taxonID	type	description	language	source
83552453FFF79113640EA590FF55F7BD.taxon	description	Hexakistra Hook. f. (1887): 283, nom. inval., in syn. Thelypetalum Gagnep. (1925 ‘ 1924 ’) 876. — Type: Thelypetalum pierrei Gagnep. (= Leptopus australis (Zoll. & Moritzi) Pojark.).	en	Martin, B. de Manuel, Welzen, P. C. van (2022): Revision of the genera Leptopus and Notoleptopus (Phyllanthaceae) in Malesia. Blumea 67 (1): 20-25, DOI: 10.3767/blumea.2022.67.01.05, URL: https://doi.org/10.3767/blumea.2022.67.01.05
83552453FFF79113640EA590FF55F7BD.taxon	distribution	Distribution — Nine species, ranging from the Caucasus and Iran to the Himalayas, China, S and SE Asia and into Malesia (with a single species) east to the Lesser Sunda Islands and the Moluccas.	en	Martin, B. de Manuel, Welzen, P. C. van (2022): Revision of the genera Leptopus and Notoleptopus (Phyllanthaceae) in Malesia. Blumea 67 (1): 20-25, DOI: 10.3767/blumea.2022.67.01.05, URL: https://doi.org/10.3767/blumea.2022.67.01.05
83552453FFF791116747A788FEE2FC60.taxon	description	Andrachne australis Zoll. & Moritzi var. angustifolia Müll. Arg. (1866) 235. — Lectotype (designated indirectly by Vorontsova & Hoffmann 2009): Cuming 1528 (lecto G-DC *; isolecto K 2 sheets *, L 3 sheets!), Philippines, Luzon, Prov. Batangao.	en	Martin, B. de Manuel, Welzen, P. C. van (2022): Revision of the genera Leptopus and Notoleptopus (Phyllanthaceae) in Malesia. Blumea 67 (1): 20-25, DOI: 10.3767/blumea.2022.67.01.05, URL: https://doi.org/10.3767/blumea.2022.67.01.05
83552453FFF791116747A788FEE2FC60.taxon	description	Leptopus sanjappae Sumathi, Karthig., Jayanthi & Diwakar (2006) 155, f. 1. — Leptopus calcareus (Ridl.) Pojark. var. sanjappae (Sumathi, Karthig., Jayanthi & Diwakar) Chakrab. & N. P. Balakr. (2008) 604. — Type: Sumathi 17362 (holo CAL; iso K *, MCCH, PBL), Andaman Is., North Andaman, Tekkari, Saddle Peak National Park. Subshrubs, 9 – 50 cm high, stem generally branched only at base; flowering branches 1 – 1.5 mm thick, often tinged red, densely hairy when young, glabrescent. Hairs sericeous-hirsute (base upright, upper part bent horizontally), to c. 0.5 (– 0.8) mm long. Stipules triangular to subulate, 0.8 – 2.8 by 0.2 – 0.6 mm, apex acute, densely hairy. Leaves: petiole 0.3 – 3.4 cm long, hairy; blade elliptic to somewhat obovate, 1 – 8 (– 10) by 0.5 – 4 cm; (1 –) 2 – 3.5 times longer than wide, papery, symmetric, drying brownish or greyish green, base cuneate to attenuate, margin flat, apex rounded and mucronate to shortly acuminate, upper surface glabrous to (completely) hairy on midrib and margin, dark green when fresh, lower surface subglabrous to hairy on venation, light green when fresh, margin subglabrous to hairy; nerves 4 – 6, flat above, slightly raised below, distinct. Flowers green. Staminate flowers 2 – 3.8 mm diam; pedicel terete, 2.7 – 7 mm long, slightly hairy, green to maroonish; se- pals 5, obovate, 0.8 – 2.3 by 0.7 – 1 mm, hairy outside, glabrous inside, green, veined when dry; petals 5, oblong to obovate, 0.5 – 1.2 by 0.2 – 0.3 mm, glabrous, whitish; disc of 5 bilobed glands (to seemingly 10 separate glands), strap-like, 0.7 – 1.2 by 0.1 – 0.2 mm, glabrous, whitish; stamens 5, glabrous, filaments 0.5 – 1.3 mm long, whitish, anthers c. 0.2 – 0.3 mm long, light yellow, 2 - thecate, each theca bilobed; pistillode of 3 free lobes, to 0.8 mm long, glabrous, top slightly split. Pistillate flowers 5 – 6 mm diam, disc-shaped; pedicel terete, 1.2 – 5 (– 9 in fruit) mm long, pilose, green; sepals 5, ovate to obovate, 1.5 – 3.2 (– 5.5 in fruit) by 1.5 – 2 (– 3.5 in fruit) mm, margin and outside pilose, shortly hirsute inside, green, clearly veined when dry; petals 5, minute and sticking to back of disk lobes, oblong to obovate, 0.1 – 0.4 by 0.1 – 0.2 mm, glabrous, single vein generally visible; disc lobes 5, almost completely split and V-shaped, 0.6 – 0.8 mm long, apically often somewhat erose; ovary 0.5 – 0.6 by 0.7 – 0.9 mm, densely pilose, yellowish to light greenish; style c. 0.2 mm long, white to light greenish, pilose; stigmas 0.7 – 0.8 mm long, glabrous except sometimes for some hairs on clavate lobe apices, white to greenish. Fruits triangular in transverse section, c. 5 by 2.5 mm, yellow-green to white when ripe, shortly pilose, reticulately veined, white to light greenish when young; columella 1 – 1.5 mm long. Seeds 1.2 – 2 by 1 – 1.5 by 1 – 1.5 mm, sides ribbed, back convex and smooth, brown when dry. Distribution — China (Hainan), Thailand, Andaman Islands, Vietnam; Malesia: Malay Peninsula, Sumatra, Java (including Madura), Borneo (Sabah), Philippines, Lesser Sunda Islands (Bali, Timor), Moluccas (Tanimbar Islands, Wetar). Habitat & Ecology — Locally common in primary to disturbed tropical to evergreen forest, deciduous forest, dry Eucalypt savannah; in the shade to open spots, often on steep slopes, often between rocks or even lithophytic; soil: in shallow or deep soils on limestone or loam. Altitude: 15 – 1100 m. Presumably flowering and fruiting at least twice during the whole year except for January. According to Vorontsova & Hoffmann (2009) the IUCN Red List category is Least Concern (LC), as the extent of occurrence far exceeds 20 000 km 2. Note — The young branchlets are often very hairy, but they are glabrescent. The difference with Notoleptopus decaisnei is then mainly in the subglabrous older branchlets, which are hairy in N. decaisnei.	en	Martin, B. de Manuel, Welzen, P. C. van (2022): Revision of the genera Leptopus and Notoleptopus (Phyllanthaceae) in Malesia. Blumea 67 (1): 20-25, DOI: 10.3767/blumea.2022.67.01.05, URL: https://doi.org/10.3767/blumea.2022.67.01.05
83552453FFF59111640EA22BFD89F7CE.taxon	description	— Type: Notoleptopus decaisnei (Benth.) Voronts. & Petra Hoffm. Leptopus auct. non Decne.: Airy Shaw (1980 a) 122, (1980 b) 645. Small shrubs, erect, monoecious. Indumentum of simple hairs, most parts hirsute. Stipules small, late caducous. Leaves alternate, distichous, simple; petiole reniform in transverse section, not pulvinate; blade chartaceous, eglandular, margin entire, venation pinnate, nerves looped and closed near margin, veins and veinlets reticulate, indistinct. Inflorescences axillary fascicles, with a single to few flowers of both sexes; bracts pres- ent, small, similar to stipules. Flowers 5 - merous, actinomorphic; pedicels likely articulate above base, but impossible to establish correctly; sepals 5, herbaceous; petals 5; disc outside stamens. Staminate flowers: pedicel slender; petals slightly shorter than sepals; disc glands 5, bilobed for less than 1 / 3, lobes rounded; stamens 5, free, filaments filiform, anthers opening latrorse with lengthwise slits; pistillode of 3 free segments, apices slightly bifid. Pistillate flowers: petiole sturdier than in staminate flowers, apically somewhat widening; sepals slightly accrescent in fruit; petals much shorter than sepals; disc annular, regularly and deeply divided into 5 emarginate or erose segments; ovary 3 - locular, superior, densely pilose; ovules 2 per locule; style indistinct, stigmas bifid to base, apically rounded. Fruits 3 - lobed, capsular; columella slender, persistent, apically not broadened, puberulent. Seeds naked, triangular to reniform in transverse section, abaxial side convex, all sides ribbed. Distribution — A monotypic genus, Malesia: Java, Lesser Sunda Islands, New Guinea; and Australia.	en	Martin, B. de Manuel, Welzen, P. C. van (2022): Revision of the genera Leptopus and Notoleptopus (Phyllanthaceae) in Malesia. Blumea 67 (1): 20-25, DOI: 10.3767/blumea.2022.67.01.05, URL: https://doi.org/10.3767/blumea.2022.67.01.05
83552453FFF591176747A789FF4FFEF4.taxon	description	Andrachne fruticosa Decne. ex Müll. Arg. (1866) 235, nom. illeg., non L. — Arachne fruticosa (Decne. ex Müll. Arg.) Hurus. (1954) 339. — Syntypes: Anonymous s. n. (ex Herb. P) (G-DC *, 2 sheets), Indonesia, Timor; F. von Mueller s. n. (G-DC *), New Holland (Australia), Victoria River; Zollinger 2795 (not seen), Indonesia, Java. Subshrubs, 10 ‒ 50 cm high; internodes 0.5 ‒ 2 cm long; flower- ing branchlets 0.05 ‒ 0.3 mm diam, hairy, likely not glabrescent. Indumentum hirsute hairs of c. 0.8 – 1 mm long. Stipules triangular, 1 ‒ 2.5 by 0.25 ‒ 0.5 mm, membranous, apex attenuate, sometimes slightly acute, hairy. Leaves: petiole 0.25 ‒ 3 mm long, pubescent to densely pubescent; blade elliptic to obovate, 2.5 ‒ 50 by 2 ‒ 33 mm, 1 ‒ 2 times longer than wide, symmetric, base cuneate to slightly attenuate, margin flat to slightly sinuate, densely ciliate, apex attenuate to acute, adaxial surface puberulous to pubescent, abaxial surface densely pubescent, venation of 3 ‒ 5 lateral veins per side, midrib very prominent, flat on adaxial surface and elevated abaxially. Flowers: sepals ovate, with patent simple hairs, ciliate, apex acute to slightly rounded, midrib prominent; disc glabrous. Staminate flowers 0.4 ‒ 0.8 mm diam in bud, 0.9 ‒ 1.5 mm diam when open, coro- nate, light yellowish to reddish brown; pedicel terete, slightly thickened apically and basally, 0.8 ‒ 1.7 by 0.03 ‒ 0.12 mm, hairy; sepals ovate to elliptic, 0.6 ‒ 1 by 0.25 ‒ 0.5 mm; petals elliptic to obovate, 0.75 ‒ 1.1 by 0.1 ‒ 0.2 mm, glabrous to slightly pubescent, margin entire, apex acute to attenuate, midrib prominent; disc glands in Malesian material small and indistinct, bilobed; stamens: filaments 0.25 ‒ 0.5 mm long, glabrous, an- thers 0.15 ‒ 0.22 by 0.2 ‒ 0.25 mm, yellowish brown; pistillode 0.1 ‒ 0.2 mm long, subsessile. Pistillate flowers 1.5 ‒ 3 mm diam in bud, 2.5 ‒ 3.5 mm when open, disc-like, puberulous; pedicel terete, thickened especially apically, 1.5 ‒ 4.5 (– 6.5 in fruit) mm long, pubescent to densely pubescent; sepals ovate, 2.2 ‒ 3 (– 3.4 in fruit) by 1.2 ‒ 1.9 (– 2.1 in fruit) mm; petals elliptic to ligulate to obovate, 0.35 ‒ 0.55 by 0.18 ‒ 0.3 mm, slightly pilose, margin en- tire, apex narrowly rounded, distinctly veined; disc lobes strap-like, c. 0.5 by 0.3 mm; ovary globose to slightly oblate, 0.5 ‒ 1 by 0.75 ‒ 1.3 mm, densely pubescent; stigmas 0.55 ‒ 0.8 mm long. Fruit triangular in transverse section, 1.5 ‒ 3.5 by 2.7 ‒ 4.5 mm, densely pubescent, reticulately veined; columella 1.2 ‒ 1.7 mm long. Seeds 1.35 ‒ 1.8 by 1 ‒ 1.3 by 0.75 ‒ 1 mm, black when dry. Distribution — Malesia: Java (Madura), Lesser Sunda Islands (Roti, Sumba, Timor), Papua New Guinea (Central Province); Australia (Queensland). Habitat & Ecology — Scarce to locally fairly common in usu- ally (more) open places in rocky surroundings between grass tussocks and in open Eucalypt woodland. The plants appear to be much cropped by wallabies (Pullen 9616). Altitude: 1 ‒ 500 m. Flowering and fruiting: February to May. Note — A geocline may be present as staminate disc lobes and pistillode seem to be smaller on Madura (near Java) and much larger in Australia. However, the material was too scarce to be certain. 24 Blumea – Volume 67 / 1, 2022 Acknowledgements We thank Jan van Os (†) and Esmée Winkel for the two beautiful drawings and the director of Naturalis Biodiversity Center for facilities and use of the collections. The other herbaria mentioned are thanked for providing high resolution images of the types. The last author thanks the Treub Maatschappij (the Society for the Advancement of Research in the Tropics) for their support of the Ornstein chair in Tropical Plant Biogeogra- phy. Two reviewers are thanked for their helpful suggestions that improved this manuscript.	en	Martin, B. de Manuel, Welzen, P. C. van (2022): Revision of the genera Leptopus and Notoleptopus (Phyllanthaceae) in Malesia. Blumea 67 (1): 20-25, DOI: 10.3767/blumea.2022.67.01.05, URL: https://doi.org/10.3767/blumea.2022.67.01.05
