taxonID	type	description	language	source
0EF552F542975BBDA9EB77CCD2E97ED6.taxon	description	Figs 25 – 31	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
0EF552F542975BBDA9EB77CCD2E97ED6.taxon	description	Description. Female. Body length 1.17 – 1.9 mm; fore wing length 1.3 – 1.5 mm. Colour. Head and mesosoma blue green with metallic diffuse coppery lustr; antenna with scape yellowi-brown; pedicel, anelli, F 1 – F 6 and clava brown. Fore and hind coxae dark blue with metallic violet lustre, mid coxa brown; all femora dark brown, all tibiae brown, all tarsi yellow-brown. Fore wing hyaline, venation brown. Metasoma dorsally dark brown with metallic green and coppery lustre; ovipositor sheath brown. Sculpture. Head, clypeus and mesosoma reticulate; propodeum alutaceous; petiole smooth; metasoma smooth and shiny. Head. Head in dorsal view 2 – 2.06 × as broad as long and 1.26 – 1.3 × as broad as mesoscutum; in frontal view 1.2 – 1.3 × as broad as high. POL 2 – 2.2 × as long as OOL. Eye height 1.08 – 1.11 × eye length and 1.49 – 1.57 × as long as malar space. Distance between antennal toruli and lower margin of clypeus 1.18 – 1.25 × distance between antennal toruli and median ocellus. Antenna with scape 0.92 – 0.93 × as long as eye height and 1 – 1.03 × as long as eye length; pedicel 1.9 – 2.0 × as long as broad; combined length of pedicel and flagellum 1.09 – 1.1 × breadth of head; F 1 – F 6 longer than broad, F 1 1.4 – 1.6 × as long as broad and with 1 row of sensilla; clava 2.16 – 2.2 × as long as broad, with small micropilose area on each C 3 and C 4. Lower margin of clypeus bidentate. Mesosoma. Mesosoma 1.46 – 1.52 × as long as broad. Scutellum moderately arched, as long as broad, frenal area not distinct by sculpture. Propodeum 0.38 – 0.42 × as long as scutellum, without costula, median carina and nucha, with irregular transverse carina. Fore wing 2.3 – 2.5 × as long as its maximum width; basal cell with 0 – 3 setae; basal vein pilose; speculum open below; M 0.85 – 0.93 × as long as PM and 1.82 – 1.92 × as long as S; stigma small. Metasoma. Metasoma 3.1 – 3.15 × as long as broad, 1.68 – 1.73 × as long as mesosoma, 1.18 – 1.22 × as long as mesosoma and head combined. Petiole transverse. Ovipositor sheath projecting slightly beyond apex of metasoma. Male. Body length 1.6 – 1.7 mm; fore wing length 1.2 – 1.4 mm. Distance between antennal toruli and lower margin of clypeus 1.3 – 1.55 × distance between antennal toruli and median ocellus. Combined length of pedicel and flagellum 1.26 – 1.29 × breadth of head; clava 2.9 – 3.2 × as long as broad. Metasoma 2.22 – 2.33 × as long as broad, 1.11 – 1.19 × as long as mesosoma, 0.8 – 0.84 × as long as mesosoma and head. Otherwise similar to female.	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
0EF552F542975BBDA9EB77CCD2E97ED6.taxon	etymology	Etymology. From the Latin « minutus » (= small), referring to the small size of body of this species.	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
0EF552F542975BBDA9EB77CCD2E97ED6.taxon	distribution	Distribution. South Korea.	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
DE14747AF1CD5D118BBF1767AE760635.taxon	description	Figs 6 – 16	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
DE14747AF1CD5D118BBF1767AE760635.taxon	description	Description. Female. Body length 1.63 – 1.88 mm; fore wing length 1.26 – 1.31 mm. Coloration. Head with areas around ocellus and occiput yellowish brown. Antenna with scape and pedicel reddish yellow, flagellum dark brown. Posterior margin of pronotum, median part of scutellum, dorsellum, median part of propodeum and metasoma from brownish-black to fuscous. All legs yellow. Tegulae brown; wings hyaline, venation pale yellow. Sculpture. Mesoscutum moderately shiny, with delicate superficial reticulation; propodeum shiny with smooth engraved reticulation; metasoma with Mt 2 - Mt 4 smooth, Mt 5 - Mt 6 alutaceous. Head. Head in dorsal view 2.42 × as broad as long and 1.05 – 1.13 × as broad as mesosoma, in frontal view 1.15 × as broad as height. POL 1.67 × OOL; OOL 2 × OD. Face medially concave. Malar space 0.58 – 0.6 × as long as eye height; malar sulcus moderately curved. Mouth 1.42 – 1.45 × as wide as height of malar space. Antenna with scape 0.8 × as long as eye height; 3.56 × as long as broad and reaching of median ocellus; combined length of pedicel and flagellum 1.34 – 1.41 × breadth of head; 1.5 – 1.53 × breadth of mesoscutum; pedicel 1.75 × as long as broad and 0.57 × as long as F 1; funicle proximally not stouter than pedicel; F 1 3 – 3.25 ×, F 2 2.22 – 2.4 ×, F 3 1.67 – 2 × as long as broad. Clava slightly broader than F 3; 0.76 – 0.78 × as long as F 2 – F 3 and 2.37 × as long as broad; articulation of claval segments not visible. Mesosoma. Mesosoma in dorsal view 1.53 × as long as broad. Pronotum medially 0.3 – 0.42 × as long as mesoscutum. Mid lobe of mesoscutum 1.07 × as broad as long; median line fine but traceable in some lights, with 4 adnotaular setae on each side. Scutellum 0.62 × as long as mid lobe of mesoscutum, 1.32 × as broad as long; distance between SMG equal distance SMG to SLG, enclosing space 2.29 × as long as broad; have the anterior pair of setae clearly visible behind the middle of scutellum; distance between anterior pairof setae and front edge of scutellum 2 × distance between anterior and posterior pair of setae. Dorsellum 0.37 × as long as scutellum and 2 × as broad as long. Propodeum medially as long as dorsellum; median carina distinct; callus with 4 setae. Legs slender; hind femur 3.13 × as long as broad; spur of mid tibia as long as basitarsus. Fore wing 1.96 – 2.1 × as long as broad; costal cell 0.71 × length of M, 12.5 × as long as broad; SM with 4 dorsal setae; M 3.5 × length of S, its front edge with 13 setae; ST thin proximally, expanding gradually forming rather indistinct stigma; PM rudimentary; speculum small, not extending below M; cilia 0.3 × length of S. Hind wing obtuse; cilia 0.26 × breadth of wing. Metasoma. Metasoma 1.33 – 1.5 × as long as mesosoma; 1.09 – 1.18 × as long as head and mesosoma combined. Gaster longitudinal-oval, 1.96 – 2.25 × as long as broad, acuminate; its last tergite as long as broad; ovipositor sheath projecting by 0.1 length of Mt 8; cerci with one pair of setae 2 × as long as second pair, slightly curved; tip of hypopygium at 0.51 – 0.6 length of gaster. Male. Body length 1.53 mm; fore wing length 1.26 mm. Mid lobe of mesoscutum 1.04 × as long as broad; with 3 adnotaular setae on each side. Antenna with scape 2.14 × as long as broad, with ventral plaque 0.53 length of scape; combined length pedicel and flagellum 2.19 × breadth of mesoscutum; pedicel 1.56 × as long as broad, and as long as F 1; F 1 quadrate, F 2 2.2 × as long as broad, F 3 and F 4 2.6 × as long as broad; clava 1.07 × as long as F 3 – F 4, 7 × as long as broad, with C 1 and C 2 subequal and 2.5 × as long as broad; C 3 slightly shorter; whorled setae long. Metasoma as long as mesosoma, 0.78 × as long as head and mesosoma combined; gaster 1.89 × as long as broad. Otherwise, similar to female.	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
DE14747AF1CD5D118BBF1767AE760635.taxon	etymology	Etymology. The species is named in honour of the Russian hymenopterist, expert of Pteromalidae, and main collector of the types, Dr Ekaterina V. Tselikh (ZISP).	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
DE14747AF1CD5D118BBF1767AE760635.taxon	distribution	Distribution. Republic of Korea.	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
F2D49C1D970757DE8D8A484DC1211CB8.taxon	description	Fig. 19	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
F2D49C1D970757DE8D8A484DC1211CB8.taxon	distribution	Distribution. Moldova, Poland (Bouček 1965; UCD Community 2025), Japan, Republic of Korea (Kamijo 1979), Russia (Kosheleva 2023).	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
15AA234D880E5324BF41F05821203303.taxon	description	Fig. 18	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
15AA234D880E5324BF41F05821203303.taxon	distribution	Distribution. Argentina, Australia, Belgium, Bulgaria, Canada, Chile, China, Croatia, Czech Republic, Egypt, England, France, Georgia, Germany, Hungary, India, Iran, Israel, Italy, Japan, Kazakhstan, Moldova, Netherlands, New Zealand, Poland, Republic of Korea (new record), Romania, Russia, Serbia, Slovakia, Spain, Sweden, Tajikistan, Tunisia, Turkey, Ukraine, United Kingdom, USA, Uzbekistan (Tselikh 2010, 2019; UCD Community 2025).	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
CB3E34275C8B50D6B38C73717CAAE235.taxon	description	Fig. 20	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
CB3E34275C8B50D6B38C73717CAAE235.taxon	distribution	Distribution. Australia, Austria, Belgium, Bosnia and Herzegovina, Bulgaria, Canada, China, Croatia, Czech Republic, Finland, France, Georgia, Germany, Greece, Guatemala, Hungary, Israel, Italy, Japan, Kazakhstan, Lithuania, Mexico, Netherlands, Norway, Poland, South Korea (new record), Romania, Russia, Spain, Sweden, Switzerland, United Kingdom, USA (Tselikh 2010, 2019; UCD Community 2025).	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
573CEAA6165653F993FCCC8B9032261F.taxon	description	Figs 32 – 39	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
573CEAA6165653F993FCCC8B9032261F.taxon	distribution	Distribution. Czech Republic, Republic of Korea (new record), Sweden, (Hedqvist 1974; Bouček and Rasplus 1991).	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
D01CD61798ED5242A8507CE3E2E27424.taxon	diagnosis	Diagnosis. Head without occipital carina (Fig. 35). Gena without malar depression; genal lamina absent (Fig. 34). Clypeal margin slightly produced and subtruncate (Fig. 37). Antennal formula 11264 (Fig. 32); flagellum short and slightly clavate; area of micropilosity small and extending C 3 – C 4 (Fig. 32). Antenna inserted above lower ocular line; antennal protuberance small; scrobes shallow (Fig. 33). Pronotum short with collar not carinate (Fig. 39); notauli not complete (Fig. 39); prepectus longer than tegula. Mesoscutum and scutellum flat (Fig. 34). Propodeum reticulate with median carina and plicae; costula and nucha absent; propodeal spiracle inserted close to anterior propodeal margin (Fig. 39). Fore wing hyaline, with closed speculum; M not widened and longer than S and P (Fig. 36). Hind coxa dorsally bare. Petiole in dorsal view smooth and transverse (Fig. 39). Cerci with setae subequal in length, ovipositor shortly protruding.	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
6A7759624ED056AFBD8953024CC31222.taxon	description	Fig. 17	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
6A7759624ED056AFBD8953024CC31222.taxon	distribution	Distribution. Taiwan, Japan, South Korea; Russia (south of Far East) (Belokobylskij et al. 2024).	en	Tselikh, Ekaterina V., Kosheleva, Oksana V., Ku, Deok-Seo, Lee, Jaehyeon, Belokobylskij, Sergey A. (2025): The hymenopteran parasitoid complex (Hymenoptera, Braconidae, Eulophidae, Pteromalidae) of the pine bark beetle Cryphalus fulvus Niisima, 1908 (Curculionidae, Scolytinae) in South Korea. Journal of Hymenoptera Research 98: 637-652, DOI: 10.3897/jhr.98.155465
