>Planche 58.</p></div>	https://treatment.plazi.org/id/857E879E8E55963BFEB1F7E0FC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E56963BFF32F7EAFB13F781.text	857E879E8E56963BFF32F7EAFB13F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus atridorsis d'Orbigny 1902	<div><p>3:  Onthophagus (s. l.) atridorsis d'Orbigny, 1902, P. N. du W, chutes de Koudou, 12 X 2004, 2,9 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E56963BFF32F7EAFB13F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E56963BFF32F78AFB72F7E1.text	857E879E8E56963BFF32F78AFB72F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>4:  Onthophagus (s. l.) complexe pullus Roth, 1851, ♀, Karimama, P. N. du W, 28 IX 2015, 3 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E56963BFF32F78AFB72F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E56963BFF32F82FFB1BF7A2.text	857E879E8E56963BFF32F82FFB1BF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus paravinctus Tagliaferri et Moretto 2012	<div><p>1:  Onthophagus (s. l.) paravinctus Tagliaferri et Moretto, 2012, ♂, Batia, P. N. Pendjari, Camp Numi, 03 XI 2015, 6 mm. 2:  Onthophagus (s. l.) picatus d'Orbigny, 1902, Houéyogbé, forêt de Houéyogbé, 02 IV 2013, 6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E56963BFF32F82FFB1BF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E57963DFEE9F82FFCAFF877.text	857E879E8E57963DFEE9F82FFCAFF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus lioides d'Orbigny 1902	<div><p>1:  Onthophagus (s. l.) lioides d'Orbigny, 1902, ♂, Karimama, P. N. du W, 28 IX 2015, 4 mm. 2:  Onthophagus (s. l.) notatus d'Orbigny, 1902, ♀, Batia, P. N. Pendjari, 27 VII 2009, 4 mm. 3:  Onthophagus tersipennis d'Orbigny, 1902, ♂, Karimama, P. N. du W, 28 IX 2015, 3 mm. 4:  Onthophagus (s. l.) brevipennis d'Orbigny, 1902, [Ifan 1948, 5 mm].</p><p>Planche 60.</p></div>	https://treatment.plazi.org/id/857E879E8E57963DFEE9F82FFCAFF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E50963DFEA9F82FFB3FF781.text	857E879E8E50963DFEA9F82FFB3FF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus Latreille 1802	<div><p>1:  Onthophagus (s. l.) complexe  cupreus von Harold, 1880, ♂, Batia, P.N.Pendjari, 26VIII 2010, 11 mm. 2:  Onthophagus (s. l.) complexe  cupreus von Harold, 1880, ♀, Batia, P.N.Pendjari, 18VIII 2010, 11 mm. 3:  Onthophagus (s. l.) feai d'Orbigny, 1905, Zogbodomey, forêt de la Lama, X 2010, 5,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E50963DFEA9F82FFB3FF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E50963CFEA9F78AFC80F877.text	857E879E8E50963CFEA9F78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus sellatus Klug 1845	<div><p>4:  
Onthophagus (s. l.) sellatus 
Klug, 1845, ♂,  Sahé, bord du Couffo, 14 V 2013, 5 mm.</p><p>Planche 61.</p></div>	https://treatment.plazi.org/id/857E879E8E50963CFEA9F78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E51963CFF67F82FFB1BF7A2.text	857E879E8E51963CFF67F82FFB1BF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus ecopas Josso et Prevost 2006	<div><p>1:  Onthophagus (s. l.) ecopas Josso et Prévost, 2006, ♀, Mamassi Gourma, P. N. du W, 30 IX 2015, 6,4 mm. 2:  Onthophagus (s. l.) latiggiber d'Orbigny, 1902, Attogon, forêt de Niaouli, 14 VI 2016, 8 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E51963CFF67F82FFB1BF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E51963CFF67F7EAFB1BF781.text	857E879E8E51963CFF67F7EAFB1BF781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus latiggiber d'Orbigny 1902	<div><p>3:  Onthophagus (s. l.) latiggiber d'Orbigny, 1902, Attogon, forêt de Niaouli, 07 VI 2016, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E51963CFF67F7EAFB1BF781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E51963FFF67F78AFC87F877.text	857E879E8E51963FFF67F78AFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus trinominatus Goidanich 1926	<div><p>4:  Onthophagus (s. l.) trinominatus Goidanich, 1926, Houéyogbé, forêt de Houéyogbé, 25 VI 2016, 7,5 mm.</p><p>Planche 62.</p></div>	https://treatment.plazi.org/id/857E879E8E51963FFF67F78AFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E52963FFF4AF7CFFB8DF7A2.text	857E879E8E52963FFF4AF7CFFB8DF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus bimarginatus d'Orbigny 1902	<div><p>2:  Onthophagus (s. l.) bimarginatus d'Orbigny, 1902, Bohicon, 15 VI 2009, 9 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E52963FFF4AF7CFFB8DF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E52963EFF4AF7EAFB25F842.text	857E879E8E52963EFF4AF7EAFB25F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus bimarginatus d'Orbigny 1902	<div><p>3:  Onthophagus (s. l.) bimarginatus d'Orbigny, 1902, Houéyogbé, forêt de Houéyogbé, 08 V 2013, 10 mm. 4:  Onthophagus (s. l.) callosipennis Boucomont, 1930, Pénéssoulou, forêt de Pénélan, 02 VII 2013, 5 mm. 1:  Onthophagus (s. l.) ghanensis Balthasar, 1966, Batia, P. N. Pendjari, 26 VIII 2010, 4,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E52963EFF4AF7EAFB25F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E52963FFF4AF82FFAC3F842.text	857E879E8E52963FFF4AF82FFAC3F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus trinominatus Goidanich 1926	<div><p>1:  Onthophagus (s. l.) trinominatus Goidanich, 1926, Pénéssoulou, forêt de Pénélan, 03 VII 2013, 7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E52963FFF4AF82FFAC3F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E53963EFEAFF7CFFBB0F7A2.text	857E879E8E53963EFEAFF7CFFBB0F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus grandifrons d'Orbigny 1907	<div><p>2:  Onthophagus (s. l.) grandifrons d'Orbigny, 1907, Lokossa, 18 III 2006, 5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E53963EFEAFF7CFFBB0F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E53963EFEAFF7EAFB26F781.text	857E879E8E53963EFEAFF7EAFB26F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus mankonoensis Balthasar 1966	<div><p>3:  Onthophagus (s. l.) mankonoensis Balthasar, 1966, Batia, P. N. Pendjari, 08 X 2013, 9 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E53963EFEAFF7EAFB26F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E53963EFEAFF78AFAA0F7E1.text	857E879E8E53963EFEAFF78AFAA0F7E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus mankonoensis Balthasar 1966	<div><p>4:  
Onthophagus (s. l.) mankonoensis 
Balthasar, 1966, Burkina Faso, Parc W, Tapoa Djerma, 01 06 2004, 9 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E53963EFEAFF78AFAA0F7E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5C9631FEA9F82FFB02F842.text	857E879E8E5C9631FEA9F82FFB02F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus obenbergeri Balthasar 1965	<div><p>1:  Onthophagus (s. l.) obenbergeri Balthasar, 1965, Batia, P. N. Pendjari, VI 2005, 13,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E5C9631FEA9F82FFB02F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5C9631FEA9F7CFFAD9F7A2.text	857E879E8E5C9631FEA9F7CFFAD9F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus semivirescens d'Orbigny 1902	<div><p>2:  Onthophagus (s. l.) semivirescens d'Orbigny, 1902, ♀, Pobé, forêt de Pobé, 13 II 2017, 5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E5C9631FEA9F7CFFAD9F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5C9630FEA9F7EAFCBFF877.text	857E879E8E5C9630FEA9F7EAFCBFF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus viridicatus d'Orbigny 1902	<div><p>3:  Onthophagus (s. l.) viridicatus d'Orbigny, 1902, ♂, Attogon, forêt de Niaouli, 14 VI 2016, 5 mm. 4:  Onthophagus (s. l.) sp., Batia, P. N. Pendjari, 24 VIII 2010, 3,5 mm.</p><p>Planche 65.</p></div>	https://treatment.plazi.org/id/857E879E8E5C9630FEA9F7EAFCBFF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5D9630FF52F7CFFB3AF7A2.text	857E879E8E5D9630FF52F7CFFB3AF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus circumdatus d'Orbigny 1913	<div><p>2:  Onthophagus (s. l.) circumdatus d'Orbigny, 1913, chaine de l'Atakora, 01 VI 2004, 4,8 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E5D9630FF52F7CFFB3AF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5D9630FF52F82FFB2AF842.text	857E879E8E5D9630FF52F82FFB2AF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus margaritifer d'Orbigny 1898	<div><p>1:  Onthophagus (s. l.) margaritifer d’Orbigny, 1898, Karimama, P. N. du W, 28 IX 2015, 4 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E5D9630FF52F82FFB2AF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5D9630FF52F7EAFAF9F781.text	857E879E8E5D9630FF52F7EAFAF9F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus tripartitus d'Orbigny 1902	<div><p>3:  Onthophagus (s. l.) tripartitus d'Orbigny, 1902, Bohicon, quartier Soglogon, 27 IV 2013, 4,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E5D9630FF52F7EAFAF9F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5D9633FF52F78AFCA3F877.text	857E879E8E5D9633FF52F78AFCA3F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus fimetarius Roth 1851	<div><p>4:  Onthophagus (s. l.) fimetarius Roth, 1851, ♂, Batia, P. N. Pendjari, Camp Numi, 07 XI 2015, 6 mm.</p><p>Planche 66.</p></div>	https://treatment.plazi.org/id/857E879E8E5D9633FF52F78AFCA3F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5E9633FF5FF82FFB7CF842.text	857E879E8E5E9633FF5FF82FFB7CF842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus lobi Cambefort 1984	<div><p>1:  Onthophagus (s. l.) lobi Cambefort, 1984, ♂, Karimama, P.N.duW, 28IX 2015, 6 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E5E9633FF5FF82FFB7CF842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5E9633FF5FF7CFFAC2F7A2.text	857E879E8E5E9633FF5FF7CFFAC2F7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus lobi Cambefort 1984	<div><p>2:  Onthophagus (s. l.) lobi Cambefort, 1984, ♂, Pénéssoulou, forêt de Pénélan, 15 VI 2016, 6,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E5E9633FF5FF7CFFAC2F7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5E9632FF5FF7EAFC80F877.text	857E879E8E5E9632FF5FF7EAFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus lutaticollis d'Orbigny 1907	<div><p>3:  Onthophagus (s. l.) lutaticollis d'Orbigny, 1907, ♀, Pénéssoulou, forêt de Saramanga, 25 V 2013, 5,5 mm. 4:  Onthophagus (s. l.) lutaticollis d'Orbigny, 1907, ♂, Batia, P.N.Pendjari, 27VII 2009, 6 mm.</p><p>Planche 67.</p></div>	https://treatment.plazi.org/id/857E879E8E5E9632FF5FF7EAFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5F9632FF19F82FFB01F842.text	857E879E8E5F9632FF19F82FFB01F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus mucronifer d'Orbigny 1905	<div><p>1:  Onthophagus (s. l.) mucronifer d'Orbigny, 1905, ♀, Bohicon, quartier Soglogon, 27 IV 2013, 5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E5F9632FF19F82FFB01F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5F9632FF19F7CFFB1BF7A2.text	857E879E8E5F9632FF19F7CFFB1BF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus proximus d'Orbigny 1908	<div><p>2:  Onthophagus (s. l.) proximus d'Orbigny, 1908, ♂, P.N.Pendjari,hôtelPendjari, VI2005, 6,7 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E5F9632FF19F7CFFB1BF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5F9635FF19F7EAFC87F877.text	857E879E8E5F9635FF19F7EAFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus proximus d'Orbigny 1908	<div><p>3:  Onthophagus (s. l.) proximus d'Orbigny, 1908, ♀, P.N.duW, réservedelaDjona,AlphaKouara, IX2003, 6,5 mm. 4:  Onthophagus (s. l.) sellatulus d'Orbigny, 1902, ♂, Tanguiéta, chaîne de l’Atakora, 01VI2004, 4,8 mm.</p><p>Planche 68.</p></div>	https://treatment.plazi.org/id/857E879E8E5F9635FF19F7EAFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E589635FF71F7CFFBB0F781.text	857E879E8E589635FF71F7CFFBB0F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus bandamai Cambefort 1984	<div><p>2:  Onthophagus (s. l.) bandamai Cambefort, 1984, ♂, Pénéssoulou, forêt de Goutansoukpa, 30 V 2012, 6 mm. 3:  Onthophagus (s. l.) bidens (Olivier, 1789), ♂, Pobé, forêt de Pobé, 18 VI 2013, 8 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E589635FF71F7CFFBB0F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E589635FF71F82FFAC0F842.text	857E879E8E589635FF71F82FFAC0F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus sellatulus d'Orbigny 1902	<div><p>1:  Onthophagus (s. l.) sellatulus d'Orbigny, 1902, ♀, P.N.duW,chutesdeKoudou, 12 X 2003, 4,5 mm.</p></div>	https://treatment.plazi.org/id/857E879E8E589635FF71F82FFAC0F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E589634FF71F78AFC80F877.text	857E879E8E589634FF71F78AFC80F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus bidens (Olivier 1789)	<div><p>4:  Onthophagus (s. l.) bidens (Olivier, 1789), ♀, Pobé, forêt de Pobé, 13 II 2017, 8 mm.</p><p>Planche 69.</p></div>	https://treatment.plazi.org/id/857E879E8E589634FF71F78AFC80F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E599634FF5FF82FFAD4F781.text	857E879E8E599634FF5FF82FFAD4F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus loxodontae Cambefort 1984	<div><p>1:  Onthophagus (s. l.) loxodontae Cambefort, 1984, P. N, Pendjari, hôtel Pendjari, VI 2005, 4,3 mm. 2:  Onthophagus (s. l.) mucronatus Thomson, 1858, ♂, Attogon, forêt de Niaouli, 07 VI 2016, 5,5 mm. 3:  Onthophagus (s. l.) mucronatus Thomson, 1858, ♀, Attogon, forêt de Niaouli, 07 VI 2016, 5 mm</p></div>	https://treatment.plazi.org/id/857E879E8E599634FF5FF82FFAD4F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E599637FF5FF78AFCBBF877.text	857E879E8E599637FF5FF78AFCBBF877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onitis unguiculatus (Olivier 1789)	<div><p>4:  Onitis unguiculatus (Olivier, 1789),   ♀, [Sénégal, Fatick,  Diouroup, 14-18 VIII 2007, 24 mm]  .</p><p>Planche 70.</p></div>	https://treatment.plazi.org/id/857E879E8E599637FF5FF78AFCBBF877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5A9637FF6AF7EAFAF9F781.text	857E879E8E5A9637FF6AF7EAFAF9F781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius morettoi Walter 2014	<div><p>3:  
Caccobius (s. l.) morettoi 
Walter, 2014, [Côte d’Ivoire,  Dabakalla, Ouandérama, 23-29 V 2016, 3 mm].</p></div>	https://treatment.plazi.org/id/857E879E8E5A9637FF6AF7EAFAF9F781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5A9637FF6AF7CFFBFAF7A2.text	857E879E8E5A9637FF6AF7CFFBFAF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Euoniticellus nasicornis (Reiche 1849)	<div><p>2:  Euoniticellus nasicornis (Reiche, 1849), ♂, [Kitega, Burindi, V 1969, 12 mm].</p></div>	https://treatment.plazi.org/id/857E879E8E5A9637FF6AF7CFFBFAF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5A9636FF6AF78AFC87F877.text	857E879E8E5A9636FF6AF78AFC87F877.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Parascatonomus (Morettoeus) imbellis (d'Orbigny 1905)	<div><p>4:  Parascatonomus (Morettoeus) imbellis (d’Orbigny, 1905),   [ Camerooun, Mts. Mandaea, Watir, VI 2001, 5 mm]  .</p><p>Planche 71.</p></div>	https://treatment.plazi.org/id/857E879E8E5A9636FF6AF78AFC87F877	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5B9636FF5AF82FFB72F842.text	857E879E8E5B9636FF5AF82FFB72F842.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus chrysoderus (d'Orbigny 1905)	<div><p>1:  Tiaronthophagus chrysoderus (d’Orbigny, 1905),  Kandi, NE Bénin, 21-23 VI 2001, 7 mm].</p></div>	https://treatment.plazi.org/id/857E879E8E5B9636FF5AF82FFB72F842	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5B9636FF5AF7CFFB2AF7A2.text	857E879E8E5B9636FF5AF7CFFB2AF7A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Tiaronthophagus rougonorum (Cambefort 1984)	<div><p>2:  Tiaronthophagus rougonorum (Cambefort, 1984),   [Sénégal,  Niokolo Koba, VII 1995, 6 mm]  .</p></div>	https://treatment.plazi.org/id/857E879E8E5B9636FF5AF7CFFB2AF7A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E5B9629FF5AF7EAFC80FBC9.text	857E879E8E5B9629FF5AF7EAFC80FBC9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus nodieri d'Orbigny 1908	<div><p>3:  
Onthophagus (s. l.) nodieri 
d'Orbigny, 1908, [Sénégal, Kedougou,  Dindéfélo, 18-19 XI 2011, 4 mm]. 4:  Onthophagus (s. l.) wensis Josso &amp; Prévost, 2006, [Côte d’Ivoire, Touba, 4-5 V 2018, 5 mm].</p><p>Planche 72.</p></div>	https://treatment.plazi.org/id/857E879E8E5B9629FF5AF7EAFC80FBC9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E449629FF32FB48FAF5FB27.text	857E879E8E449629FF32FB48FAF5FB27.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus mocquerysi d'Orbigny 1902	<div><p>1:  
Onthophagus (s. l.) mocquerysi 
d'Orbigny, 1902, [Côte d’Ivoire,  Korhogo, Koko, 22-23 VII 2014, 10 mm].</p></div>	https://treatment.plazi.org/id/857E879E8E449629FF32FB48FAF5FB27	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E449628FF32FB68FC87F821.text	857E879E8E449628FF32FB68FC87F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus cyanochlorus d'Orbigny 1902	<div><p>2:  
Onthophagus (s. l.) cyanochlorus 
d'Orbigny, 1902, ♂, [Côte d’Ivoire,  Boundiali, M’banto, 13-14 VII 2013, 5,5 mm].</p><p>Planche 73.</p></div>	https://treatment.plazi.org/id/857E879E8E449628FF32FB68FC87F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E45962BFFFFF860FC87F821.text	857E879E8E45962BFFFFF860FC87F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Parateuchus palemo (Olivier 1789)	<div><p>1.  Parateuchus palemo (Olivier, 1789) . 2.  Scarabaeus goryi (de Castelnau, 1840), ♂. 3.  Allogymnopleurus olivieri (de Castelnau, 1840), ♂. 4.  Allogymnopleurus umbrinus (Gerstaecker, 1871), ♂. 5.  Sisyphus (Neosisyphus) gladiator Arrow, 1927, ♀. 6.  Sisyphus (Neosisyphus) gladiator Arrow, 1927, ♂. 7.  Sisyphus (Neosisyphus) youngai Endrödi, 1973, ♂. 8.  Sisyphus (Neosisyphus) youngai Endrödi, 1973, ♀. 9.  Catharsius eteocles (de Castelnau, 1840), ♂. 10.  Catharsius guineensis Ferreira, 1963, ♂. 11.  Catharsius pseudolycaon Ferreira, 1960, ♂. 12.  Catharsius complexe sesostris Waterhouse, 1891, ♂.</p><p>Planche 74.</p></div>	https://treatment.plazi.org/id/857E879E8E45962BFFFFF860FC87F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E46962AFFFFF860FC87F821.text	857E879E8E46962AFFFFF860FC87F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris angustus Nguyen-Phung et Cambefort 1986	<div><p>1.  Copris angustus Nguyen-Phung et Cambefort, 1986, ♂. 2.  Copris angustus Nguyen-Phung et Cambefort, 1986, ♀. 3.  Copris coriarius Gillet, 1907, ♂. 4.  Copris interioris Kolbe, 1897, ♂. 5.  Copris interioris Kolbe, 1897, ♀. 6.  Copris laius laius von Harold, 1868, ♂. 7.  Copris laius laius von Harold, 1868, ♀. 8.  Copris megaceratoides Waterhouse, 1891, ♂. 9.  Copris megaceratoides Waterhouse, 1891, ♀. 10.  Copris moffartsi Gillet, 1907, ♂. 11.  Copris moffartsi Gillet, 1907, ♀. 12.  Copris orion orion Klug, 1835, ♂.</p><p>Planche 75.</p></div>	https://treatment.plazi.org/id/857E879E8E46962AFFFFF860FC87F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E47962DFFFFF860FC87F821.text	857E879E8E47962DFFFFF860FC87F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Copris orphanus Guerin-Meneville 1847	<div><p>1.  Copris orphanus Guérin-Méneville, 1847 . 2.  Copris sinon Olivier, 1789, ♂. 3.  Heliocopris antenor (Olivier, 1789), ♀. 4.  Heliocopris cf dilloni Guérin-Méneville, 1847, ♂. 5.  Heliocopris cf dilloni Guérin-Méneville, 1847, ♀. 6.  Heliocopris myrmidon Kolbe, 1893, ♀. 7.  Pedaria estellae Josso &amp; Prévost, 2006, ♂. 8.  Pedaria humana Cambefort, 1982 . 9.  Heteronitis tridens (de Castelnau, 1840), ♂. 10.  Heteronitis tridens (de Castelnau, 1840), ♀. 11.  Onitis guineensis Gillet, 1918, ♂. 12.  Onitis guineensis Gillet, 1918, ♀.</p><p>Planche 76.</p></div>	https://treatment.plazi.org/id/857E879E8E47962DFFFFF860FC87F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E40962CFFFFF860FC87F821.text	857E879E8E40962CFFFFF860FC87F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Clypeodrepanus strigatus (Janssens 1953)	<div><p>1.  Clypeodrepanus strigatus (Janssens, 1953) . 2.  Euoniticellus nasicornis (Reiche, 1849), ♂. 3.  Euoniticellus triangulatus (von Harold, 1873), ♂. 4.  Ixodina endroedyi (Endrödi, 1976) . 5.  Ixodina szunyoghyi occidentalis (Endrödi, 1976) . 6.  Liatongus militaris (de Castelnau, 1840) . 7.  Tibiodrepanus sulcicollis (de Castelnau, 1840), ♂. 8.  Tibiodrepanus sulcicollis (de Castelnau, 1840), ♀. 9.  Pseudosaproecius portentosus (Felsche, 1907) . 10.  Pseudosaproecius complexe validicornis (Quedenfeldt,1884), ♂. 11  Stiptopodius doriae vonHarold, 1871. 12.  Caccobius (s. l.) cavatus d'Orbigny, 1908, ♂.</p><p>Planche 77.</p></div>	https://treatment.plazi.org/id/857E879E8E40962CFFFFF860FC87F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E41962FFFFFF860FC87F821.text	857E879E8E41962FFFFFF860FC87F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Caccobius ivorensis Cambefort 1984	<div><p>1.  Caccobius (s. l.) ivorensis Cambefort 1984, ♂. 2.  Caccobius (s. l.) ivorensis Cambefort 1984, ♀. 3.  Caccobius (s. l.) signatipennis von Harold, 1867 . 4.  Caccobius (s. l.) villiersi Cambefort, 1975 . 5  Cleptocaccobius balthasarianus (Cambefort, 1971) . 6.  Cleptocaccobius convexifrons (Raffray, 1877), ♂. 7.  Cleptocaccobius dorbignyi Cambefort, 1984 . 8-9.  Proagoderus auratus (Fabricius, 1801), ♂. 10.  Proagoderus rarus Guérin-Méneville, 1829, ♂. 11.  Afrostrandius adspersus (d’Orbigny, 1908) . 12.  Bicornonthophagus cornifrons (Thomson, 1858), ♂.</p><p>Planche 78.</p></div>	https://treatment.plazi.org/id/857E879E8E41962FFFFFF860FC87F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E42962EFFFFF860FC87F821.text	857E879E8E42962EFFFFF860FC87F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Bicornonthophagus decorsei (d'Orbigny 1908)	<div><p>1.  Bicornonthophagus decorsei (d'Orbigny, 1908), ♂. 2.  Bicornonthophagus decorsei (d'Orbigny, 1908), ♀. 3. Bicornonthophagusreticulatus (d'Orbigny, 1902). 4. Digitontophagus eucata Génier, 2017, ♂. 5.  Digitonthophagus falciger Génier, 2017, ♂. 6.  Digitonthophagus fimator Génier, 2017, ♂. 7.  Furconthophagus monforti Josso &amp; Prévost, 2006, ♂. 8.  Furconthophagus monforti Josso &amp; Prévost, 2006, ♀. 9.  Furcontophagus flaviclava (d'Orbigny, 1902), ♂. 10.  Furconthophagus variegatus (Fabricius, 1798), ♂. 11.  Hyalonthophagus nigroviolaceus (d'Orbigny, 1902), ♂. 12.  Hyalonthophagus nigroviolaceus (d'Orbigny, 1902), ♀.</p><p>Planche 79.</p></div>	https://treatment.plazi.org/id/857E879E8E42962EFFFFF860FC87F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E439621FFFFF860FC87F821.text	857E879E8E439621FFFFF860FC87F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Hyalonthophagus pseudoalcyon (d'Orbigny 1913)	<div><p>1-2.  Hyalonthophagus pseudoalcyon (d'Orbigny, 1913), ♂. 3.  Hyalonthophagus pseudoalcyon (d'Orbigny, 1913), ♀. 4.  Parascatonomus (Morettoeus) maculatus (Fabricius, 1801) . 5.  Parascatonomus (Morettoeus) chloroderus (d’Orbigny, 1905) . 6.  Tiaronthophagus curtipilis d'Orbigny, 1905, ♂. 7.  Tiaronthophagus liberianus (van Lansberge, 1883), ♂. 8.  Tiaronthophagus flexicornis (d'Orbigny, 1902), ♂. 9.  Tiaronthophagus viridiaerus (d’Orbigny, 1908), ♂. 10.  Tiaronthophagus viridiaerus (d’Orbigny, 1908), ♂. 11.  Phalops batesi batesi von Harold, 1867, ♂. 12.  Phalops fimbriatus (Klug, 1835), ♂.</p><p>Planche 80.</p></div>	https://treatment.plazi.org/id/857E879E8E439621FFFFF860FC87F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E4C9620FFFFF860FC87F821.text	857E879E8E4C9620FFFFF860FC87F821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Trichonthophagus triacanthus (Laporte 1840)	<div><p>1.  Trichonthophagus triacanthus (Laporte, 1840), ♂. 2.  Trichontophagus gonzalohalffteri Moretto, 2023, ♂. 3.  Onthophagus (s. l.) atridorsis d'Orbigny, 1902 . 4.  Onthophagus (s. l.) brevipennis d'Orbigny, 1902 . 5.  Onthophagus (s. l.) sellatus Klug, 1845, ♂. 6.  Onthophagus (s. l.) ecopas Josso et Prévost, 2006, ♂. 7.  Onthophagus (s. l.) latiggiber d'Orbigny, 1902 . 8.  Onthophagus (s. l.) trinominatus Goidanich, 1926 . 9.  Onthophagus (s. l.) bimarginatus d'Orbigny, 1902 . 10.  Onthophagus (s. l.) mankonoensis Balthasar, 1966 . 11.  Onthophagus (s. l.) obenbergeri Balthasar, 1965 . 12.  Onthophagus (s. l.) circumdatus d'Orbigny, 1913 .</p><p>Planche 81.</p></div>	https://treatment.plazi.org/id/857E879E8E4C9620FFFFF860FC87F821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
857E879E8E4D9622FFFFF860FAA9F798.text	857E879E8E4D9622FFFFF860FAA9F798.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	fr	Onthophagus tripartitus d'Orbigny 1902	<div><p>1.  Onthophagus (s. l.) tripartitus d'Orbigny, 1902 . 2.  Onthophagus (s. l.) lobi Cambefort, 1984, ♂. 3 -4.  Onthophagus (s. l.) lutaticollis d'Orbigny, 1907, ♂. 5.  Onthophagus (s. l.) lutaticollis d'Orbigny, 1907, ♀. 6.  Onthophagus (s. l.) proximus d'Orbigny, 1908, ♂. 7.  Onthophagus (s. l.) proximus d'Orbigny, 1908, ♀. 8.  Onthophagus (s. l.) bandamai Cambefort, 1984, ♂. 9.  Onthophagus (s. l.) bidens (Olivier, 1789), ♂. 10.  Onthophagus (s. l.) bidens (Olivier, 1789), ♀. 11.  Onthophagus (s. l.) loxodontae Cambefort, 1984 . 12.  Onthophagus (s. l.) mucronatus Thomson, 1858, ♀.</p><p>Derniers articles publiés</p><p>Lourenço W. R. &amp; Ythier E., 2025. – A new cave species of  Isometrus Ehrenberg 1828 from Myanmar ( Scorpiones:  Buthidae). Faunitaxys, 13(04): 1 – 6. https://doi.org/10.57800/faunitaxys-13(04)</p><p>Platia G. &amp; Pedroni G., 2025. – Five new species and new records of click beetles from Saudi Arabia with an updated catalogue ( Coleoptera,  Elateridae). Faunitaxys, 13(05): 1 – 12. https://doi.org/10.57800/faunitaxys-13(05)</p><p>Háva J. &amp; Hodeček J., 2025. – Two additional new species of Cryptorhopalum ( Coleoptera:  Dermestidae: Megatominae) from the Dominican Republic. Faunitaxys, 13(06): 1 – 5. https://doi.org/10.57800/faunitaxys-13(06)</p><p>Vitali F., 2025. – The genus  Epicasta Thomson, 1864 and the limits of the morphological systematics in the study of Lamiinae ( Coleoptera Cerambycidae). Faunitaxys, 13(07): 1 – 5. https://doi.org/10.57800/faunitaxys-13(07)</p><p>Ythier E., Sadine S. E., Alioua Y. &amp; Lourenço W. R., 2025. – A new species of  Androctonus Ehrenberg, 1828 from the Tassili n'Ajjer, Algeria ( Scorpiones:  Buthidae). Faunitaxys, 13(08): 1 – 8. https://doi.org/10.57800/faunitaxys-13(08)</p><p>Strümpher W. P. &amp; Stals R., 2025. – A world list of known cave- and bat guano-associated  Trogidae ( Coleoptera: Scarabaeoidea), and further range extension of  Omorgus lindemannae (Petrovitz) . Faunitaxys, 13(09): 1 – 11. https://doi.org/10.57800/faunitaxys-13(09)</p><p>Lourenço W. R. &amp; Velten J., 2025. – Description of a second species for the genus  Serratochaerilobuthus Lourenço, 2024 ( Scorpiones:  Chaerilobuthidae). Faunitaxys, 13(10): 1 – 6. https://doi.org/10.57800/faunitaxys-13(10)</p><p>Jarzabek-Müller A., 2025. – A new  Idolus Desbrochers des Loges, 1875 for the Caucasus region ( Coleoptera,  Elateridae, Pomachiliini). Faunitaxys, 13(11): 1 – 6. https://doi.org/10.57800/faunitaxys-13(11)</p><p>Laguerre M., 2025. – New species of Arctiinae east of the Canal Zone, Panama ( Lepidoptera,  Erebidae, Arctiinae,  Arctiini). Faunitaxys, 13(12): 1 – 17. https://doi.org/10.57800/faunitaxys-13(12)</p><p>Vlasak J., Botero J. P. &amp; Santos-Silva A., 2025. –  Cerambycidae and  Disteniidae ( Coleoptera) from Ecuador: new genus, new species, new records, and correction in species-group names. Faunitaxys, 13(13): 1 – 14. DOI: https://doi.org/10.57800/faunitaxys-13(13)</p><p>Vives E., 2025. – Una nueva especie de  Pseudelasma Breuning, 1968 ( Coleoptera,  Cerambycidae: Lamiinae).  Cerambycidae de Vietnam (Pars 12). Faunitaxys, 13(14): 1 – 2. https://doi.org/10.57800/faunitaxys-13(14)</p><p>Audureau A., 2025. – A propos de quelques  Hemilophini nouveaux ou peu connus ( Coleoptera,  Cerambycidae, Lamiinae). Faunitaxys, 13(14): 1 – 6. https://doi.org/10.57800/faunitaxys-13(15)</p><p>Faunitaxys</p><p>Volume 13, Numéro 16, Mars 2025</p><p>SOMMAIRE</p><p>Première évaluation de la faune des  Scarabaeidae de la République du Bénin ( Coleoptera,  Scarabaeidae).</p><p>Alain Coache, Michael Dierkens, Bernard Rainon, Lionel Delaunay, Jean-François Josso</p><p>&amp; Pascal Deschamps ................................................. 1 – 120</p><p>CONTENTS</p><p>First assessment of the  Scarabaeidae fauna of the Republic of Benin ( Coleoptera,  Scarabaeidae).</p><p>Alain Coache, Michael Dierkens, Bernard Rainon, Lionel Delaunay, Jean-François Josso</p><p>&amp; Pascal Deschamps ................................................. 1 – 120</p><p>Illustration de la couverture:</p><p>Chalconotus sp. et sa "boule" d'excrément (ici excrément humain). Forêt de Houéyogbé, Bénin.</p><p>Crédits:</p><p>Lionel Delaunay, Pascal Deschamps &amp;  Alain Coache.</p><p>Publié par l’Association Française de Cartographie de la Faune et de la Flore (AFCFF)</p></div>	https://treatment.plazi.org/id/857E879E8E4D9622FFFFF860FAA9F798	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Coache, Alain;Dierkens, Michael;Rainon, Bernard;Delaunay, Lionel;Josso, Jean-François;Deschamps, Pascal	Coache, Alain, Dierkens, Michael, Rainon, Bernard, Delaunay, Lionel, Josso, Jean-François, Deschamps, Pascal (2025): Première évaluation de la faune des Scarabaeidae de la République du Bénin (Coleoptera, Scarabaeidae). Faunitaxys 13 (16): 1-120, DOI: 10.57800/faunitaxys-13(16), URL: http://dx.doi.org/10.5281/zenodo.15376970
