taxonID	type	description	language	source
6C303602FFA70959FF69F8846CB6DE01.taxon	diagnosis	Diagnosis. Antennomere 3 of males strongly modified, with massive subtriangular dorsal projection; antennae gradually thickened toward apices; frons of males in some species modified; aedeagus of simonis form, with approximately drop-shaped capsular region of median lobe, setae on subapical area of dorsal wall, and generally subtriangular distal plate. Species of this group are large-bodied, and morphologically similar to those of the C. impressifrons and C. taurus groups. The modification of antennomere 3 in males is a unique synapomorphy. This group includes seven species described previously:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFA40955FF69FB4D6C5ADE25.taxon	description	(Figs 1 – 11)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFA40955FF69FB4D6C5ADE25.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA [22] - Yunnan, / SE Pingbian, primary / forest, 22 º 54 ’ 31 ’’ N, / 103 º 41 ’ 44 ’’ E, 2100 m / 27. VIII. 2014, V. Assing ” [white, printed]; “ CEPHENNODES (s. str.) / triangulifer m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW). Paratypes (15 exx.): 2 ♀♀, same data as for holotype; 4 ♂♂, 2 ♀♀, “ CHINA: Yunnan, SE Pingbian, / 22 º 54 ’ 31 ’’ N, 103 º 41 ’ 44 ’’ E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 28. VIII. 2014, / leg. M. Schülke [CH 14 - 22 a] ” [white, printed]; 3 ♂♂, 4 ♀♀, “ CHINA: Yunnan, SE Pingbian, / 22 º 54 ’ 31 ’’ N, 103 º 41 ’ 44 ’’ E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 27. VIII. 2014, / leg. M. Schülke [CH 14 - 22] ” [white, printed]; each paratype with yellow “ PARATYPUS ” label (MNB, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFA40955FF69FB4D6C5ADE25.taxon	diagnosis	Diagnosis. Body very large, BL> 1.9 mm; antennae gradually thickening; punctures on frontovertexal region of head and on pronotum fine but still more distinct than those on elytra; pronotal lateral carinae single; subhumeral lines short, as long as 0.35 × EL. Male: antennomere 3 (Figs 3, 7) with mesal subtriangular projection only slightly longer than length of antennomere, with pointed apex and anterior margin indistinctly sinuate; frontal region of head (Fig. 2) anterior to supraantennal tubercles with pair of small, shallow and diffuse lateral impressions; aedeagus (Figs 4 – 5, 8 – 11) in ventral view with apex of capsular region subtriangular and blunt, distal plate elongate, subtriangular and strongly curved dorsad.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFA40955FF69FB4D6C5ADE25.taxon	description	Description. Body of male (Figs 1, 6) strongly convex, oval, with weakly marked constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 2.01 – 2.13 mm. Head (Figs 2, 7) broadest across small but strongly convex and coarsely faceted eyes, HL 0.29 – 0.35 mm, HW 0.50 – 0.53 mm; frontovertexal region weakly convex at middle, anterior region of frons with pair of small and shallow lateral impressions; supraantennal tubercles weakly elevated; anterior margin of clypeus slightly thickened. Punctures on frontovertexal area fine but sharply marked and separated by spaces 2 – 3 × as wide as their diameters, clypeus with sparser punctures at middle, frontovertexal region with barely discernible median longitudinal line composed of minute punctures; setae on frons and vertex sparse, long and suberect. Antennae (Figs 1, 3) long and moderately slender, gradually thickened, AnL 1.05 – 1.13 mm, antennomere 1 and 2 each about as long as broad, 3 modified, with large mesal triangular projection slightly longer than length of antennomere, 4 – 6 each about as long as broad, 7 indistinctly elongate, 8 and 9 each weakly transverse, 10 about as long as broad, 11 about as long as 9 and 10 combined, twice as long as broad. Pronotum in strictly dorsal view transversely subrectangular (in Figs 1 and 6 pronotum is slightly tilted downwards anteriorly), strongly convex at middle and flattened near anterior and hind corners, broadest near middle; PL 0.63 – 0.68 mm, PW 0.90 – 1.00 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded and distinctly microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc fine but distinct, with sharp margins, separated by spaces subequal to diameters of punctures; punctures becoming smaller towards all pronotal margins, except for those in anterior corners, which are much denser and coarse. Setae on pronotal disc long and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior third; EL 1.08 – 1.25 mm, EW 0.95 – 1.00 mm, EI 1.10 – 1.25; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.35 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located slightly closer to subhumeral line than to lateral margin of mesoscutellum; elytral apices unmodified, separately rounded. Punctures on elytra fine, unremarkable; setae distinctly longer than those on pronotum, dense and erect. Hind wings functional. Metaventrite with diffuse lateral impressions as long as 1 / 3 of its length. Legs long and slender, unmodified. Aedeagus (Figs 4 – 5, 8 – 11) of simonis form, AeL 0.33 mm; capsular region of median lobe in ventral view approximately drop-shaped, stout, with subtriangular and blunt apex shifted to left side; dorsal apical projection (= distal plate) in ventral view slender, elongate and tapering distad, in lateral view hook-shaped, curved dorsad at a nearly right angle; parameres long and slender, not reaching apex of aedeagus, each with one apical and two subapical long setae. Female. Externally similar to male, except for antennomere 3 and frontal region of head unmodified, and relatively slightly shorter antennae; wingless; BL 1.93 – 2.10 mm; HL 0.30 – 0.35 mm, HW 0.50 – 0.55 mm, AnL 1.03 – 1.05 mm; PL 0.58 – 0.70 mm, PW 0.95 – 1.03 mm; EL 1.00 – 1.13 mm, EW 0.93 – 1.03 mm, EI 1.08 – 1.13.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFA40955FF69FB4D6C5ADE25.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFA40955FF69FB4D6C5ADE25.taxon	etymology	Etymology. The adjective triangulifer refers to the modification of the antennomere 3 in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFA40955FF69FB4D6C5ADE25.taxon	discussion	Remarks. Species of the C. bos group are difficult to identify, their aedeagi are poorly differentiated, so external structures are important to diagnose them, especially head modifications in males. Males of Cephennodes triangulifer differ from all remaining Chinese species of this complex (C. aschnae is excluded from the discussion) in the following combination of characters: the frontal region of head with shallow and indistinct lateral impressions without elevated anterior margins and lacking a median impunctate area; and the apex of projection on antennomere 3 not abruptly narrowed to needle-sharp point. The remaining (i. e., non-Chinese) species included in this group, the Vietnamese C. bos, C. bubalus, and C. buceros, have the subtriangular projection of the antennomere 3 much longer than the length of this antennomere, and its mesal margin strongly sinuate, clearly differing in shape from the short projection of C. triangulifer. Among the Chinese species, C. pseudobos and C. impressibos also have the projections and antennomeres 3 of a different shape, and only C. brachyspinosus, as its name implies, has the spinelike projection similarly short as that of C. triangulifer. The aedeagi of these two species are highly similar, but they differ in the shape of the distal plate in ventral view: it is narrowing distad in C. triangulifer, but parallel-sided in C. brachyspinosus. A clear external feature to distinguish these two species is the frontal area of the head in males: in C. triangulifer the very faint and barely discernible median longitudinal group of densely set minute punctures is surrounded by large punctures distributed all over the frontovertexal region, whereas in C. brachyspinosus this median line is surrounded by a large oval impunctate area. Cephennodes brachyspinosus also has a more strongly punctate pronotum. Within the studied sample of specimens identified as C. triangulifer, variability in the shape of the median lobe and distal projections was found, illustrated in Figs 4 – 5, 8 – 11. All specimens come from the same collecting spot, and externally they do not differ in any features, also the variability in the body pigmentation (dark brown shown in Fig. 1 and slightly lighter, as in Fig. 6) is continuous. The impressions on the frontal region are difficult to study, and if the head is observed in posterodorsal view, as illustrated in Fig. 7, it is possible to overlook these small structures.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFA80955FF69FE466C76DB51.taxon	diagnosis	Diagnosis. Aedeagus of modified latus form, with generally oval large and broad median lobe, its membranous area relatively large, slant and exposed (i. e., not concealed by ventral wall of median lobe), apical projections subtriangular in shape, and (most importantly) endophallic structures include two long strongly elongate elements: anchoring device with bifurcate apex, and thick modified flagellum (in a fully erected condition flagellum projecting distad, and bifurcate anchoring device directed proximad). Moreover, in some species apices of parameres are variously modified, in some cases bearing a tooth-like projection, and the pigmentation in many cases is darker on the head and pronotum, and distinctly lighter on the elytra. In many species males have odd (sometimes multiple) modifications of the elytra (lateral impressions covered with short setae, apical impressions or cavities, setal tufts), antennomeres, legs (tibiae and / or tarsi) or the last exposed abdominal sternite. The excavatus group is similar to the C. longipes group, the latter also occurring in China, but no new species have been found in the material from Yunnan studied for the present publication. These groups can in future be merged, based on a highly similar structure of the aedeagus. This group is particularly species-rich and currently includes 25 described species distributed in China, Nepal, and Bhutan:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAE0951FF69FC4F6854DC1D.taxon	description	(Figs 12 – 17)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAE0951FF69FC4F6854DC1D.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan [CH 07 - 22], / Nujiang Lisu Aut. Pref., / Gaoligong Shan, valley 21 km W / Gongshan, 27 º 47 ’ 03 ’’ N, / 98 º 27 ’ 39 ’’ E, 3320 m, moss / alder, bamboo, Rhodod., sifted, / 6. VI. 2007, leg. A. Pütz ” [white, printed]; “ CEPHENNODES (s. str.) / curvitibia m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (cAP).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAE0951FF69FC4F6854DC1D.taxon	diagnosis	Diagnosis. Body small, around 1 mm in length; antennae with trimerous clubs; punctures on frontovertexal region of head stronger than on pronotum and elytra; pronotal lateral carinae single; subhumeral lines short, as long as 0.30 × EL. Male: metatibiae distinctly curved (Fig. 12), and elytral apices modified (Fig. 13): each apex with small asetose oval impression, and subapical region of elytral suture with row of setae directed posterolaterad; aedeagus (Figs 14 – 15) in ventral view with apical projections about as long as broad, in lateral view distal (dorsal) plate weakly curved, median lobe in ventral view with short and rounded lobe at each distolateral corner of capsular region; parameral apices strongly modified, in lateral view (Fig. 17) apex broadened and with elongate lateral subapical process, distal region microdentate.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAE0951FF69FC4F6854DC1D.taxon	description	Description. Body of male (Fig. 12) strongly convex, oval, with shallow but distinct constriction between pronotum and elytra, dark chestnut brown with slightly lighter tarsi, covered with light brown setae. BL 1.11 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.15 mm, HW 0.25 mm; frontovertexal region weakly convex at middle, unmodified; supraantennal tubercles weakly elevated. Punctures on frontovertexal area distinct, sharply marked and separated by spaces equal to or slightly narrower than their diameters; setae sparse, short and suberect. Antennae moderately long and moderately slender, with trimerous clubs, AnL 0.55 mm, antennomere 1 and 2 each distinctly elongate, 3 indistinctly elongate, 4 about as long as broad, 5 indistinctly elongate, 6 and 7 each about as long as broad, 8 distinctly transverse, 9 and 10 each weakly transverse, 11 about as long as 9 and 10 combined, nearly twice as long as broad. Pronotum semi-oval, strongly convex at middle and weakly flattened near hind corners, broadest near middle; PL 0.34 mm, PW 0.48 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded and finely, barely noticeably microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equally distant from posterior and lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, weakly suberect. Elytra broader than pronotum, together oval, broadest slightly anterior to middle; EL 0.63 mm, EW 0.55 mm, EI 1.14; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.30 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices (Fig. 13) modified, separately rounded, each with small asetose oval impression, and subapical region of elytral suture with row of setae directed posterolaterad. Punctures on elytra fine, unremarkable; setae similar to those on pronotum. Hind wings functional. Metaventrite lacking lateral impressions. Legs long and slender, modified: metatibiae strongly curved mesad. Aedeagus (Figs 14 – 17) of modified latus form, AeL 0.25 mm; capsular region of median lobe in ventral view oval, with pair of short rounded distolateral lobes; apical projections subtriangular and about as long as broad, in lateral view distal (dorsal) plate weakly curved; parameres short and massif, each with strongly modified apex (Figs 16 – 17), bearing elongate lateral projection and with microdentate distal process, each paramere with two similarly long subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAE0951FF69FC4F6854DC1D.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAE0951FF69FC4F6854DC1D.taxon	etymology	Etymology. The noun in apposition curvitibia refers to the curved metatibiae in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAE0951FF69FC4F6854DC1D.taxon	discussion	Remarks. This is a remarkable species that can be easily identified based on the small body size, punctures on the head dorsum stronger than those on the pronotum and elytra, unmodified antennae, modified metatibiae and modified elytral apices; shapes of highly modified parameral apices are also unique for C. curvitibia.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAC0950FF69FC5E69A7DCE5.taxon	description	(Figs 18 – 24)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAC0950FF69FC5E69A7DCE5.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan [CH 07 - 24] Nujiang / Lisu Aut. Pref., Gaoligong Shan, valley 18 / km W Gongshan, 3020 m, 27 º 47 ’ 54 ’’ N, / 98 º 30 ’ 13 ’’ E, mixed forest, litter, moss / wood sifted, 7. VI. 2007, M. Schülke ” [white, printed]; “ CEPHENNODES (s. str.) / rufus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (3 exx.): 3 ♀♀, same data as for holotype, each with yellow “ PARATYPUS ” label (MNB, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAC0950FF69FC5E69A7DCE5.taxon	diagnosis	Diagnosis. Body moderately large, around 1.5 mm in length; antennae with pentamerous clubs; punctures on head, pronotum and elytra equally fine and inconspicuous; pronotal lateral carinae single; subhumeral lines short, as long as 0.30 × EL. Male lacking secondary sexual characters; aedeagus (Figs 23 – 24) in ventral view with apical projections about as long as broad, in lateral view distal (dorsal) plate weakly curved, in ventral view rounded lobe at each distolateral corner of capsular region lacking; parameral apices strongly modified, in lateral view (Fig. 22) apical region with minute lateral denticles, subapical lateral process short, subtriangular and distant from apex.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAC0950FF69FC5E69A7DCE5.taxon	description	Description. Body of male (Fig. 18) strongly convex, oval, with indistinct constriction between pronotum and elytra, light reddish-brown with darker pronotum and lighter tarsi, covered with yellowish setae. BL 1.45 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.18 mm, HW 0.35 mm; frontovertexal region weakly convex at middle, unmodified; supraantennal tubercles weakly elevated. Punctures on frontovertexal area fine and inconspicuous; setae sparse, short and suberect. Antennae long and slender, with pentamerous clubs, AnL 0.75 mm, antennomere 1 and 2 each distinctly elongate, 3 indistinctly elongate, 4 about as long as broad, 5 and 6 each distinctly elongate, 7 enlarged, elongate and slightly broader than 8, 8 distinctly transverse, 9 and 10 each about as long as broad, 11 indistinctly shorter than 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and weakly flattened near hind corners, equally broad between middle and base; PL 0.45 mm, PW 0.65 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and parallel posteriorly, microserration extremely fine and barely discernible; posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each slightly closer to posterior than lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, weakly suberect. Elytra broader than pronotum, together oval, broadest near anterior third; EL 0.83 mm, EW 0.75 mm, EI 1.10; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.30 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located distinctly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra fine, unremarkable; setae similar to those on pronotum. Hind wings functional. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 19 – 24) of modified latus form, AeL 0.40 mm; capsular region of median lobe in ventral view oval, lacking pair of distolateral lobes; apical projections (Fig. 19) subtriangular and about as long as broad, in lateral view distal (dorsal) plate weakly curved; parameres short and massif, each with strongly modified apical region (Figs 20 – 22), in lateral view (Fig. 22) apical region with minute lateral denticles, subapical lateral process short, subtriangular and distant from apex, each paramere with two similarly long apical setae. Female. Externally similar to male but slightly stouter and wingless; BL 1.55 – 1.58 mm; HL 0.23 mm, HW 0.38 mm, AnL 0.73 – 0.75 mm; PL 0.45 – 0.48 mm, PW 0.68 – 0.70 mm; EL 0.85 – 0.88 mm, EW 0.75 – 0.78 mm, EI 1.13 – 1.17.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAC0950FF69FC5E69A7DCE5.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAC0950FF69FC5E69A7DCE5.taxon	etymology	Etymology. The adjective rufus refers to the reddish pigmentation.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAC0950FF69FC5E69A7DCE5.taxon	discussion	Remarks. This species is unremarkable, lacking any secondary sexual characters in males. The light pigmentation seems to be typical of fully pigmented adults, with the pronotum slightly darker than head and elytra. The most reliable diagnostic character is the unique modification of parameral apices (Figs 27 – 29), which have a small subapical process far from the insertion of setae and the subapical region is covered with asperities.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAD094EFF69FC066EF9D96D.taxon	description	(Figs 25 – 31)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAD094EFF69FC066EF9D96D.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Dali Bai Aut. Pref., / Zhemo Shan, 7 km SW Xiaguan, / 25 º 32 - 33 ’ N, 100 º 10 - 11 ’ E, 2870 - 2970 / m, scrub with bamboo, oaks & / Rhododendr., litter sifted, 18. IX. / 2009, leg. M. Schülke [CH 09 - 60] ” [white, printed]; “ CEPHENNODES (s. str.) / deinonychus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (24 exx.): 14 ♂, 4 ♀♀, 2 exx. of unknown sex (mounted with fore legs folded under the body), same data as for holotype; 1 ♂, “ CHINA: Yunnan, Dali Bai Aut. Pref., / Jizu Shan, path to cable car, 37 km NE / Dali, 25 º 58 ’ N, 100 º 23 ’ E, / 2450 m, mixed forest, sifted from litter, / moss & pine apples, 5. IX. 2009. / leg. M. Schülke [CH 09 - 29] ” [white, printed]; 2 ♂♂, 1 ♀, “ CHINA: Yunnan, Dali Bai Aut. Pref., / mount. range E Weishan, 12 km NE / Weishan, 25 º 17 ’ 02 - 15 ’’ N, 100 º 22 ’ / 23 - 30 ’’ E, 2630 - 2660 m, scrub with / pines and bamboo, litter sifted, 15. IX. / 2009, leg. M. Schülke [CH 09 - 54] ” [white, printed]; each with yellow “ PARATYPUS ” label (MNB, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAD094EFF69FC066EF9D96D.taxon	diagnosis	Diagnosis. Body moderately large, around 1.5 mm in length; antennae with pentamerous clubs; punctures on head small but sharply marked and more distinct than those on pronotum and elytra; pronotal lateral carinae single; subhumeral lines short, as long as about 0.25 × EL. Male: antennomere 7 enlarged, longer and broader than 8; protarsi (Fig. 27) conspicuously modified, each with tarsomeres 1 – 3 bearing minute ventral subtriangular projection, tarsomere 4 transverse, projecting ventrad to form a large subtriangular lobe, tarsomere 5 almost as long as 1 – 4 combined, with one claw extremely enlarged, longer than tarsomere, the other claw of normal size; aedeagus (Figs 28 – 31) in ventral view with apical projections slightly longer than wide, in lateral view distal (dorsal) plate weakly curved, capsular region in ventral view lacking rounded lobe at each distolateral corner; parameral apices unmodified.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAD094EFF69FC066EF9D96D.taxon	description	Description. Body of male (Figs 25 – 26) strongly convex, oval, with shallow constriction between pronotum and elytra, light to dark brown, in lightly pigmented specimens pronotum and head slightly darker than elytra, in fully pigmented specimens tarsi lighter than rest of body, vestiture light brown. BL 1.38 – 1.50 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.18 – 0.23 mm, HW 0.34 – 0.35 mm; frontovertexal region weakly convex at middle, unmodified; supraantennal tubercles weakly elevated. Punctures on frontovertexal area small but sharply marked, dense, separated by spaces subequal to their diameters, clypeus with finer and sparser punctures; setae sparse, short and suberect. Antennae long and slender, with pentamerous clubs, AnL 0.78 – 0.83 mm, antennomere 1 weakly elongate, 2 strongly elongate, 3 – 6 each weakly elongate, 7 enlarged, slightly elongate and slightly broader than 8, 8 and 9 each weakly transverse, 10 about as long as broad or weakly transverse, 11 about as long as 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and weakly flattened near hind corners, equally broad between middle and base or in some specimens indistinctly narrowing posterad from near middle; PL 0.43 – 0.50 mm, PW 0.63 – 0.65 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and parallel or nearly parallel posteriorly, microserration extremely fine and barely discernible; posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each slightly closer to posterior than lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, weakly suberect. Elytra indistinctly broader than pronotum, together oval, broadest between middle and anterior third; EL 0.75 – 0.88 mm, EW 0.65 – 0.73 mm, EI 1.07 – 1.25; subhumeral lines carinate but relatively blunt, as long as about 0.25 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located distinctly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra fine, unremarkable; setae similar to those on pronotum. Hind wings in some males functional, in others obliterated (but only a few have been checked). Metaventrite with shallow lateral impressions about as long as half of its length. Legs long and slender, with conspicuously modified protarsi (Fig. 27): each of tarsomeres 1 – 3 bearing minute ventral subtriangular projection, tarsomere 4 transverse, projecting ventrad to form a large subtriangular lobe, tarsomere 5 almost as long as 1 – 4 combined, with one claw extremely enlarged, longer than length of tarsomere, the other claw of normal size. Aedeagus (Figs 28 – 31) of modified latus form, AeL 0.33 mm; capsular region in ventral view oval, lacking pair of distolateral lobes; apical projections subtriangular and slightly elongate, in lateral view distal (dorsal) plate weakly curved; parameres short and thick, their apical regions unmodified, each paramere with two similarly long apical setae. Female. Externally similar to male but with unmodified protarsi and with slightly shorter antennae in relation to BL, wingless; BL 1.46 – 1.55 mm; HL 0.18 – 0.23 mm, HW 0.34 – 0.35 mm, AnL 0.75 – 0.78 mm; PL 0.48 – 0.50 mm, PW 0.65 – 0.70 mm; EL 0.80 – 0.85 mm, EW 0.73 – 0.75 mm, EI 1.07 – 1.17.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAD094EFF69FC066EF9D96D.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAD094EFF69FC066EF9D96D.taxon	etymology	Etymology. The noun deinonychus is borrowed from the name of a dinosaur and refers to a similar feature — the ‘ terrible claw’ (Greek deinós and ónyx), as in Fig. 27.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFAD094EFF69FC066EF9D96D.taxon	discussion	Remarks. Cephennodes deinonychus is the only species of the genus (and of Scydmaeninae) with an extremely enlarged one protarsal claw. Some variability in the body pigmentation (dark or light brown, see Figs 25 and 26) and in the shape of the distal plates of the median lobe (Figs 28 and 30) was found, but a relatively large number of specimens (18 males) makes it possible to interpret these differences as intraspecific or, in the case of the aedeagus, resulting from a slightly different orientation in the mounting medium.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB3094CFF69F98E6934DEC9.taxon	description	(Figs 32 – 37)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB3094CFF69F98E6934DEC9.taxon	materials_examined	Material studied. 1 ♂, Yunnan, Nujiang Lisu Aut. Pref., Gaoligong Shan, valley 18 km W Gongshan, 3020 m, 27 º 47 ’ 54 ’’ N, 98 º 30 ’ 13 ’’ E, mixed forest, litter, moss, wood sifted, 6. VI. 2007, leg. M. Schülke; 1 ♂, 3 ♀♀, Yunnan, Nujiang Lisu Aut. Pref., Gaoligong Shan, valley 21 km W Gongshan, 27 º 47 ’ 03 ’’ N, / 98 º 27 ’ 39 ’’ E, 3320 m, moss, alder, bamboo, Rhodod., sifted, 6. VI. 2007, leg. M. Schülke (MNB, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB3094CFF69F98E6934DEC9.taxon	discussion	Remarks. Cephennodes spinosus was described based on a male and female collected in Nujiang Lisu Aut. Pref., Gongshan Co., Gaoligong Shan, valley, 3000 – 3050 m, 27 ° 47.90 ’ N, 98 ° 30.19 ’ E, in a coniferous forest with Rhododendron and broad-leaved bushes, sifted from leaf litter, moss and dead wood along a creek and snowfields. The newly examined specimens come from close proximities of this locality. The most conspicuous and unique diagnostic character of this species is a subtriangular process, a mucro, on the posterior margin of the male sternite VI (Fig. 35). Although the aedeagus, as in most members of the C. excavatus group, shows some variability in the shape of distal projections (see Figs 36 – 37, vs. figs 88 – 90 in Jałoszyński (2007 a )), there is no doubt that all examined specimens belong in the same species. Fully pigmented specimens (Fig. 32) have dark brown, nearly black head and pronotum and distinctly lighter, reddish-brown elytra with infuscate suture, and males have a large and unique set of secondary sexual characters: antennomere 7 enlarged; protibia with a submedian mesal broadening; metatibia curved; elytral apex modified (Figs 33 – 34); and sternite VI with the posteromedian mucro (Fig. 35). Although the apical region of the aedeagus shows some variability within the studied sample, the presence of a pair of distolateral rounded lobes on the capsular region, and the conspicuously modified parameral apices, each with a subapical lateral process, are constant features.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB1094AFF69FE276F3ADCE5.taxon	description	(Figs 38 – 44)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB1094AFF69FE276F3ADCE5.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Nujiang Lisu Pref., / Gaoligong Shan, “ Cloud pass ”, / 21 km NW Liuku, 25 º 58 ’ 11 ’’ N, / 98 º 41 ’ 01 ’’ E, 3150 m, shrubs & / bamboo, litter sifted, 3. IX. 2009, / leg. M. Schülke [CH 09 - 22 a] ” [white, printed]; “ CEPHENNODES (s. str.) / cavatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (12 exx.): 3 ♀♀, same data as for holotype; 1 ♂, “ CHINA: Yunnan, Nujiang Lisu Pref., / Gaoligong Shan, “ Cloud pass ” 3150 m / 21 km NW Liuku (shrubs, / Vaccinium, bamboo, litter sifted) / 25 º 58 ’ 21 ’’ N, 98 º 41 ’ 01 ’’ E / 2. IX. 2009 D. W. Wrase [22 A] ” [white, printed]; 1 ♂, “ CHINA: Yunnan [CH 07 - 27], Nujiang / Lisu Aut. Pref., Gaoligong Shan, creek / valley 20 km NW Liuku, 25 º 58 ’ 49 ’’ N, / 98 º 41 ’ 48 ’’ E, 3000 m, bamboo, shrubs, / litter sifted, 9. VI. 2007, M. Schülke ” [white, printed]; 2 ♂♂, 5 ♀♀, “ CHINA: Yunnan, Nujiang Lisu Pref., / Gaoligong Shan, “ Cloud pass ”, / 21 km NW Liuku, 3150 m, / 25 º 58 ’ 21 ’’ N, / 98 º 41 ’ 01 ’’ E, / shrubs & bamboo, litter sifted, / 2. IX. 2009, leg. M. Schülke [CH 09 - 22] ” [white, printed]; all paratypes with yellow “ PARATYPUS ” label (MNB, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB1094AFF69FE276F3ADCE5.taxon	diagnosis	Diagnosis. Body moderately large, slightly over 1.5 mm in length; antennae with pentamerous clubs; punctures on head, pronotum and elytra equally fine and inconspicuous; pronotal lateral carinae single; subhumeral lines short, as long as 0.25 × EL. Male: antennomere 7 enlarged, longer and thicker than 8; apex of each elytron with oval asetose cavity (Fig. 39); aedeagus (Figs 40 – 43) with pair of rounded distolateral lobes on capsular region; each paramere with lateral subapical process (Fig. 44).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB1094AFF69FE276F3ADCE5.taxon	description	Description. Body of male (Fig. 38) strongly convex, oval, with shallow but distinct constriction between pronotum and elytra, umbra brown with slightly darker pronotum and lighter tarsi, covered with light brown setae. BL 1.55 – 1.66 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.21 – 0.25 mm, HW 0.38 mm; frontovertexal region weakly convex at middle, unmodified; supraantennal tubercles weakly elevated. Punctures on frontovertexal area fine and inconspicuous; setae sparse, short and suberect. Antennae long and slender, with pentamerous clubs, AnL 0.95 – 1.00 mm, antennomere 1 about as long as broad, 2 and 3 each distinctly elongate, 4 indistinctly elongate, 5 and 6 each distinctly elongate, 7 enlarged, elongate and slightly broader than 8, 8 slightly transverse, 9 and 10 each about as long as broad, 11 nearly as long as 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and weakly flattened near hind corners, broadest near middle but barely discernibly narrowing posterad; PL 0.48 – 0.50 mm, PW 0.65 – 0.70 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and nearly straight posteriorly, microserration extremely fine and barely discernible; posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each distinctly closer to posterior than lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, weakly suberect. Elytra clearly broader than pronotum, together oval, broadest near anterior third; EL 0.83 – 0.95 mm, EW 0.75 – 0.80 mm, EI 1.06 – 1.19; subhumeral lines carinate but blunt, as long as 0.25 × EL and weakly divergent posterad, mesally accentuated by short basal impression; basal elytral fovea on each elytron located distinctly closer to lateral margin of mesoscutellum than to subhumeral line and situated in large oval impression extending posterad; elytral apices (Fig. 39) modified, separately rounded, each with oval asetose and impunctate impression. Punctures on elytra fine, unremarkable; setae similar to those on pronotum, setae surrounding apical impressions slightly longer than on remaining surface. Hind wings functional. Metaventrite with indistinct lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 40 – 44) of modified latus form, AeL 0.35 mm; capsular region in ventral view oval, with pair of small and rounded distolateral lobes; apical projections subtriangular and about as long as broad, in ventral view distal (dorsal) plate with narrow and curved apical region, in lateral view distal plate weakly curved; parameres short and thick, each with modified apical region, in lateral view (Fig. 44) with elongate lateral subapical process, each paramere with two similarly long apical setae. Female. Externally similar to male but lacking apical elytral modifications and with shorter antennae in relation to BL, wingless; BL 1.55 – 1.58 mm; HL 0.23 mm, HW 0.38 mm, AnL 0.73 – 0.75 mm; PL 0.45 – 0.48 mm, PW 0.68 – 0.70 mm; EL 0.85 – 0.88 mm, EW 0.75 – 0.78 mm, EI 1.13 – 1.17.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB1094AFF69FE276F3ADCE5.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB1094AFF69FE276F3ADCE5.taxon	etymology	Etymology. The adjective cavatus refers to the apical elytral cavities.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB1094AFF69FE276F3ADCE5.taxon	discussion	Remarks. Cephennodes cavatus belongs to a group of species with a lateral finger-like process in the subapical region of each paramere (Fig. 44). This unusual character is shared with C. superlatus (which differs in recurved metatibiae and much more extensive elytral modifications, comprising not only large apical impressed and concave apical regions, but also large lateral patches of modified setae), C. excavatus (which has the lateral subapical parameral process shifted more proximad and directed distad, a slenderer body, and the apical elytral impressions facing dorsad), C. spinosus (which has a conspicuous mucro on the sternite VI, submedially thickened protibiae, and curved metatibiae), C. curvitibia (which has a different shape of the parameral apices and strongly curved metatibiae), and C. rufus (which also has different shapes of parameral apices, with the subapical lateral process shifted proximad, and unmodified elytra).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB70948FF69FC0669E9DF95.taxon	description	(Figs 45 – 47)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB70948FF69FC0669E9DF95.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Lincang Pref., Xue / Shan, 11 km ENE Lincang, 2510 m, / 23 º 55 ’ 01 ’’ N, / 100 º 11 ’ 17.5 ’’ E, second. / pine forest with Rhodod., small cleft with / water, litter & mushrooms sifted, / 10. IX. 2009, / leg. M. Schülke [CH 09 - 39] ” [white, printed]; “ CEPHENNODES (s. str.) / xueshanus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB70948FF69FC0669E9DF95.taxon	diagnosis	Diagnosis. Body moderately large, slightly over 1.5 mm in length; antennae gradually thickened; punctures on head small but dense, slightly more distinct than those on pronotum, punctures on pronotum small but sharply marked and more distinct than those on elytra; pronotal lateral carinae single; subhumeral lines short, as long as 0.25 × EL. Male: antennomere 7 not enlarged; aedeagus (Figs 46 – 47) with capsular region of median lobe in ventral view distally deeply divided by a long notch into lateral subtriangular elongate halves; parameral apices unmodified.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB70948FF69FC0669E9DF95.taxon	description	Description. Body of male (Fig. 45) strongly convex, oval, with shallow but distinct constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.53 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.23 mm, HW 0.36 mm; frontovertexal region weakly convex at middle, unmodified; supraantennal tubercles weakly elevated. Punctures on frontovertexal area small and shallow but distinct and dense, separated by spaces subequal to their diameters; setae sparse, short and suberect. Antennae moderately long and slender, gradually thickened, AnL 0.78 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 – 6 each about as long as broad, 7 weakly elongate and about as broad as 8, 8 weakly transverse, 9 and 10 each about as long as broad, 11 as long as 9 and 10 combined, slightly more twice as long as broad. Pronotum semi-oval, strongly convex at middle and weakly flattened near hind corners, broadest slightly anterior to middle but weakly narrowing posterad; PL 0.48 mm, PW 0.68 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and nearly straight posteriorly, microserration extremely fine and barely discernible; posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each similarly distant from posterior and lateral pronotal margin. Punctures on pronotal disc similar to those on head but slightly less distinct; setae short and moderately dense, weakly suberect. Elytra slightly broader than pronotum, together oval, broadest near anterior third; EL 0.83 mm, EW 0.73 mm, EI 1.14; subhumeral lines carinate but blunt, as long as 0.25 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra fine, unremarkable; setae similar to those on pronotum. Hind wings functional. Metaventrite with short and shallow lateral impressions occupying only its anterior 1 / 4. Legs long and slender, unmodified. Aedeagus (Figs 46 – 47) of modified latus form, AeL 0.45 mm; capsular region in ventral view oval, lacking distolateral lobes, but distally deeply divided by a long notch into lateral subtriangular halves; apical projections forming two asymmetrical groups, in ventral view distal (dorsal) plate with narrow and nearly straight subtriangular apical region, in lateral view distal plate weakly curved; parameres long and moderately thick, with unmodified apices, each paramere with two similarly long apical setae. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB70948FF69FC0669E9DF95.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB70948FF69FC0669E9DF95.taxon	etymology	Etymology. The adjective xueshanus refers to Lincang Daxueshan (= Hunhua) Mountains.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB70948FF69FC0669E9DF95.taxon	discussion	Remarks. Cephennodes xueshanus is externally unremarkable and, because males do not have any secondary sexual characters, only its genital features classify this species as a member of the C. excavatus group. The aedeagus is unique and allows for unambiguous identification. No other species has the capsular region of the median lobe deeply cleaved distally, in ventral view divided by a long notch into lateral subtriangular lobes.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB50947FF69FED66828DD85.taxon	description	(Figs 48 – 56)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB50947FF69FED66828DD85.taxon	discussion	Remarks. Cephennodes hamatus was described based on specimens collected in Nujiang Lisu Aut. Pref., Gongshan Co., Gaoligong Shan, 3000 – 3050 m, 27 ° 47.90 ’ N, 98 ° 30.19 ’ E ”, in a valley with coniferous forest with Rhododendron and broad-leaved bushes, sifted along creek and snowfields from litter, moss and dead wood. The newly recorded specimens come from a close proximity. Cephennodes hamatus is one of the most easily identifiable species of the C. excavatus group. Males have a minute ventral denticle on the protarsomere 5 (illustrated in Jałoszyński (2007 a): fig. 75), the protibia with a submedian mesal thickening (Fig. 48), and conspicuously modified elytra (Figs 49 – 51), not only with apical asetose impressions accompanied by setal penicilli, but also with large patches of modified setae on slightly flattened posterolateral regions. The distal region of the aedeagus is somewhat variable; constant features, visible also when endophallic structures are everted (Figs 52 – 55) are: a pair of rounded lobes in distolateral regions of the capsular region (clearly visible in ventral view), only slightly asymmetrical, triangular ventral apical plate about as long as broad and with a pointed apex, and parameres with unmodified apices (Fig. 56) bearing a pair of apical setae of similar lengths. External characters of males are more important for correct identification than the rather unremarkable aedeagus.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBA0947FF69FCE66E82DA97.taxon	description	(Figs 57 – 64)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBA0947FF69FCE66E82DA97.taxon	materials_examined	Material studied. CHINA (YUNNAN Prov.): 2 ♂♂, Yunnan, Dali Bai Nat. Aut. Pref., Diancang Shan, W of Dali, 25 ° 41.09 ’ N, 100 ° 06.32 ’ E, 3000 - 3200 m, sifted from litter debris in a cleft in mixed forest, 27. V. 2007, leg. A. Pütz (cAP, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBA0947FF69FCE66E82DA97.taxon	discussion	Remarks. Cephennodes hamatus was described based on specimens collected in Yunnan, Dali Bai Nat. Aut. Pref., Diancang Shan, 3 km W of Dali old town, in pine forest at „ Cloud Road ”, right upper chairlift station, 25 ° 41.1 ’ N, 100 ° 06.8 ’ E, 2650 – 2750 m, sifted from pine needles, dry moss, in ditches, and from mushrooms. The newly recorded specimens come from a close proximity, but the collecting site is at a higher altitude. Cephennodes bicolor is a relatively unremarkable species, difficult to identify. Fully pigmented males (Figs 57 – 58) are not so ‘ bicolor ’ as the specimens in the type series, but still the head and pronotum is somewhat darker than the elytra. The main diagnostic characters are: body slender; antennomere 6 slightly asymmetrical, so that antennae appear as slightly kinked (Fig. 58; in the specimen showed in Fig 57 antennae were forcefully straighten up and glued to mounting card); legs, elytra and other body parts unmodified; aedeagus with a pair of small rounded lobes in distolateral regions of the capsular region; apical structures distad the lobes longer than wide; and parameres with slightly modified apices (Figs 63 – 64), with a rapid narrowing forming a small lateral ‘ platform’ where a short and thin subapical seta is inserted, and the apex with a longer and thicker seta.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBA0947FF69F9D86FCCD87F.taxon	diagnosis	Diagnosis. Aedeagus of simonis form; median lobe asymmetrical with drop-shaped capsular region and asymmetrical parameres; among apical structures of aedeagus, tubular component situated in front of apical projection (in ventral view) is bifurcate at apex; males with unmodified abdominal sternites and unmodified metatibiae, with elytral subapical region covered with setae distinctly denser than those on remaining areas and directed posteromesad. This group is similar to the C. inflatipes, C. spatulipes and C. simplicipes groups. It currently includes two described species occurring in the Himalayas:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB80943FF69FF67682FDE5D.taxon	description	(Figs 65 – 72)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB80943FF69FF67682FDE5D.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, SE Pingbian, / 22 º 54 ’ 31 ’’ N, 103 º 41 ’ 44 ’’ E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 28. VIII. 2014, / leg. M. Schülke [CH 14 - 22 a] ” [white, printed]; “ CEPHENNODES (s. str.) / subapicalis m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (11 exx.): 1 ♂, 1 ♀, “ CHINA: Yunnan, mountain W / Gejiu, 23 º 24 ’ 13 ’’ N, 103 º 07 ’ 28 ’’ E, / 1990 m, mixed forest, litter and / various debris sifted, 23. VIII. 2014, / leg. M. Schülke [CH 14 - 20] ” [white, printed]; 1 ♂, 1 ♀, “ CHINA: Yunnan, Baoshan Pref., Gao- / ligong Shan, 65 km NNE Tengchong, / 1750 m, 25 º 35 ’ 20 ’’ N, 98 º 40 ’ 21 ’’ E, / sec. mixed forest, overgrown stone, / debris, litter and moss sifted, 31. VIII. / 2009, leg. M. Schülke [CH 09 - 10 b] ” [white, printed]; 1 ♂, 3 ♀♀, “ CHINA: Yunnan [CH 07 - 20], / Nujiang Lisu Aut. Pref., creek / valley 3 km SE Gongshan, / 1450 - 1500 m, 27 º 43 ’ 02 ’’ N, / 98 º 41 ’ 27 ’’ E, litter, moss, sifted, / 5. VI. 2007, leg. A. Pütz ” [white, printed]; 1 ♂, 1 ♀, “ CHINA: Yunnan (CH 07 - 25) / Nujiang Lisu Aut. Pref., / Salween side valley 5 km S / Fugong, road SS 228, / under stones, moist / enbarkment {sic!}, 8. VI. 2007 / leg. A. Pütz ” [white, printed]. Paratypes each with yellow “ PARATYPUS ” label (MNB, cPJ, cAP).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB80943FF69FF67682FDE5D.taxon	diagnosis	Diagnosis. Body small, around 1.1 – 1.2 mm in length; antennae with tetramerous clubs; punctures on head, pronotum and elytra small but distinct and dense, those on pronotum slightly smaller and slightly denser than those on elytra; pronotal lateral carinae single; subhumeral lines long, as long as about 0.4 × EL; in both sexes subapical adsutural region of elytra slightly flattened and covered with setae directed mesad (Fig. 66). Aedeagus (Figs 67 – 72) in ventral view with sharp-angled subtriangular apex of capsular region of median lobe and approximately trapezoidal group of apical structures strongly projecting distad.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB80943FF69FF67682FDE5D.taxon	description	Description. Body of male (Fig. 65) strongly convex, oval, with distinct constriction between pronotum and elytra, moderately light brown with tarsi lighter than rest of body, vestiture yellowish. BL 1.11 – 1.18 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.15 mm, HW 0.28 mm; frontovertexal region weakly convex at middle, unmodified; supraantennal tubercles weakly elevated. Punctures on frontovertexal area small but sharply marked, dense, separated by spaces subequal to their diameters; setae sparse, short and suberect. Antennae moderately long and slender, with tetramerous clubs, AnL 0.55 – 0.65 mm, antennomere 1 about as long as broad, 2 strongly elongate, 3 – 6 each about as long as broad, 7 slightly elongate, 8 – 10 each about as long as broad, 11 about as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and weakly flattened near hind corners, broadest near middle and slightly narrowing posterad; PL 0.35 – 0.38 mm, PW 0.50 – 0.53 mm. Anterior margin in strictly dorsal view weakly rounded; lateral margins strongly rounded in anterior half and weakly so posteriorly, microserration extremely fine and barely discernible; posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each slightly closer to posterior than lateral pronotal margin. Punctures on pronotal disc small but sharply marked and dense, in median area separated by spaces slightly narrower than diameters of punctures, area near each anterior corner covered with denser and coarser punctures; setae short and moderately dense, weakly suberect. Elytra slightly broader than pronotum, together oval, broadest near anterior third; EL 0.61 – 0.65 mm, EW 0.55 – 0.58 mm, EI 1.09 – 1.14; subhumeral lines carinate and sharp, as long as about 0.40 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located distinctly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices unmodified, separately rounded, but subapical adsutural region (Fig. 66) slightly flattened and covered with setae directed mesad. Punctures on elytra slightly larger and slightly sparser than those on pronotum; setae similar to those on pronotum. Hind wings functional. Metaventrite with shallow lateral impressions about as long as half of its length. Legs moderately long and slender, unmodified, except slightly recurved mesotibiae. Aedeagus (Figs 67 – 72) of simonis form, AeL 0.11 mm; capsular region of median lobe in ventral view drop-shaped with clearly defined subtriangular blunt apex; apical projections subtrapezoidal; parameres short and slender, in ventral view left paramere longer, and right one shorter than capsular region of median lobe, each paramere with one apical seta. Female. Externally similar to male but with slightly less distinct subapical flattening on elytra, winged; BL 1.10 – 1.13 mm; HL 0.15 mm, HW 0.28 mm, AnL 0.55 – 0.60 mm; PL 0.35 mm, PW 0.50 mm; EL 0.60 – 0.63 mm, EW 0.55 mm, EI 1.09 – 1.14.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB80943FF69FF67682FDE5D.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB80943FF69FF67682FDE5D.taxon	etymology	Etymology. The adjective subapicalis refers to the subapical elytral modification.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFB80943FF69FF67682FDE5D.taxon	discussion	Remarks. Aedeagi of all three members of the C. gokarnaensis group are very similar. In C. gokarnaensis, the capsular region of the median lobe in ventral view is more oval and more elongate than that in the two remaining species, with the apex poorly marked as an obtuse-angled rounded corner. In C. pseudogokarnaensis and C. subapicalis the capsular region is more drop-shaped, with a clearly defined sharp-angled (but still blunt) apex. Even though there is a variability in the shape of the aedeagus (Figs 67 – 72), in all studied males of C. subapicalis the distal region of the aedeagus in ventral view is invariantly longer than in C. pseudogokarnaensis. Cephennodes subapicalis is also slightly smaller than C. pseudogokarnaensis, 1.10 – 1.18 mm vs. 1.23 – 1.25 mm, respectively.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBE0943FF69FE9E6CCEDD94.taxon	diagnosis	Diagnosis. Aedeagus of modified simonis form, median lobe conspicuously large and slightly or moderately asymmetrical, with capsular region drop-shaped, oval or subrectangular, with ventral membranous area concealed by ventral wall; parameres asymmetrical, apical processes with dorsally curved hook, subapical area of dorsal wall bearing numerous minute setae. In males head modified, with convexities and / or impressions. This group is highly similar to the C. bos and C. taurus groups, it differs in unmodified antennomere 3 in males and the lack of a pair of long thickened setae on the frontovertexal region. This is a moderately species-rich group that currently includes 13 described species distributed in China, Thailand and Vietnam:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBF0940FF69FDB26F06DC55.taxon	description	(Figs 73 – 80)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBF0940FF69FDB26F06DC55.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, SE Pingbian, / 22 º 54 ’ 31 ’’ N, 103 º 41 ’ 44 ’’ E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 28. VIII. 2014, / leg. M. Schülke [CH 14 - 22 a] ” [white, printed]; “ CEPHENNODES (s. str.) / asymmetricus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (37 exx.): 6 ♂♂, 23 ♀♀, same data as for holotype; 4 ♂♂, 4 ♀♀, “ CHINA: Yunnan, SE Pingbian, / 22 º 54 ’ 31 ’’ N, 103 º 41 ’ 44 ’’ E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 27. VIII. 2014, / leg. M. Schülke [CH 14 - 22] ” [white, printed]; each with yellow “ PARATYPUS ” label (MNB, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBF0940FF69FDB26F06DC55.taxon	diagnosis	Diagnosis. Body large, BL> 1.6 mm; antennal club pentamerous; punctures on frontal region of head and on pronotum larger and deeper than those on elytra; pronotal lateral carinae single; subhumeral lines short, as long as 0.25 × EL. Male: antennomere 7 (Figs 74 – 75) longer and wider than 8, asymmetrical, with flat round mesal region delimited by sharp ridge; frontal region of head between eyes with elongate median longitudinal elevation broadening anterad and densely covered with fine punctures (Fig. 76); aedeagus (Figs 77 – 80) in ventral view with apex of capsular region broadly subtriangular and rounded, distal plate elongate, subtriangular and strongly curved dorsad. Female: antennomere 7 distinctly enlarged but symmetrical, longer and slightly broader than 8.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBF0940FF69FDB26F06DC55.taxon	description	Description. Body of male (Fig. 73) strongly convex, oval, with barely marked constriction between pronotum and elytra, light brown with slightly lighter distal regions of legs, covered with yellowish setae. BL 1.68 – 1.83 mm. Head (Fig. 76) broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.25 – 0.28 mm, HW 0.43 – 0.45 mm; frontovertexal region weakly convex; frons weakly convex, between eyes with elongate median longitudinal elevation broadening anterad and densely covered with fine punctures (Fig. 76, indicated by arrowhead); supraantennal tubercles weakly elevated. Punctures on area anterior to eyes distinct and sharply marked, unevenly distributed, on clypeus sparser, close to antennal fossae denser; setae on frons and vertex sparse, long and suberect. Antennae (Figs 73 – 75) long and slender, with pentamerous clubs, AnL 0.95 – 1.00 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 – 6 each about as long as broad, 7 enlarged and asymmetrical, longer and broader than both 6 and 8, with flat round area on mesal side delimited by sharp ridge, 8 weakly transverse, 9 and 10 each indistinctly transverse, 11 nearly as long as 9 and 10 combined, about twice as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, broadest just anterior to base; PL 0.50 – 0.55 mm, PW 0.75 – 0.83 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded and barely noticeably microserrate, posterior corners indistinctly obtuse-angled; posterior margin shallowly bisinuate; lateral marginal carinae narrow and mesally not accentuated by groove, simple (i. e., not doubled). Punctures on median region of pronotal disc large and deep, with sharp margins, separated by spaces subequal to diameters of punctures; punctures reducing in size and depth toward anterior and posterior pronotal margins, those in anterior corners indistinct, in posterior corners denser than those at middle. Setae on pronotal disc long and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior third; EL 0.93 – 1.03 mm, EW 0.80 – 0.83 mm, EI 1.15 – 1.28; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.25 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located at middle between lateral margin of mesoscutellum and subhumeral line; elytral apices unmodified, separately rounded. Punctures on elytra fine, unremarkable; setae distinctly longer than those on pronotum, dense and erect. Hind wings lacking. Metaventrite lacking lateral impressions, convex. Legs long and slender, unmodified. Aedeagus (Figs 77 – 80) of modified simonis form, AeL 0.48 mm; capsular region of median lobe in ventral view approximately drop-shaped, stout, with subtriangular and broadly rounded apex shifted to the left side; dorsal apical projection (= distal plate) in ventral view slender, elongate and tapering distad, in lateral view hook-shaped, curved dorsad at weakly obtuse angle; parameres long and slender, not reaching apex of aedeagus, each with three long subapical setae. Female. Externally similar to male, except for antennomere 7 distinctly enlarged but symmetrical, head dorsum unmodified, and slightly shorter antennae in relation to BL; wingless; BL 1.65 – 1.78 mm; HL 0.25 mm, HW 0.53 – 0.58 mm, AnL 0.90 – 0.93 mm; PL 0.55 – 0.58 mm, PW 0.78 – 0.85 mm; EL 0.88 – 0.98 mm, EW 0.78 – 0.83 mm, EI 1.09 – 1.19.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBF0940FF69FDB26F06DC55.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBF0940FF69FDB26F06DC55.taxon	etymology	Etymology. The adjective asymmetricus refers to the asymmetrical antennomere 7 in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBF0940FF69FDB26F06DC55.taxon	discussion	Remarks. Only three species of the C. impressifrons group have the antennomere 7 modified in males, and only two of them have this antennomere bearing a round mesal flat area: C. antennatus and C. asymmetricus (the third species, C. triangularis described below, has a subtriangular mesal process on antennomere 7). The Chinese Cephennodes asymmetricus and Vietnamese C. antennatus have clearly different modifications of the head in males. In the latter species (illustrated in Jałoszyński & Nomura (2009): fig. 110), there is a pair of minute frontal tubercles at middle of a large virtually impunctate area delimited anteriorly from a strongly punctate clypeofrontal region by an arcuate transverse ridge, whereas in C. asymmetricus there is neither a ridge nor tubercles (Fig. 76). Moreover, in C. antennatus, the apex of the capsular region of the median lobe forms a strongly elongate and slender rod-like process, whereas in C. asymmetricus the apex is broadly rounded; the distal plate in C. antennatus is broadening distad, vs. tapering in C. asymmetricus.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBD097EFF69FC966822DCE5.taxon	description	(Figs 81 – 84)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBD097EFF69FC966822DCE5.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA [21] - Yunnan, / Pingbian, broad- / leaved for., 23 º 00 ’ 39 ’’ N, / 103 º 42 ’ 10 ’’ E, 1500 m / 26. VIII. 2014, V. Assing ” [white, printed]; “ CEPHENNODES (s. str.) / transversicarinatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW). Paratypes (22 exx.): 5 ♂♂, 3 ♀♀, same data as for holotype; 7 ♂♂, 7 ♀♀, “ CHINA: Yunnan, NNE Pingbian, / 23 º 00 ’ 39 ’’ N, 103 º 42 ’ 10 ’’ E, 1500 / m, subtropical broad-leaved forest, / litter sifted, 26. VIII. 2014, / leg. M. Schülke [CH 14 - 21] ” [white, printed]. Each paratype with yellow “ PARATYPUS ” label (MNB, MNHW, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBD097EFF69FC966822DCE5.taxon	diagnosis	Diagnosis. Body moderately large, BL around 1.5 mm; antennae gradually thickened; punctures on head, pronotum and elytra small but distinct and dense, those on clypeus and frontovertexal region denser than those on remaining body parts; pronotal lateral carinae single; subhumeral lines short, as long as 0.3 × EL. Male: head modified (Fig. 82), clypeus demarcated from frontal region by weakly arcuate (anteriorly convex) transverse carina; scape with dorsal longitudinal ridge demarcating mesal flat surface expanding mesad; aedeagus (Figs 83 – 84) conspicuously elongate, in ventral view with distal margin of capsular region nearly symmetrical and broadly rounded, distal plate elongate and strongly curved dorsad; aedeagus with elongate endophallic structure bearing lateral denticles on its distal head-like portion.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBD097EFF69FC966822DCE5.taxon	description	Description. Body of male (Fig. 81) strongly convex, oval, with shallow constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.45 – 1.53 mm. Head (Fig. 82) broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.20 – 0.21 mm, HW 0.35 – 0.38 mm; clypeus strongly convex, so that in posterodorsal view appears as short subtrapezoidal convexity (Fig. 81); frontovertexal region weakly convex, demarcated from clypeus by arcuate transverse carina; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region small but sharply marked and dense, separated by spaces subequal to their diameters; setae sparse, long and suberect. Antennae long and slender, gradually thickened, AnL 0.73 – 0.80 mm, antennomere 1 modified, wider than long, with dorsal longitudinal ridge demarcating subrectangular mesal flat surface expanding mesad over lateral margins of clypeus and mandibular base, 2 weakly elongate, 3 – 6 each about as long as broad, 7 weakly to distinctly elongate, 8 weakly transverse, 9 and 10 each about as long as broad, 11 nearly as long as 9 and 10 combined, about twice as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, broadest near middle and barely discernibly narrowing posterad or equally broad between base and middle; PL 0.45 – 0.50 mm, PW 0.68 – 0.73 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and nearly straight or weakly rounded posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located distinctly closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc small but sharply marked, separated by spaces subequal to their diameters; punctures near anterior angles denser and somewhat coarse. Setae on pronotal disc long and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior third; EL 0.78 – 0.88 mm, EW 0.70 – 0.75 mm, EI 1.07 – 1.21; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as about 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron located slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra distinctly shallower than those on pronotum but distinct and similarly dense; setae distinctly longer than those on pronotum, dense and erect. Hind wings functional. Metaventrite with diffuse lateral impressions about as long as half of its length. Legs long and slender, unmodified. Aedeagus (Figs 83 – 84) of strongly modified simonis form, AeL 0.60 mm; capsular region of median lobe in ventral view extremely elongate, with distal margin nearly symmetrical and weakly rounded; dorsal apical projection (= distal plate) in ventral view slender, elongate and tapering distad, in lateral view hook-shaped, curved dorsad at nearly right angle; endophallus with elongate structure bearing lateral denticles on its distal head-like portion; parameres long and slender, not reaching apex of aedeagus, each with one long apical and two similarly long subapical setae. Female. Externally similar to male, except for unmodified head; wingless; BL 1.43 – 1.53 mm; HL 0.20 – 0.23 mm, HW 0.34 – 0.38 mm, AnL 0.70 – 0.73 mm; PL 0.45 – 0.50 mm, PW 0.68 – 0.70 mm; EL 0.75 – 0.85 mm, EW 0.70 – 0.75 mm, EI 1.03 – 1.14.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBD097EFF69FC966822DCE5.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBD097EFF69FC966822DCE5.taxon	etymology	Etymology. The adjective transversicarinatus refers to the transverse frontal carina on the head in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFBD097EFF69FC966822DCE5.taxon	discussion	Remarks. The unique head and scape modifications in male, and the endophallic structure with a denticulate head-like distal portion are unique features and allow for unambiguous identification of C. transversicarinatus.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF83097DFF69FC066C6ADA15.taxon	description	(Figs 85 – 88)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF83097DFF69FC066C6ADA15.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan [CH 07 - 35] Dali Bai / Auton. Pref., Wuliang Shan, 9 km SW / Weishan, 2450 – 2500 m, 25 º 10 ’ 14 ’’ N, W / slope, 100 º 14 ’ 22 ’’ E, oaks and pines, / sifted, 13. VI. 2007, / M. Schülke ” [white, printed]; “ CEPHENNODES (s. str.) / glabrifrons m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (13 exx.): 4 ♂♂, 6 ♀♀, same data as for holotype; 1 ♂, “ CHINA: Yunnan Dali Bai Auton. Pref., / Wuliang Shan, 11 km SW Weishan, / 25 º 08 ’ 46.7 ’’ N, 100 º 14 ’ 14.1 ’’ E, / 2520 m, pine forest, litter & / dead wood sifted, 14. IX. 2009, / leg. M. Schülke [CH 09 - 52] ” [white, printed]; 2 ♂♂, “ CHINA: Yunnan [CH 07 - 35], / Dali Bai Auton. Pref., Wuliang / Shan, 9 km SW Weishan, 2450 - / 2500 m, 25 º 10 ’ 14.7 ’’ N, W slope. / 100 º 14 ’ 22 ’’ E, oaks and pines, / sifted, 13. VI. 2007, leg. A. Pütz ” [white, printed]. Each paratype with yellow “ PARATYPUS ” label (MNB, cAP, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF83097DFF69FC066C6ADA15.taxon	diagnosis	Diagnosis. Body large, BL around 1.6 – 1.7 mm; antennae gradually thickened; punctures on clypeus, lateral areas of frontovertexal region, pronotum and elytra small but distinct and dense, those on pronotum more distinct than on elytra; pronotal lateral carinae single; subhumeral lines short, as long as 0.3 × EL. Male: head modified (Fig. 86), large elongate and rhomboidal median region of frontovertexal area asetose and impunctate, posteriorly flat and anteriorly slightly impressed, its anterior margin subtriangular and projecting anterad as short horn-like ridge, impunctate area with deep longitudinal median groove; aedeagus (Figs 87 – 88) strongly elongate, in ventral view with distal margin of capsular region asymmetrical and broadly subtriangular, distal plate short and strongly curved dorsad.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF83097DFF69FC066C6ADA15.taxon	description	Description. Body of male (Fig. 85) strongly convex, oval, with very shallow constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.63 – 1.70 mm. Head (Fig. 86) broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.23 – 0.25 mm, HW 0.39 – 0.40 mm; frontovertexal region modified, with large elongate and rhomboidal median region asetose and impunctate, posteriorly flat and anteriorly slightly impressed, its anterior margin subtriangular and projecting anterad as short horn-like ridge; supraantennal tubercles weakly elevated. Punctures on clypeus and lateral areas of frontovertexal region small but sharply marked and dense, separated by spaces subequal to their diameters; setae on lateral areas sparse, long and suberect. Antennae long and slender, gradually thickened, AnL 0.85 – 0.93 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 weakly transverse, 4 – 6 each about as long as broad, 7 weakly elongate, 8 – 10 each about as long as broad, 11 as long as 9 and 10 combined, about twice as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, broadest near middle but barely discernibly narrowing posterad, so in some specimens it appears as equally wide between base and middle; PL 0.50 – 0.55 mm, PW 0.75 – 0.78 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc small but sharply marked, separated by spaces subequal to their diameters; punctures near anterior angles denser. Setae on pronotal disc long and moderately dense, suberect. Elytra as wide as pronotum, together oval, broadest near anterior third; EL 0.83 – 0.90 mm, EW 0.75 – 0.78 mm, EI 1.10 – 1.17; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as about 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron similarly distant from lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra distinctly shallower and slightly sparser than those on pronotum but distinct; setae distinctly longer than those on pronotum, dense and erect. Hind wings in some specimens functional, in others obliterated (but not in all specimens were the wings studied). Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 87 – 88) of modified simonis form, AeL 0.48 mm; capsular region of median lobe in ventral view strongly elongate, with distal margin asymmetrical and broadly subtriangular; dorsal apical projection (= distal plate) in ventral view short and tapering distad, in lateral view hook-shaped, curved dorsad at nearly right angle; parameres long and slender, not reaching apex of aedeagus, each with one long apical and two similarly long subapical setae. Female. Externally similar to male, except for unmodified head and slightly shorter antennae in relation to BL; wingless; BL 1.58 – 1.68 mm; HL 0.23 – 0.25 mm, HW 0.39 – 0.43 mm, AnL 0.80 – 0.85; PL 0.50 – 0.58 mm, PW 0.75 – 0.83 mm; EL 0.85 – 0.88 mm, EW 0.75 – 0.80 mm, EI 1.06 – 1.17.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF83097DFF69FC066C6ADA15.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF83097DFF69FC066C6ADA15.taxon	etymology	Etymology. The adjective glabrifrons refers to the glabrous median area of the frontovertexal region in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF83097DFF69FC066C6ADA15.taxon	discussion	Remarks. The head modifications in male composed of an elongate flat asetose and impunctate area divided medially by a groove and anteriorly projecting into a subtriangular process allow for unambiguous identification of C. glabrifrons.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF80097AFF69FA566989DE91.taxon	description	(Figs 89 – 92)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF80097AFF69FA566989DE91.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA [3] - Yunnan, E / Kunming, 24 º 55 ’ 43 ’’ N, / 103 º 05 ’ 22 ’’ E, 2110 m / Xiaobailong Forest Park / 10. VIII. 2014, V. Assing ” [white, printed]; “ CEPHENNODES (s. str.) / tuberculatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW). Paratypes (75 exx.): 6 ♂♂, 17 ♀♀, same data as for holotype; 9 ♂♂, 41 ♀♀, “ CHINA: Yunnan, E Kunming, / Xiaobailong Forest Park, 2110 m, 24 º 55 ’ 43 ’’ N, 103 º 05 ’ 22 ’’ E, sec. / pine forest margin, litter sifted, / 10. VIII. 2014, Schülke [CH 14 - 03] ” [white, printed]; 1 ♂, 1 ♀, “ CHINA: Yunnan, E Kunming, / 25 º 09 ’ 07 ’’ N, 102 º 53 ’ 46 ’’ E, 2280 m / secondary pine forest, with scattered old alder, litter sifted, / 11. VIII. 2014, Schülke [CH 14 - 04] ” [white, printed]. Each paratype with yellow “ PARATYPUS ” label (MNB, MNHW, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF80097AFF69FA566989DE91.taxon	diagnosis	Diagnosis. Body large, BL around 1.6 mm; antennae with indistinctly delimited trimerous clubs; punctures on vertexal region, pronotum and elytra small but distinct and dense, those on pronotum distinctly deeper than those on elytra; pronotal lateral carinae single; subhumeral lines short, as long as 0.3 × EL. Male: head modified (Fig. 90), with pair of transverse subtriangular lateral asetose and impunctate impressions between eyes, elongate median impunctate tubercle just anterior to anteromesal margins of impressions bearing posteriorly elongate group of setae, and pair of large lateral tubercles adjacent to supraantennal tubercles and covered with setae, flanking median asetose and impunctate area where tubercle is situated; aedeagus (Figs 91 – 92) in ventral view with distal margin of capsular region broadly rounded, distal plate elongate and strongly curved dorsad.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF80097AFF69FA566989DE91.taxon	description	Description. Body of male (Fig. 89) strongly convex, oval, with very shallow constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.56 – 1.65 mm. Head (Fig. 90) broadest across moderately large and conspicuously weakly convex, coarsely faceted eyes, HL 0.21 – 0.25 mm, HW 0.36 – 0.39 mm; frontovertexal region with pair of transverse subtriangular lateral impressions between eyes, elongate median tubercle, and pair of large lateral tubercles adjacent to supraantennal tubercles; supraantennal tubercles weakly elevated. In not fully pigmented specimens, a pair of minute dark tubercles can be seen on posterior frontal region (Fig. 90), in fully pigmented males these tubercles are difficult to observe. Punctures on clypeus sparse and mainly situated on its lateral regions; median tubercle and surrounding area and lateral subtriangular impressions impunctate and asetose, punctures on areas posterior to impressions small but sharply marked and dense, separated by spaces subequal to their diameters; setae on punctate areas unevenly distributed, those on lateral tubercles dense, on clypeus and posterior frontovertexal region sparse and moderately long, median tubercle and median longitudinal area behind it with group of long dense setae. Antennae long and slender, with indistinctly demarcated trimerous clubs, AnL 0.75 – 0.78 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 weakly transverse, 4 – 6 each about as long as broad, 7 weakly elongate, 8 – 10 each about as long as broad, 11 as long as 9 and 10 combined, nearly twice as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, broadest near middle or equally broad between base and middle; PL 0.50 mm, PW 0.68 – 0.73 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc small but sharply marked, separated by spaces subequal to their diameters; punctures near anterior angles denser. Setae on pronotal disc long and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior third; EL 0.85 – 0.93 mm, EW 0.73 – 0.78 mm, EI 1.13 – 1.23; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as about 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron almost equidistant from lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra distinctly shallower and slightly sparser than those on pronotum but distinct; setae distinctly longer than those on pronotum, dense and erect. Hind wings in some specimens functional, in others distinctly shortened (but only in a few specimens wings have been studied). Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 91 – 92) of modified simonis form, AeL 0.30 mm; capsular region of median lobe in ventral view oval, with distal margin broadly rounded; dorsal apical projection (= distal plate) in ventral view slender, elongate and tapering distad, in lateral view hook-shaped, curved dorsad at nearly right angle; parameres long and slender, not reaching apex of aedeagus, each with one long apical and two similarly long subapical setae. Female. Externally similar to male, except for unmodified head; wingless; BL 1.58 – 1.63 mm; HL 0.23 – 0.25 mm, HW 0.38 – 0.40 mm, AnL 0.68 – 0.75 mm; PL 0.49 – 0.53 mm, PW 0.70 – 0.75 mm; EL 0.83 – 0.90 mm, EW 0.73 – 0.80 mm, EI 1.09 – 1.20.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF80097AFF69FA566989DE91.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF80097AFF69FA566989DE91.taxon	etymology	Etymology. The adjective tuberculatus refers to the median convexity between supraantennal tubercles in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF80097AFF69FA566989DE91.taxon	discussion	Remarks. Cephennodes tuberculatus can be easily identified by unique head modifications in males: a pair of transverse subtriangular lateral asetose and impunctate impressions between eyes, an elongate median impunctate tubercle just anterior to impressions bearing a dorsal group of setae, and a pair of large lateral tubercles adjacent to supraantennal tubercles and covered with setae, flanking median asetose and impunctate area where the tubercle is situated. Moreover, the eyes of males and females of this species are conspicuously weakly convex, much less so than in all other species of the C. impressifrons group. The aedeagus of C. tuberculatus within the type series from Kunming has an oval capsular region of the median lobe (Fig. 91), and although clearly differs from the genitalia of all remaining members of the C. impressifrons group in a broadly rounded, not subtriangular apex of the capsular region of the median lobe, this character makes it highly similar to some species of the closely related C. bos (C. pseudobos, C. buceros) and C. taurus (C. clypeicornis, C. qiongdaoanus) groups, which clearly differ in secondary male dimorphic characters of the head. One male from a mountain near Yuxi has the capsular region of the aedeagus drop-shaped, with a clearly defined subtriangular apex, and somewhat different shape of the apical group of projections (Figs 93 – 94). Externally, this specimen does not differ in any characters from those from Kunming, and all structures on the head in males appear identical. This male may represent a separate cryptic species, but a larger sample from Yuxi is required to test this hypothesis. The specimens from Yuxi are not included in the type series of C. tuberculatus, and they comprise 1 ♂ and 5 ♀♀, with the following data: “ CHINA: Yunnan, mountain W Yuxi, 24 º 27 ’ 11 ’’ N, / 102 º 29 ’ 58 ’’ E, 2250 m, secondary mixed forest, litter, roots and moss sifted, 31. VIII. 2014, M. Schülke [CH 14 - 23] ” (cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF870978FF69FDD26804DCE5.taxon	description	(Figs 95 – 103)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF870978FF69FDD26804DCE5.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA [1] - Yunnan, / Wuding, Lion Mountain / 2200 m, 25 º 31 ’ 59 ’’ N, / 102 º 22 ’ 36 ’’ E, sifted / 17. VIII. 2014, V. Assing ” [white, printed]; “ CEPHENNODES (s. str.) / triangularis m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW). Paratypes (21 exx.): 3 ♀♀, same data as for holotype; 2 ♂♂, 4 ♀♀, “ CHINA [20] - Yunnan, / mt. W Gejiu, mixed / forest, 23 º 24 ’ 13 ’’ N, / 103 º 07 ’ 28 ’’ E, 1990 m, / 23. VIII. 2014, V. Assing ” [white, printed]; 5 ♂♂, 2 ♀♀, “ CHINA: Yunnan, mountain W / Gejiu, 23 º 24 ’ 13 ’’ N, 103 º 07 ’ 28 ’’ E, / 1990 m, mixed forest, litter and / various debris sifted, 23. VIII. 2014, / leg. M. Schülke [CH 14 - 20] ” [white, printed]; 1 ♂, 2 ♀♀, “ CHINA: Yunnan, mountain W / Gejiu, 23 º 24 ’ 13 ’’ N, 103 º 07 ’ 28 ’’ E, / 1990 m, mixed forest, litter and / various debris sifted, 24. VIII. 2014, / leg. M. Schülke [CH 14 - 20 a] ” [white, printed]; 2 ♀♀, “ CHINA: Yunnan, mountain W / Gejiu, 23 º 24 ’ 13 ’’ N, 103 º 07 ’ 28 ’’ E, / 1990 m, mixed forest, litter and / various debris sifted, 25. VIII. 2014, / leg. M. Schülke [CH 14 - 20 b] ” [white, printed]. Each paratype with yellow “ PARATYPUS ” label (MNB, MNHW, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF870978FF69FDD26804DCE5.taxon	diagnosis	Diagnosis. Body large, BL around 1.5 – 1.6 mm; antennae with pentamerous clubs; punctures on frontovertexal region smaller than those on pronotum and elytra, punctures on pronotum small but sharply marked and dense, those on elytra slightly shallower, larger and sparser than those on pronotum; pronotal lateral carinae single; subhumeral lines short, as long as 0.3 × EL. Male: head modified (Figs 96 – 98), with pair of slightly transverse oval lateral asetose and impunctate impressions between antennal insertions and broad, dense, erect median tuft of setae between eyes appearing as curved ‘ horn’ (Fig. 98); antennae modified, with antennomere 7 strongly enlarged and mesally projecting into subtriangular convexity; aedeagus (Figs 100 – 103) in ventral view with distal margin of capsular region subtriangular. Female: antennomere 7 enlarged, distinctly longer than 8 and subequal in width.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF870978FF69FDD26804DCE5.taxon	description	Description. Body of male (Fig. 95) strongly convex, oval, with very shallow constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.48 – 1.66 mm. Head (Figs 96 – 98) broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.24 – 0.25 mm, HW 0.38 – 0.43 mm; frontovertexal region with pair of slightly transverse oval lateral asetose and impunctate impressions between antennal insertions and broad, dense median erect tuft of setae between eyes appearing as curved ‘ horn’ (Fig. 98). Punctures on clypeus large and dense, those behind impressions slightly smaller and sparser; setae on punctate areas moderately long, sparse, suberect. Antennae (Fig. 99) long and slender, with pentamerous clubs, AnL 0.88 – 0.93 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 – 6 each about as long as broad, 7 strongly enlarged, asymmetrical, with large mesal subtriangular convexity, 8 – 10 each indistinctly elongate, 11 as long as 9 and 10 combined, slightly more than twice as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, broadest slightly posterior to middle and weakly but distinctly narrowing posterad; PL 0.43 – 0.53 mm, PW 0.70 – 0.78 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and weakly so posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc small but sharply marked and deep, separated by spaces subequal to their diameters; punctures near anterior angles denser and coarse. Setae on pronotal disc long and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior third; EL 0.78 – 0.93 mm, EW 0.70 – 0.78 mm, EI 1.06 – 1.23; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as about 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron almost equidistant from lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra distinctly shallower and slightly sparser than those on pronotum but distinct; setae distinctly longer than those on pronotum, dense and erect. Hind wings lacking. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 100 – 103) of modified simonis form, AeL 0.43 mm; capsular region of median lobe in ventral view approximately drop-shaped, with distal margin subtriangular; dorsal apical projection (= distal plate) in ventral view slender, elongate and tapering distad, in lateral view hook-shaped, curved dorsad at nearly right angle; parameres long and slender, not reaching apex of aedeagus, each with one long apical and two similarly long subapical setae. Female. Externally similar to male, except for unmodified head and antennae, and antennae slightly shorter in relation to BL; wingless; BL 1.54 – 1.63 mm; HL 0.21 – 0.25 mm, HW 0.35 – 0.43 mm, AnL 0.70 – 0.83 mm; PL 0.48 – 0.53 mm, PW 0.68 – 0.78 mm; EL 0.83 – 0.90 mm, EW 0.70 – 0.78 mm, EI 1.07 – 1.20.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF870978FF69FDD26804DCE5.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF870978FF69FDD26804DCE5.taxon	etymology	Etymology. The adjective triangularis refers to the triangular antennomere 7 in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF870978FF69FDD26804DCE5.taxon	discussion	Remarks. Cephennodes triangularis is the only species of the genus with the antennomere 7 bearing a large mesal subtriangular convexity. A small variability in the median lobe of aedeagus was found in this species (Figs 100 – 103); all males have the capsular region distally subtriangular, but the shape of the apex slightly varies.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF850976FF69FC066836DF95.taxon	description	(Figs 104 – 108)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF850976FF69FC066836DF95.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ China: Yunnan Prov. / Nabanhe N. R. / Benggangxinzhai / alt. 1750 m, 03 - V - 2009 / Jia-Yao Hu & / Zi-Wei Yin leg. ” [white, printed]; “ CEPHENNODES (s. str.) / biimpressus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (SNUC). Paratypes (12 exx.): 8 ♂♂, 2 ♀♀, same data as for holotype; 2 ♂♂, same data except for 1900 m and 01 - V- 2009. Each paratype with yellow “ PARATYPUS ” label (SNUC, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF850976FF69FC066836DF95.taxon	diagnosis	Diagnosis. Body large, BL around 1.6 – 1.7 mm; antennae gradually thickened; punctures on clypeus and frontovertexal region outside transverse impressions sharply marked and dense, separated by spaces narrower than their diameters, punctures on pronotum larger and deeper than those on head, also conspicuously dense, those on elytra contrastingly superficial; pronotal lateral carinae single; subhumeral lines short, as long as 0.3 × EL. Male: head modified (Figs 105 – 106), with pair of transverse impunctate and asetose impressions between antennal insertions, each with sharply marked ridge-like anterior margin, and posteriorly gradually becoming shallower; aedeagus (Figs 107 – 108) in ventral view strongly elongate, with distal margin of capsular region only slightly asymmetrical and not triangular.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF850976FF69FC066836DF95.taxon	description	Description. Body of male (Fig. 104) strongly convex, oval, with very shallow constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.63 – 1.73 mm. Head (Figs 105 – 106) broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.23 – 0.25 mm, HW 0.43 mm; frontovertexal region with pair of large transverse impunctate and asetose impressions between antennal insertions, each with sharply marked ridge-like anterior margin and connected at middle, posteriorly each impression gradually becoming shallower. Punctures on clypeus and frontovertexal region behind impressions conspicuously distinct, sharply marked and dense, punctures separated by spaces narrower than their diameters; setae on punctate areas moderately long, sparse, suberect. Antennae long and slender, gradually thickened, AnL 0.90 – 0.95 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 and 4 each about as long as broad, 5 and 6 each weakly elongate, 7 strongly elongate, 8 – 10 each indistinctly elongate, 11 as long as 9 and 10 combined, slightly more than twice as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, broadest near middle and weakly but distinctly narrowing posterad; PL 0.50 – 0.53 mm, PW 0.73 – 0.78 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and weakly so posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc small but sharply marked and deep, separated by spaces slightly narrower than their diameters; punctures near anterior angles denser and coarse. Setae on pronotal disc long and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior third; EL 0.88 – 0.98 mm, EW 0.78 – 0.80 mm, EI 1.09 – 1.26; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as about 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron almost equidistant from lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra much shallower and slightly sparser than those on pronotum, superficial; setae distinctly longer than those on pronotum, dense and erect. Hind wings functional. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 107 – 108) of strongly modified simonis form, AeL 0.60 mm; capsular region of median lobe in ventral view strongly elongate and oval, with distal margin weakly asymmetrical and not triangular; dorsal apical projection (= distal plate) in ventral view slender, elongate and tapering distad, in lateral view hook-shaped, curved dorsad at nearly right angle; parameres long and slender, not reaching apex of aedeagus, each with one long apical and two similarly long subapical setae. Female. Externally similar to male, except for unmodified head, and antennae slightly shorter in relation to BL; wings not studied; BL 1.65 mm; HL 0.25 mm, HW 0.44 mm, AnL 0.88 mm; PL 0.55 mm, PW 0.80 mm; EL 0.85 mm, EW 0.80 mm, EI 1.06.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF850976FF69FC066836DF95.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF850976FF69FC066836DF95.taxon	etymology	Etymology. The adjective biimpressus refers to a pair of frontal impressions in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF850976FF69FC066836DF95.taxon	discussion	Remarks. Cephennodes biimpressus can be easily identified by male head modifications: a pair of transverse impunctate and asetose impressions in front of eyes, with a sharply marked anterior margin developed as a biarcuate ridge, and posteriorly gradually becoming shallower. Moreover, the strongly elongate aedeagus with only weakly asymmetrical distal margin of the capsular region without a defined apex is also characteristic of this species.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8B0975FF69FED669EFDB17.taxon	description	(Figs 109 – 114)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8B0975FF69FED669EFDB17.taxon	materials_examined	Material studied. CHINA (YUNNAN Prov.): 1 ♂, 1 ♀, Baoshan Pref., Gaoligong Shan, 29 km ESE Tengchong, 24 º 55 ’ 37 ’’ N, 98 º 45 ’ 09 ’’ E, 2350 m, devastated deciduous forest, litter, wood and fungi, 1. VI. 2007, leg. M. Schülke; 3 ♂♂, 5 ♀♀, Baoshan Pref., Gaoligong Shan, E pass, 36 km SE Tengchong, 24 º 49 ’ 32 ’’ N, 98 º 46 ’ 06 ’’ E, 2200 m, devastated deciduous forest, litter, wood and fungi, 31. V. 2007, leg. M. Schülke; 1 ♂, Baoshan Pref., Gaoligong Shan, 33 km SE Tengchong, 24 º 51 ’ 02 ’’ N, 98 º 45 ’ 36 ’’ E, 2100 - 2200 m, deciduous forest, litter, wood and fungi, 31. V. 2007, leg. M. Schülke; 2 ♂♂, 2 ♀♀, Baoshan Pref., Gaoligong Shan, W pass 35 km SE Tengchong, 24 º 50 ’ 18 ’’ N, 98 º 45 ’ 43 ’’ E, 2100 m, devastated primeval deciduous forest, litter, wood and fungi, 25. VIII. 2009, leg. M. Schülke; 3 ♂♂, 1 ♀, Baoshan Pref., Gaoligong Shan, W pass 35 km SE Tengchong, 24 º 50 ’ 18 ’’ N, 98 º 45 ’ 43 ’’ E, 2100 m, devastated primeval deciduous forest, litter, wood and fungi, 28. VIII. 2009, leg. M. Schülke; 2 ♂♂, 2 ♀♀, Baoshan Pref., Gaoligong Shan, 32 km SE Tengchong, 24 º 51 - 53 ’ N, 98 º 45 ’ E, 2150 - 2250 m, devastated primeval deciduous forest, litter, wood and fungi, 26. VIII. 2009, leg. M. Schülke; 1 ♂, Baoshan Pref., mountain range 22 km S Tengchong, 24 º 49 ’ 29 ’’ N, 98 º 29 ’ 30 ’’ E, 1800 m, devastated primeval deciduous forest, litter, wood and fungi, 30. VIII. 2009, leg. M. Schülke; 1 ♂, 3 ♀♀, Baoshan Pref., Gaoligong Shan, 29 km ESE Tengchong, 24 º 55 ’ 37 ’’ N, 98 º 45 ’ 09 ’’ E, 2350 m, devastated deciduous forest, litter, wood and fungi, 1. VI. 2007, leg. A. Pütz; 1 ♂, Baoshan Pref., Gaoligong Shan, near Xiaoheishan N. R., 35 km SE Tengchong, 24 º 50 ’ 16 ’’ N, 98 º 45 ’ 43 ’’ E, 2110 m, deciduous forest, litter, 30. V. 2007, leg. A. Pütz. Specimens in MNB, cAP, cPJ.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8B0975FF69FED669EFDB17.taxon	discussion	Remarks. Cephennodes carinifrons was described based on specimens collected in Dabei (2,430 m), Gaoligongshan Mts at altitudes above 2200 m. The newly recorded specimens come from the same area and demonstrate that this species is relatively common at high altitudes of Gaoligong Shan. Examination of a large sample of males revealed that the aedeagus of C. carinifrons shows a low variability, mainly restricted to the length and shape of the distal processes. The distal margin of the capsular region in ventral view is quite stable and the aedeagi examined during the present study differ only slightly from those illustrated in Jałoszyński (2007 a): figs 52 – 54. Externally, males of this species are most similar to those of C. triangulifrons, differences in head modifications are illustrated in Figs 110 (C. carinifrons; median impression broader and anteriorly arcuate) and Fig. 116 (C. triangulifrons; median impression narrower and anteriorly angulate).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF880975FF69FB586E3BD9F5.taxon	description	(Figs 115 – 118)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF880975FF69FB586E3BD9F5.taxon	materials_examined	Material studied. CHINA (YUNNAN Prov.): 1 ♂, Baoshan Pref., 10 km SE Kambaiti Pass, 45 km NW Tengchong, 25 º 21 ’ 13 - 29 ’’ N, 98 º 13 ’ 39 - 54 ’’ E, 1700 - 1800 m, primary forest, litter, fungi, 29. VIII. 2009, leg. M. Schülke (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF880975FF69FB586E3BD9F5.taxon	discussion	Remarks. Cephennodes carinifrons was described based on specimens collected in Huanxipo and on Mt. Dayingshan, Tengchong X. (1950 – 1990 m). The newly recorded specimen comes also from Tengchong county. The aedeagus of the newly examined specimen is slightly different from that illustrated in Jałoszyński (2007 a): figs 49 – 51, mainly in the shape of the apical region. However, the shape of the distal region of the basal capsule is very similar, and differences can be attributed to delicate structures being slightly misshaped during the preparation and mounting. Externally, this male is identical to the holotype and can be easily distinguished from the similar C. carinifrons by the male head modification illustrated in Fig. 116.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF880974FF69F8F76C2FDEED.taxon	diagnosis	Diagnosis. Externally unremarkable, lacking male secondary sexual characters; aedeagus of modified simonis form: in ventral view, median lobe drop-shaped but with apical portion of basal capsule strongly elongate and forming nearly rod-like projection; apical projection of median lobe in ventral view broadly T-shaped, its apex with two clearly defined distolateral corners developed as subtriangular lobe (in ventral view left one) and scythe-shaped projection (right one); parameres asymmetrical, each with three apical / subapical setae. Species of this group are very difficult to identify. In many cases, the aedeagi are highly similar between species, but external characters (especially body size and proportions of body parts) differ markedly. Moreover, the shape of the aedeagal structures strongly depends on the orientation in mounting medium, and even a slight rotation can cause the aedeagus of the same specimen look different (Figs 120, 122 and 123 show such differences). Great care should be taken to compare specimens with the type material of previously described species, and to compare all measurements, punctuation pattern, proportions of antennomeres et cet., to avoid misidentifications. This is a moderately species-rich group that currently includes 12 described species distributed in Nepal, Thailand and Japan:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF890972FF69FACA6F24DA15.taxon	description	(Figs 119 – 123)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF890972FF69FACA6F24DA15.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan [CH 07 - 13], Baoshan / Pref., Gaoligong Shan, E pass, 36 km SE / Tengchong, 2200 m, 24 º 49 ’ 32 ’’ N, / 98 º 46 ’ 06 ’’ E, decid. forest, litter, wood, / fungi sifted, 31. V. 2007, M. Schülke ” [white, printed]; “ CEPHENNODES (s. str.) / unus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (2 exx.): 1 ♂, “ CHINA: Yunnan, Baoshan Pref., Gao- / ligong Shan, 33 km SE / Tengchong, / 2150 m, 24 º 51 ’ 22 ’’ N, 98 º 45 ’ 36 ’’ E, / devast. primary deciduous forest, / litter, wood, mushrooms sifted, 28. VII. / 2009, leg. M. Schülke [CH 09 - 08 a] ” [white, printed] (cPJ); 1 ♂, “ CHINA: Yunnan [CH 07 - 11], / Baoshan Pref., Gaoligong Shan, / nr. Xiaoheishan N. R., 35 km SE / Tengchong, 2110 m, / 24 º 50 ’ 16 ’’ N, 98 º 45 ’ 43 ’’ E, decid. / forest, litter, sifted, 30. V. 2007, / leg. A. Pütz ” [white, printed] (cAP). Each paratype with yellow “ PARATYPUS ” label.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF890972FF69FACA6F24DA15.taxon	diagnosis	Diagnosis. Body small, BL around 1.2 – 1.3 mm; antennae gradually and strongly thickened; punctures on clypeus fine and inconspicuous, on frontovertexal region more distinct, small but sharply marked and dense, separated by spaces subequal to their diameters, punctures on pronotum and elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: lacking secondary sexual characters; aedeagus (Figs 120 – 123) in ventral view with strongly elongate and narrow apical region of capsular portion tapering distad, parameres not exceeding apex of median lobe.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF890972FF69FACA6F24DA15.taxon	description	Description. Body of male (Fig. 119) strongly convex, oval, with very shallow constriction between pronotum and elytra, light brown with slightly lighter tarsi, covered with yellowish brown setae. BL 1.21 – 1.28 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.15 – 0.16 mm, HW 0.29 – 0.30 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus fine and inconspicuous, those on frontovertexal region more distinct, small, sharply marked and dense, separated by spaces subequal to their diameters; setae short, sparse, suberect. Antennae long and slender, gradually and strongly thickened, AnL 0.58 – 0.60 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 – 7 each about as long as broad, 8 – 10 each transverse, 11 about as long as 9 and 10 combined, about 1.6 × as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, equally broad between base and middle; PL 0.35 mm, PW 0.53 – 0.58 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous except for a few distinct small punctures near each hind corner. Setae on pronotal disc short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.70 – 0.78 mm, EW 0.60 – 0.63 mm, EI 1.17 – 1.24; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as about 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum In the three specimens studied, hind wings are functional in one, vestigial in the second, and completely obliterated in the third male. Metaventrite with lateral impressions about as long as 3 / 4 of its length. Legs long and slender, unmodified. Aedeagus (Figs 120 – 123) of modified simonis form, AeL 0.20 mm; capsular region of median lobe in ventral view strongly elongate and subtriangular with strongly elongate and narrow distal portion tapering towards apex; dorsal apical projection (= distal plate) in ventral view with rounded left distolateral lobe and long, pointed right scythe-shaped process; parameres long and slender, not reaching apex of aedeagus, each with one long apical and two similarly long subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF890972FF69FACA6F24DA15.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF890972FF69FACA6F24DA15.taxon	etymology	Etymology. The Latin adjective unus means one, a single. Cephennodes unus is the only Chinese species that belongs in the C. oblongopunctatus group.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF890972FF69FACA6F24DA15.taxon	discussion	Remarks. Among the Nepalese species of this group, the main genital diagnostic character is the shape of the apical region of the capsular portion of the median lobe in ventral view. Within the Himalayan group, this apical region is commonly broadening distad, and in a few cases nearly parallel-sided with rounded or truncate distal margin. In contrast, C. unus has the apical region tapering distad. In C. falcipenis and C. oblongopunctatus (Thailand), the apical region is either straight, parallel-sided and obliquely truncate, or curved mesad, respectively, none of the features occurring in C. unus. Unfortunately, the way the aedeagi of the Japanese species C. kotori and C. miharu have been illustrated makes it not possible to compare the newly described C. unus with these species. Cephennodes miharu was described with BL 1.35 ‒ 1.53 mm, and C. kotori 1.34 ‒ 1.50 mm, whereas C. unus is distinctly smaller, with BL 1.21 ‒ 1.28 mm.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8F0971FF69FA5668CADC21.taxon	diagnosis	Diagnosis. Conspicuously shortened maxillary palpomere 4, not elongate and strongly narrowing towards truncate apex (Fig. 124), but about as long as broad or even slightly transverse and subcylindrical (Figs 125, 126); aedeagus of modified simonis form, with elongate drop-shaped capsular region of median lobe, its distal region subtriangular, and apical dorsal plate distally also subtriangular, ventral wall lacking setae; males lacking secondary sexual characters. This is a newly established species group. The structure of the maxillary palp is relatively stable within all remaining genera of Scydmaeninae, and the difference between the C. palpalis group and all remaining species of Cephennodes could justify a separate status of these three species, even in a separate genus. The maxillary palpomere 4 in all remaining species of Cephennodes is elongate, as in Fig. 124, and it seems to strongly differ from that in Figs 125 and 126. However, all remaining groundplan morphological characters of the new group are identical to those in all other Cephennodes spp., and also the maxillary palpomere 4 even within the palpalis group is clearly variable in width and length in relation to the penultimate segment. It seems that this modification, restricted to a closely related three species (judging from their very similar aedeagi) is rather a transformation that has evolved within the Cephennodes clade, and not even within its most basal branches. This conclusion is made based on the fact that in all remaining genera of Cephenniini the aedeagus is symmetrical, except for Hlavaciellus, a sister group of Cephennodes (Jałoszyński 2011 b; a, 2012 a, 2018), and also the symmetrical form is preserved in some groups of species within Cephennodes. Therefore, the asymmetry, as that in the C. palpalis group, is a derived condition. The ancestor of Cephennodes and Hlavaciellus had a symmetrical aedeagus. For this reason, a group of species with a strongly asymmetrical aedeagus and a transformed maxillary palpomere 4 is very unlikely a sister group of all remaining Cephennodes species. This is a small group that includes the Chinese C. palpalis Jałoszyński, 2015 c and two newly described species.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8C096FFF69FC436E67DCE5.taxon	description	(Figs 125, 127 – 129)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8C096FFF69FC436E67DCE5.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan / Nujiang Lisu Aut. Pref., / Salween side valley 5 km S / Fugong, road SS 228, / km 223 (creek bank, litter sifted) / 8. VI. 2007 D. W. Wrase [25] ” [white, printed]; “ CEPHENNODES (s. str.) / pseudopalpalis m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (NKME).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8C096FFF69FC436E67DCE5.taxon	diagnosis	Diagnosis. Body small, BL around 1.15 mm; antennae gradually thickened; punctures on head and pronotum fine and inconspicuous, those on adsutural half of each elytron distinct, sharply marked and conspicuously dense, separated by spaces slightly narrower than their diameters, punctures becoming smaller towards lateral and posterior elytral margins; subhumeral lines short, as long as 0.2 × EL; elytra with conspicuous setal pattern: setae on densely punctate adsutural halves directed posterad, those on outer halves directed laterad. Male: lacking secondary sexual characters (unless setal elytral pattern is such a character); aedeagus (Figs 128 – 129) in ventral view with nearly straight and conspicuously narrow apical region of capsular portion pointed at apex; distal margin of apical plate nearly straight.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8C096FFF69FC436E67DCE5.taxon	description	Description. Body of male (Fig. 127) strongly convex, oval, with very shallow constriction between pronotum and elytra, light brown with slightly lighter tarsi, covered with yellowish brown setae. BL 1.13 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.15 mm, HW 0.28 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region fine and inconspicuous; setae short, sparse, suberect. Antennae long and slender, gradually and strongly thickened, AnL 0.60 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 – 7 each about as long as broad, 8 – 10 each weakly transverse, 11 slightly longer than 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, equally broad between base and middle; PL 0.35 mm, PW 0.50 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each almost equidistant to posterior and lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae on pronotal disc short and moderately dense, suberect. Elytra as wide as pronotum, together oval, broadest near anterior third; EL 0.63 mm, EW 0.50 mm, EI 1.25; subhumeral lines marked as borders between lower mesal and elevated humeral regions, only as long as about 0.2 × EL and distinctly divergent posterad; basal elytral fovea on each elytron situated at middle between lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra conspicuously distinct, those on mesal half of each elytron small but sharply marked and appearing somewhat coarse, separated by spaces smaller than puncture diameters, punctures gradually becoming smaller and shallower towards lateral and posterior elytral margins; setae forming conspicuous patter, those on densely punctate adsutural halves directed posterad and suberect, those on outer halves directed laterad and erect. Hind wings functional. Metaventrite with lateral impressions about as long as 3 / 4 of its length. Legs long and slender, unmodified. Aedeagus (Figs 128 – 129) of modified simonis form, AeL 0.20 mm; capsular region of median lobe in ventral view drop-shaped with moderately strongly elongate and conspicuously narrow distal region tapering towards pointed apex; dorsal apical projection (= distal plate) in ventral view with nearly straight distal (oblique) margin; parameres long and slender, not reaching apex of aedeagus, each with one short apical and two similarly short subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8C096FFF69FC436E67DCE5.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8C096FFF69FC436E67DCE5.taxon	etymology	Etymology. The adjective pseudopalpalis refers to the similarity to C. palpalis.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF8C096FFF69FC436E67DCE5.taxon	discussion	Remarks. Cephennodes pseudopalpalis clearly differs from C. palpalis (Fig. 133) in a much stouter body, and from C. micrus described below in less convex lateral margins of elytra. All three species have clearly different aedeagi, although representing variations of the same form.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF92096EFF69FC066CBADC55.taxon	description	(Figs 126, 130 – 132)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF92096EFF69FC066CBADC55.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Lincang Pref., / Bangma Shan, E pass, 7 km NW / Lincang, 23 º 57 ’ 31 ’’ N, 99 º 56 ’ 13 ’’ E, / 2040 m, secondary pine forest, litter, / dead wood & mushrooms sifted, / 9. IX. 2009, leg. M. Schülke [CH 09 - 36] ” [white, printed]; “ CEPHENNODES (s. str.) / micrus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF92096EFF69FC066CBADC55.taxon	diagnosis	Diagnosis. Body extremely small, BL 1 mm; antennae gradually thickened; punctures on head fine and inconspicuous, on pronotum small but distinct, sharply marked and dense, separated by spaces slightly narrower than their diameters, those on elytra dense but very shallow, superficial; subhumeral lines short, as long as 0.3 × EL. Male: lacking secondary sexual characters; aedeagus (Figs 131 – 132) in ventral view with slightly recurved and conspicuously elongate, rod-like apical region of capsular portion pointed at apex; distal margin of apical plate distinctly concave.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF92096EFF69FC066CBADC55.taxon	description	Description. Body of male (Fig. 130) strongly convex, oval, with shallow but distinct constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with yellowish brown setae. BL 1.00 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.13 mm, HW 0.25 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region fine and inconspicuous; setae short, sparse, suberect. Antennae long and slender, gradually and strongly thickened, AnL 0.55 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 – 7 each about as long as broad, 8 strongly, 9 and 10 weakly transverse, 11 slightly longer than 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, equally broad between base and middle; PL 0.33 mm, PW 0.48 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each distinctly closer to posterior than lateral pronotal margin. Punctures on pronotal disc small and shallow but sharply marked and dense, separated by spaces slightly shorter than their diameters; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest between middle and anterior third; EL 0.55 mm, EW 0.50 mm, EI 1.10; subhumeral lines marked as borders between lower mesal and elevated humeral regions, as long as about 0.3 × EL and weakly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra very shallow, superficial, slightly larger and sparser than those on pronotum; setae similar to those on pronotum. Hind wings functional. Metaventrite with lateral impressions about as long as half of its length. Legs long and slender, unmodified. Aedeagus (Figs 131 – 132) of modified simonis form, AeL 0.18 mm; capsular region of median lobe in ventral view drop-shaped with conspicuously strongly elongate, rod-like distal region largely parallel-sided except for tapered apex; dorsal apical projection (= distal plate) in ventral view with strongly concave distal (oblique) margin; parameres long and slender, not reaching apex of aedeagus, in studied specimen only one paramere is discernible, with two short apical setae and one long subapical seta. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF92096EFF69FC066CBADC55.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF92096EFF69FC066CBADC55.taxon	etymology	Etymology. Micrus is here treated as a noun in apposition; it refers to the extremely small body size.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF92096EFF69FC066CBADC55.taxon	discussion	Remarks. Cephennodes micrus is the stoutest of all members of the C. palpalis group, with most strongly convex lateral elytral margins.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF93096EFF69FC9669C6DB75.taxon	diagnosis	Diagnosis. Lack of modifications on head and other body parts in males; aedeagus of simonis form with drop-shaped or oval capsular region of median lobe, subtriangular apical projection, and dorsal wall lacking setae. This is a newly established and species-rich group that includes 17 previously described species distributed in Africa, China, Hawaii, Japan, Malaysia, Russian Far East, Taiwan, Thailand, and Vietnam:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF93096DFF69FB9368CADCB2.taxon	description	The C. simonis group may be a garbage dump for species that do not show any peculiar external characters, are small-bodied, and their aedeagi are similar in a general form. These species were previously treated as not placed in any group, but the newly proposed (not yet final) division facilitates identifications and reduces the number of unassigned species. A phylogenetic analysis is required to test the monophyly of this assemblage.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF90096BFF69FBB5696DDDE1.taxon	description	(Figs 134 – 138)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF90096BFF69FBB5696DDDE1.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan / Nujiang Lisu Aut. Pref., / Salween side valley 5 km S / Fugong, road SS 228, / km 223 (creek bank, litter sifted) / 8. VI. 2007 D. W. Wrase [25] ” [white, printed]; “ CEPHENNODES (s. str.) / planus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (NKME). Paratype: ♂, “ CHINA: Yunnan (CH 07 - 25) / Nujiang Lisu Aut. Pref., / Salween side valley 5 km S / Fugong, road SS 228, / km 223, under stones, moist / enbarkment {sic!} / 8. VI. 2007 / leg. A. Pütz ” [white, printed], and yellow PARATYPUS label (cAP).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF90096BFF69FBB5696DDDE1.taxon	diagnosis	Diagnosis. Body extremely small, BL around 1 mm and conspicuously flattened; antennae gradually thickened; punctures on head, pronotum and elytra fine and inconspicuous; subhumeral lines only as long as 0.2 × EL. Male: lacking secondary sexual characters; aedeagus (Figs 135 – 138) in ventral view with oval capsular region of median lobe, lacking defined apex, apical projections elongate; each paramere with 3 setae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF90096BFF69FBB5696DDDE1.taxon	description	Description. Body of male (Fig. 134) weakly convex, oval, lacking constriction between pronotum and elytra, moderately light brown, covered with yellowish setae. BL 1.03 – 1.05 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.13 mm, HW 0.25 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region fine and inconspicuous; setae short, sparse, suberect. Antennae moderately long and slender, gradually and strongly thickened, AnL 0.50 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 – 6 each about as long as broad, 7 slightly elongate, 8 – 10 each weakly transverse, 11 almost as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, broadest near middle but barely discernibly narrowing posterad; PL 0.30 – 0.33 mm, PW 0.45 – 0.48 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and only slightly rounded posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each much closer to posterior than lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior fourth; EL 0.60 mm, EW 0.45 mm, EI 1.33; subhumeral lines very short, as long as about 0.2 × EL but sharply marked, weakly divergent posterad; basal elytral fovea on each elytron situated in middle between lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite with indistinct lateral impressions about as long as 1 / 3 of its length. Legs long and slender, unmodified. Aedeagus (Figs 135 – 138) of simonis form, AeL 0.15 mm; capsular region of median lobe in ventral view oval without defined apex; dorsal apical projection (= distal plate) in ventral view elongate and tapering distad, in lateral view curved dorsad at nearly right angle; parameres long and slender, not reaching apex of aedeagus, each with one long apical and two long subapical setae. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF90096BFF69FBB5696DDDE1.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF90096BFF69FBB5696DDDE1.taxon	etymology	Etymology. The Latin adjective planus means flat; the body of C. planus is much less convex than in most congeners.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF90096BFF69FBB5696DDDE1.taxon	discussion	Remarks. Two examined specimens are externally identical and they have been collected on the same spot and on the same day. They slightly differ in some details of the aedeagus (holotype: Figs 135 – 136 vs. paratype: 137 – 138). In lateral views, the aedeagi are almost identical and the observed differences in ventral view may be results of small differences in orientation in mounting medium. Among Chinese species, C. planus has the aedeagus most similar to that of the previously known C. gaozhaianus and C. gracilis described below. Cephennodes gaozhaianus has a distinctly, densely punctate pronotum (in C. planus punctures are inconspicuous), and C. gracilis is more convex, the elytra are widest more posteriorly, the subhumeral lines are longer, over 0.3 × EL, the apical projections of aedeagus are of a different shape both in ventral and lateral views, and each paramere has an unusual number of four setae (three in C. planus).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF96096AFF69FD026F51DE91.taxon	description	(Figs 139 – 141)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF96096AFF69FD026F51DE91.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA [20] - Yunnan, / mt. W Gejiu, mixed / forest, 23 º 24 ’ 13 ’’ N, / 103 º 07 ’ 28 ’’ E, 1990 m, / 23. VIII. 2014, V. Assing ” [white, printed]; “ CEPHENNODES (s. str.) / gracilis m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF96096AFF69FD026F51DE91.taxon	diagnosis	Diagnosis. Body extremely small, BL around 1 mm; antennae gradually thickened; punctures on head, pronotum and elytra fine and inconspicuous; subhumeral lines as long as 0.35 × EL. Male: lacking secondary sexual characters; aedeagus (Figs 140 – 141) in ventral view with oval capsular region of median lobe, lacking defined apex, apical projections elongate; each paramere with 4 setae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF96096AFF69FD026F51DE91.taxon	description	Description. Body of male (Fig. 139) moderately strongly convex, oval, with barely marked constriction between pronotum and elytra, dark brown, covered with light brown setae. BL 1.03 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.13 mm, HW 0.26 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region fine and inconspicuous; setae short, sparse, suberect. Antennae moderately long and slender, gradually and strongly thickened, AnL 0.43 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 – 6 each about as long as broad, 7 slightly elongate, 8 – 10 each weakly transverse, 11 almost as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, equally broad between base and middle; PL 0.33 mm, PW 0.45 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each distinctly closer to posterior than lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, suberect. Elytra as wide as pronotum, together oval, broadest near anterior third; EL 0.58 mm, EW 0.45 mm, EI 1.28; subhumeral lines as long as 0.35 × EL, developed as sharp borders between elevated humerus and lower mesal region, weakly divergent posterad; basal elytral fovea on each elytron situated in middle between lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 140 – 141) of simonis form, AeL 0.15 mm; capsular region of median lobe in ventral view oval without defined apex; dorsal apical projection (= distal plate) in ventral view elongate and tapering distad, curved to the left, in lateral view curved dorsad at right angle; parameres long and relatively thick, not reaching apex of aedeagus, each with four moderately long setae in apical region. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF96096AFF69FD026F51DE91.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF96096AFF69FD026F51DE91.taxon	etymology	Etymology. The Latin adjective gracilis means slender, slim; C. gracilis has an elongate body.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF96096AFF69FD026F51DE91.taxon	discussion	Remarks. Among Chinese species, C. gracilis has the aedeagus most similar to that of the previously known C. gaozhaianus and the newly described C. planus. Cephennodes gaozhaianus has a distinctly and densely punctate pronotum (in C. gracilis and C. planus punctures are inconspicuous). Cephennodes gracilis differs from C. planus in a more convex body, elytra broadest more posteriorly and with longer subhumeral lines, and also in a different shape of apical projections of the aedeagus. Moreover, C. gracilis has four parameral setae, whereas C. planus and C. gaozhaianus have three setae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF970969FF69FDD26D84D89C.taxon	description	(Figs 142 – 144)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF970969FF69FDD26D84D89C.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Wuding, Lion / Mountain Scenic Area, 2200 m, / 25 º 31 ’ 59 ’’ N, 102 º 22 ’ 36 ’’ E, stream / valley with deciduous forest, moist / litter sifted 17. VIII. 2014, / leg. M. Schülke [CH 14 - 11] ” [white, printed]; “ CEPHENNODES (s. str.) / molestus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (5 exx.): 1 ♂, same data as for holotype; 3 ♂♂, 1 ♀, “ CHINA: Yunnan, Mt. W Xundian, / 25 º 34 ’ 58 ’’ N, 103 º 08 ’ 42 ’’ E, 2300 m, / mixed forest with alder, pine, / shrubs, litter, twigs, roots sifted, / 16. VIII. 2014, Schülke [CH 14 - 09 b] ” [white, printed]. Paratypes with yellow PARATYPUS labels (cPJ, MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF970969FF69FDD26D84D89C.taxon	diagnosis	Diagnosis. Body very small, BL around 1.1 – 1.2 mm; antennae gradually thickened; punctures on head and pronotum small but sharply marked and distinct, separated by spaces subequal to their diameters, punctures on elytra superficial; subhumeral lines as long as 0.35 × EL. Male: lacking secondary sexual characters; aedeagus (Figs 143 – 144) in ventral view with drop-shaped capsular region of median lobe, its distal portion projecting into broad subrectangular truncate process, apical projections much wider than long and curved towards apex of capsular region.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF970969FF69FDD26D84D89C.taxon	description	Description. Body of male (Fig. 142) moderately strongly convex, oval, with shallow but distinct constriction between pronotum and elytra, moderately light brown with lighter tarsi, covered with light brown setae. BL 1.08 – 1.18 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.15 mm, HW 0.28 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region small but distinct, separated by spaces subequal to their diameters; setae sparse, short, suberect. Antennae moderately long and slender, gradually and moderately strongly thickened, AnL 0.50 – 0.58 mm, antennomere 1 about as long as broad, 2 strongly elongate, 3 – 6 each about as long as broad, 7 slightly elongate, 8 – 10 each weakly transverse, 11 about as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, equally broad between base and middle; PL 0.30 – 0.35 mm, PW 0.48 – 0.50 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each distinctly closer to lateral than posterior pronotal margin. Punctures on pronotal disc small but distinct, separated by spaces subequal to their diameters; setae short and moderately dense, suberect. Elytra as wide as pronotum, together oval, broadest near anterior third; EL 0.60 – 0.68 mm, EW 0.50 – 0.56 mm, EI 1.12 – 1.25; subhumeral lines as long as 0.35 × EL, developed as sharp borders between elevated humerus and lower mesal region, weakly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra superficial and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite with sharply marked lateral impressions as long as about half its length. Legs long and slender, unmodified. Aedeagus (Figs 143 – 144) of simonis form, AeL 0.10 mm; capsular region of median lobe in ventral view drop-shaped, its distal portion projecting into broad subrectangular truncate process with weakly concave distal margin, apical projections much wider than long and curved towards apex of capsular region; parameres long and relatively thick, not reaching apex of aedeagus, each with one moderately long apical seta. Female. Externally indistinguishable from male; wings not studied. BL 1.13 mm; HL 0.15 mm, HW 0.28 mm, AnL 0.53 mm; PL 0.35 mm, PW 0.50 mm; EL 0.63 mm, EW 0.55 mm, EI 1.14.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF970969FF69FDD26D84D89C.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF970969FF69FDD26D84D89C.taxon	etymology	Etymology. The Latin adjective molestus means troublesome; this species is difficult to identify based on external characters.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF970969FF69FDD26D84D89C.taxon	discussion	Remarks. Besides C. molestus, only one species of the C. simonis group has the apical region of the capsular portion of the median lobe truncate, forming a subrectangular apical process. This is the Bornean C. simonis. In C. molestus, the lateral pronotal carinae are simple, i. e., developed as slightly thickened margins mesally accentuated by a step-wise difference in elevated margin and a lower discal surface. In C. simonis, the lateral pronotal carinae are doubled, i. e., in addition to a similarly developed margin, there is a sublateral carina running parallel and close to the margin. The shape of the aedeagus is also clearly different in these species, especially the apical plate, which in C. simonis is strongly curved and pointed in both ventral and lateral views (illustrated in Jałoszyński (2007 a): figs. 3 – 4).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF950968FF69FF676CAFD869.taxon	description	(Figs 145 – 147)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF950968FF69FF676CAFD869.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Baoshan Pref., Gao- / ligong Shan, W pass 35 km SE / Tengchong, 2100 m, 24 º 50 ’ 18 ’’ N, / 95 º 45 ’ 43 ’’ E, devast. prim. dec. forest, / litter, wood, mushrooms sifted, 25. VII. / 2009, leg. M. Schülke [CH 09 - 06] ” [white, printed]; “ CEPHENNODES (s. str.) / aliusmodi m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratype: 1 ♀, same data as for holotype, and with yellow PARATYPUS label (cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF950968FF69FF676CAFD869.taxon	diagnosis	Diagnosis. Body very small, BL around 1.1 mm; antennae gradually thickened; punctures on head, pronotum and elytra fine, inconspicuous; subhumeral lines as long as 0.3 × EL. Male: lacking secondary sexual characters; aedeagus (Figs 146 – 147) in ventral view with subtriangular capsular region of median lobe, its distal portion projecting into elongate triangular and apically angulate process, apical projection elongate, subtriangular, broadly separated from apex of capsular region.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF950968FF69FF676CAFD869.taxon	description	Description. Body of male (Fig. 145) moderately strongly convex, oval, with shallow but distinct constriction between pronotum and elytra, moderately light brown with lighter tarsi, covered with light brown setae. BL 1.08 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.13 mm, HW 0.28 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region fine and inconspicuous; setae sparse, short, suberect. Antennae moderately long and slender, gradually and moderately strongly thickened, AnL 0.53 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 – 7 each about as long as broad, 8 – 10 each weakly transverse, 11 slightly shorter than 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, equally broad between base and middle; PL 0.33 mm, PW 0.50 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each distinctly closer to lateral than posterior pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, suberect. Elytra as wide as pronotum, together oval, broadest between middle and anterior third; EL 0.63 mm, EW 0.50 mm, EI 1.25; subhumeral lines as long as about 0.3 × EL, developed as sharp borders between elevated humerus and lower mesal region, weakly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra superficial and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite with distinct lateral impressions as long as about half its length. Legs long and slender, unmodified. Aedeagus (Figs 146 – 147) of simonis form, AeL 0.15 mm; capsular region of median lobe in ventral view subtriangular, its distal portion projecting into elongate triangular process, apical projection elongate, subtriangular, with narrow distal portion, broadly separated from apex of capsular region; parameres long and slender, not reaching apex of aedeagus, each with one moderately long apical seta. Female. Externally indistinguishable from male; wings not studied. BL 1.10 mm; HL 0.15 mm, HW 0.25 mm, AnL 0.50 mm; PL 0.33 mm, PW 0.50 mm; EL 0.63 mm, EW 0.53 mm, EI 1.19.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF950968FF69FF676CAFD869.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF950968FF69FF676CAFD869.taxon	etymology	Etymology. The Latin alius - modi (modi being a noun) means “ of another kind ”.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF950968FF69FF676CAFD869.taxon	discussion	Remarks. Within the C. simonis group, C. aliusmodi is the only species with narrowly subtriangular apical projection in ventral view projecting distad further than apex of capsular region and parallel to it, but these two structures are widely separated.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9A0967FF69FF676CBBD8F9.taxon	description	(Figs 148 – 150)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9A0967FF69FF676CBBD8F9.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA mer.: Yunnan prov. / (pass SW from Baoshan / Gaoligong Shan) / 4. - 8.6.2005 / sporocarp of mycetes / Oto Nakladal lgt. ” [white, printed]; “ CEPHENNODES (s. str.) / parvus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW). Paratype: 1 ♀, same data as for holotype, and with yellow PARATYPUS label (cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9A0967FF69FF676CBBD8F9.taxon	diagnosis	Diagnosis. Body very small, BL around 1 mm; antennae gradually thickened; punctures on clypeus virtually absent, on frontovertexal region small but sharply marked and dense, slightly denser near mesal margins of eyes than at middle, on median area separated by spaces slightly wider, and near eyes slightly narrower than their diameters, pronotum and elytra with fine, inconspicuous punctures; subhumeral lines as long as 0.3 × EL. Male: lacking secondary sexual characters; aedeagus (Figs 149 – 150) in ventral view with subtriangular capsular region of median lobe, its distal portion projecting into elongate subtriangular and apically rounded process, apical projection much wider than long, subtriangular, broadly separated from apex of capsular region.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9A0967FF69FF676CBBD8F9.taxon	description	Description. Body of male (Fig. 148) moderately strongly convex, oval, with shallow but distinct constriction between pronotum and elytra, moderately light brown with lighter tarsi, covered with light brown setae. BL 1.05 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.13 mm, HW 0.25 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus virtually absent, on frontovertexal region small but sharply marked and dense, on median area separated by spaces slightly wider, and near eyes slightly narrower than their diameters; setae sparse, short, suberect. Antennae moderately long and slender, gradually and moderately strongly thickened, AnL 0.53 mm, antennomere 1 about as long as broad, 2 strongly elongate, 3 – 7 each about as long as broad, 8 – 10 each weakly transverse, 11 slightly shorter than 9 and 10 combined, about 1.6 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, equally broad between base and middle; PL 0.33 mm, PW 0.45 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to lateral and posterior pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.60 mm, EW 0.50 mm, EI 1.20; subhumeral lines as long as about 0.3 × EL, developed as sharp borders between elevated humerus and lower mesal region, weakly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra superficial and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite with distinct lateral impressions as long as about 3 / 4 of its length. Legs long and slender, unmodified. Aedeagus (Figs 149 – 150) of simonis form, AeL 0.13 mm; capsular region of median lobe in ventral view subtriangular, its distal portion projecting into elongate subtriangular and apically rounded process, apical projection much wider than long, subtriangular, broadly separated from apex of capsular region; parameres long and slender, not reaching apex of aedeagus, each with one moderately long apical seta. Female. Externally indistinguishable from male; wings not studied. BL 0.95 mm; HL 0.13 mm, HW 0.25 mm, AnL 0.50 mm; PL 0.20 mm, PW 0.48 mm; EL 0.63 mm, EW 0.50 mm, EI 1.25.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9A0967FF69FF676CBBD8F9.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9A0967FF69FF676CBBD8F9.taxon	etymology	Etymology. The Latin adjective parvus means little, small.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9A0967FF69FF676CBBD8F9.taxon	discussion	Remarks. Within the C. simonis group, C. parvus is most similar to C. aliusmodi. They both share the aedeagi with a subtriangular capsular region of median lobe, and a subtriangular apical projection broadly separated from the capsular region. In C. aliusmodi, the apical projection is much longer than that in C. parvus (elongate vs. wider than long), and exceeds the apex of capsular region, whereas in C. parvus the apex of capsular region projects beyond apex of the projection.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9B0966FF69FF676F95DD59.taxon	diagnosis	Diagnosis. Aedeagus small, of simonis form, ventral wall lacking setae; in males metatibiae unmodified and abdomen modified, with various projections or carinae on some sternites (only IV or IV and V). This group is closely allied to the C. inflatipes (metatibiae modified, abdomen unmodified) and C. spatulipes (both metatibiae and abdomen modified) groups. All species are relatively unremarkable, except for male abdominal modifications that should be examined in transparent slides, preferably in ventral view, and adults are small-bodied, typically around 1 mm in length, with extremely small aedeagi (around 0.15 mm), which are difficult to isolate and mount in a strictly ventral or lateral orientation. This is a species-rich group that includes 16 previously described species distributed predominantly in Nepal, with only one previously known Chinese species:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9B0964FF69F9A76E33DBA9.taxon	description	(Figs 151 – 154)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9B0964FF69F9A76E33DBA9.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan [CH 07 - 14], Baoshan / Pref., Gaoligong Shan, 33 km SE / Tengchong, 2100 - 2200 m, 24 º 51 ’ 22 ’’ N, / 98 º 45 ’ 36 ’’ E, decid. forest, litter, wood, / fungi sifted, 31. V. 2007, M. Schülke ” [white, printed]; “ CEPHENNODES (s. str.) / alius m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9B0964FF69F9A76E33DBA9.taxon	diagnosis	Diagnosis. Body very small, BL around 1 mm; antennae with indistinctly delimited trimerous clubs; punctures on clypeus virtually absent, on frontovertexal region small but sharply marked and dense, separated by spaces subequal to their diameters, pronotum and elytra with fine, inconspicuous punctures; subhumeral lines as long as about 0.4 × EL. Male: abdomen (Fig. 152) with only sternite V modified, with median transverse carina; aedeagus (Figs 153 – 154) in ventral view with subtriangular capsular region of median lobe, its distal portion projecting into elongate subtriangular and apically rounded process, apical projection subtriangular, separated from distal region of capsular portion, with narrow subtriangular apex and distal margin oblique in relation to long axis of aedeagus at ~ 45 °.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9B0964FF69F9A76E33DBA9.taxon	description	Description. Body of male (Fig. 151) strongly convex, oval, with distinct constriction between pronotum and elytra, moderately light brown with lighter tarsi, covered with light brown setae. BL 1.05 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.13 mm, HW 0.25 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus virtually absent, on frontovertexal region small but sharply marked and dense, separated by spaces subequal to their diameters; setae sparse, short, suberect. Antennae moderately long and slender, with indistinctly delimited trimerous clubs, AnL 0.53 mm, antennomere 1 about as long as broad, 2 strongly elongate, 3 – 7 each about as long as broad, 8 – 10 each weakly transverse, 11 slightly shorter than 9 and 10 combined, about 1.6 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, broadest near middle and distinctly narrowing posterad; PL 0.33 mm, PW 0.50 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and weakly so posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each slightly closer to posterior than lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.60 mm, EW 0.53 mm, EI 1.14; subhumeral lines as long as about 0.4 × EL, developed as sharp borders between elevated humerus and lower mesal region, strongly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra slightly more distinct than those on pronotum but still fine and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Abdomen (Fig. 152) with short transverse median carina on sternite V. Aedeagus (Figs 153 – 154) of simonis form, AeL 0.15 mm; median lobe in ventral view with subtriangular capsular region, its distal portion projecting into elongate subtriangular and apically rounded process, apical projection subtriangular, separated from distal region of capsular portion, with narrow subtriangular apex and distal margin oblique in relation to long axis of aedeagus at ~ 45 °; parameres long and slender, one nearly reaching apex of aedeagus, each with one moderately long subapical seta. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9B0964FF69F9A76E33DBA9.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9B0964FF69F9A76E33DBA9.taxon	etymology	Etymology. The Latin adjective alius means another, different.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9B0964FF69F9A76E33DBA9.taxon	discussion	Remarks. Among the Chinese members of the C. simplicipes group, only males of C. alius have one abdominal sternite (V) with transverse carina, whereas in the previously known C. simplicipes and the newly described C. nubis two sternites (IV and V) are modified. The aedeagus of C. simplicipes differs strongly from those of C. alius and C. nubis in the apical plate being adpressed against the distal region of the capsular portion of median lobe, with its distal margin transverse in relation to the long axis of the aedeagus. In C. alius and C. nubis, the apical plate is separated from the distal region of the capsular portion, and its distal margin is strongly oblique to the long axis of the aedeagus (in C. alius at an angle of ~ 45 °, in C. nubis much larger than 45 °). The remaining species of this group are known to occur in the Himalayas, and three of them have only the sternite V with a single transverse carina: C. tharepatianus, C. minisulcatus, and C. bahrabisensis. The modifications of abdomen in these species have different shapes than those in C. alius and C. nubis, and all of them have the apical plate of the aedeagus adpressed to the capsular region of the median lobe, not separated, as in C. alius.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF990962FF69FACA6816DA15.taxon	description	(Figs 155 – 160)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF990962FF69FACA6816DA15.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Nujiang Lisu Pref., / Gaoligong Shan, “ Cloud pass ”, / 21 km NW Liuku, 25 º 58 ’ 11 ’’ N, / 98 º 41 ’ 01 ’’ E, 3150 m, shrubs & / bamboo, litter sifted, 3. IX. 2009, / leg. M. Schülke [CH 09 - 22 a] ” [white, printed]; “ CEPHENNODES (s. str.) / nubis m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (4 exx.): 2 ♂♂, 1 ♀, same data as for holotype; 1 ♀, “ CHINA: Yunnan, Nujiang Lisu Pref., / Gaoligong Shan, “ Cloud pass ”, / 21 km NW Liuku, 25 º 58 ’ 11 ’’ N, / 98 º 41 ’ 01 ’’ E, 3150 m, shrubs & / bamboo, litter sifted, 3. IX. 2009, / leg. M. Schülke [CH 09 - 22 a] ” [white, printed]. Paratypes with yellow PARATYPUS labels (cPJ, MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF990962FF69FACA6816DA15.taxon	diagnosis	Diagnosis. Body small, BL around 1.3 mm; antennae with indistinctly delimited trimerous clubs; punctures on clypeus virtually absent, on frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters, pronotum and elytra with fine, inconspicuous punctures; subhumeral lines as long as about 0.3 × EL. Male: abdomen (Fig. 156) with sternites IV and V modified, each with median transverse carina, that on sternite IV shorter and weakly projecting posterad, that on sternite V longer and more strongly projecting posterad; aedeagus (Figs 157 – 160) in ventral view with subtriangular capsular region of median lobe, its distal portion projecting into elongate subtriangular and apically rounded process, apical projection subtriangular, separated from distal region of capsular portion, with narrow subtriangular apex and distal margin oblique in relation to long axis of aedeagus at angle much larger than 45 °.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF990962FF69FACA6816DA15.taxon	description	Description. Body of male (Fig. 155) strongly convex, oval, with distinct constriction between pronotum and elytra, moderately dark brown with lighter tarsi, covered with light brown setae. BL 1.28 – 29 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.15 mm, HW 0.30 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus virtually absent, on frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters; setae sparse, short, suberect. Antennae moderately long and slender, with indistinctly delimited trimerous clubs, AnL 0.63 – 0.65 mm, antennomere 1 indistinctly elongate, 2 strongly elongate, 3 about as long as broad, 4 – 6 each weakly transverse, 7 about as long as broad, 8 – 10 each weakly transverse, 11 almost as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, broadest near middle and distinctly narrowing posterad; PL 0.38 – 0.40 mm, PW 0.60 – 0.63 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and less strongly posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each distinctly closer to posterior than lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.73 – 0.75 mm, EW 0.63 – 0.65 mm, EI 1.13 – 1.16; subhumeral lines as long as about 0.3 × EL, developed as sharp borders between elevated humerus and lower mesal region, strongly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite with lateral impressions as long as about 3 / 4 of its length. Legs long and slender, unmodified. Abdomen (Fig. 156) with sternite V bearing short transverse median carina slightly convex posteriorly in median region; sternite VI with much longer transverse carina in median region distinctly angulate and projecting posterad. Aedeagus (Figs 157 – 160) of simonis form, AeL 0.15 mm; median lobe in ventral view with subtriangular capsular region, its distal portion projecting into elongate subtriangular and apically rounded process, apical projection subtriangular, separated from distal region of capsular portion, with narrow subtriangular apex and distal margin oblique in relation to long axis of aedeagus at angle much larger than 45 °; parameres long and slender, both nearly reaching apex of aedeagus, each with one short apical seta. Female. Externally similar to male but with unmodified abdomen, wingless. BL 1.30 – 1.33 mm; HL 0.15 mm, HW 0.30 – 0.33 mm, AnL 0.63 mm; PL 0.40 – 0.43 mm, PW 0.63 – 0.65 mm; EL 0.75 mm, EW 0.65 mm, EI 1.15.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF990962FF69FACA6816DA15.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF990962FF69FACA6816DA15.taxon	etymology	Etymology. The Latin noun nubis means a cloud and alludes to the Cloud Pass given on labels.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF990962FF69FACA6816DA15.taxon	discussion	Remarks. A slight variability in the shape of the aedeagus was observed in the studied sample (Figs 157 – 160), but differences are small and external characters, including the modifications of sternites IV and V, are the same in all specimens. Among Chinese species, C. nubis can be identified by a combination of the two modified sternites (as in C. simplicipes, but unlike C. alius), and the apical plate separated from the capsular region of median lobe (as in C. alius, but unlike C. simplicipes). Among those Himalayan species that share with C. nubis two modified abdominal sternites, all have different shapes of these modifications, and only C. bilobatus has a very similar aedeagus. The latter species, however, has the transverse carinae on sternites IV and V developed as broadly subtriangular lobes projecting posterad (especially that on sternite V), whereas carinae in C. nubis are weakly projecting posterad. Moreover, C. bilobatus is clearly smaller than C. nubis, with BL respectively 1.08 mm and 1.28 – 1.33 mm.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9F0962FF69FA566E1FD845.taxon	diagnosis	Diagnosis. Aedeagus small, of simonis form, ventral wall lacking setae; in males metatibiae modified, distally broadened, and also abdomen modified, with various projections or carinae on some sternites (only IV or IV and V). This group is closely allied to the C. inflatipes (metatibiae modified, abdomen unmodified) and C. simplicipes (metatibiae unmodified, abdomen modified) groups. As in the C. simplicipes group, adults are unremarkable, except for male abdominal modifications, and beetles are small-bodied with extremely small aedeagi. This is a moderately large group that includes 12 previously described species distributed predominantly in Nepal, with four species previously known to occur in China:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9C091FFF69FB6E6949DDE9.taxon	description	(Figs 161 – 164)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9C091FFF69FB6E6949DDE9.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan [CH 07 - 03], / Dali Bai Auton. Pref., Diancang Shan W / Dali, 25 º 41 ’ 49 ’’ N, 100 º 06 ’ 24 ’’ E, / 2970 m, sifted at rock edges and under / small shrubs, 28. V. 2007, / M. Schülke ” [white, printed]; “ CEPHENNODES (s. str.) / posterotibialis m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratype: 1 ♀, same data as for holotype, with yellow PARATYPUS label (cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9C091FFF69FB6E6949DDE9.taxon	diagnosis	Diagnosis. Body small, BL around 1.3 mm; antennae gradually thickened; punctures on clypeus virtually absent, on frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters, pronotum and elytra with fine, inconspicuous punctures; subhumeral lines as long as about 0.2 × EL. Male: abdomen (Fig. 162) with sternite V modified, bearing median subtriangular transverse carina; metatibia (Fig. 161) strongly thickened in subapical region; aedeagus (Figs 163 – 164) in ventral view broadest in sub-basal region, apex of capsular region of median lobe subtriangular, distally rounded, apical projection strongly recurved, with narrowly rounded apex situated far from long axis of aedeagus and close to apex of capsular region of median lobe.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9C091FFF69FB6E6949DDE9.taxon	description	Description. Body of male (Fig. 161) strongly convex, oval, with distinct constriction between pronotum and elytra, moderately dark brown with lighter tarsi, covered with light brown setae. BL 1.30 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.18 mm, HW 0.30 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus virtually absent, on frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters; setae sparse, short, suberect. Antennae moderately long and slender, gradually thickened, AnL 0.73 mm, antennomere 1 indistinctly elongate, 2 strongly elongate, 3 – 8 each weakly elongate, 9 – 10 each weakly transverse, 11 as long as 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, broadest near middle and distinctly narrowing posterad; PL 0.38 mm, PW 0.60 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and less strongly posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each slightly closer to posterior than lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.75 mm, EW 0.63 mm, EI 1.20; subhumeral lines as long as about 0.2 × EL, developed as sharp borders between elevated humerus and lower mesal region, strongly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite with lateral impressions as long as about half of its length. Legs long and slender, metatibiae with wide subtriangular outer lateral subapical expansion (Fig. 161). Abdomen (Fig. 162) with sternite V bearing short transverse subtriangular median carina. Aedeagus (Figs 163 – 164) of simonis form, AeL 0.15 mm; median lobe in ventral view broadest in sub-basal region, apex of capsular region of median lobe subtriangular, distally rounded, apical projection strongly recurved, with narrowly rounded apex situated far from long axis of aedeagus and close to apex of capsular region of median lobe; parameres long and slender, both nearly reaching apex of aedeagus, each with one short apical and one short subapical seta. Female. Externally similar to male but with unmodified metatibiae and abdomen, wings not studied. BL 1.30 mm; HL 0.18 mm, HW 0.30 mm, AnL 0.68 mm; PL 0.38 mm, PW 0.60 mm; EL 0.75 mm, EW 0.63 mm, EI 1.20.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9C091FFF69FB6E6949DDE9.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9C091FFF69FB6E6949DDE9.taxon	etymology	Etymology. The adjective posterotibialis refers to the modified hind tibiae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FF9C091FFF69FB6E6949DDE9.taxon	discussion	Remarks. Among Chinese species, C. posterotibialis can be identified by a combination of the following characters: in ventral view aedeagus broadest in sub-basal region, and apex of capsular region of median lobe subtriangular; apical projection strongly recurved, with narrowly rounded apex situated far from long axis of aedeagus and close to apex of capsular region of median lobe. Among the Himalayan species, C. dolakhanus, C. manangensis, C. monolaminatus, and C. thakanus have a modified sternite V and unmodified IV, as in C. posterotibialis. All these four species have clearly different shapes of the apical plates of the median lobe.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE2091DFF69FD0A69C8DE24.taxon	description	(Figs 165 – 168)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE2091DFF69FD0A69C8DE24.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA mer.: Yunnan prov. / (pass SW from Baoshan / Gaoligong Shan) / 4. - 8.6.2005 / sifted from base of old tree / Oto Nakladal lgt. ” [white, printed]; “ CEPHENNODES (s. str.) / dilatatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE2091DFF69FD0A69C8DE24.taxon	diagnosis	Diagnosis. Body small, BL around 1.2 mm; antennae gradually thickened; punctures on clypeus and frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters, pronotum with slightly larger, more distinct and similarly dense punctures, elytra with punctures similar to those on head; subhumeral lines as long as about 0.2 × EL. Male: abdomen (Fig. 166) with sternite V modified, bearing conspicuously broad median subtriangular transverse carina; metatibia (Fig. 165) strongly thickened in subapical region; aedeagus (Figs 167 – 168) in ventral view broadest in subapical region, apical projection oblique in relation to long axis of median lobe, its apex not hidden behind apex of capsular region of median lobe but adpressed against it.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE2091DFF69FD0A69C8DE24.taxon	description	Description. Body of male (Fig. 165) strongly convex, oval, with indistinct constriction between pronotum and elytra, moderately dark brown with lighter tarsi, covered with light brown setae. BL 1.20 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.18 mm, HW 0.28 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region very small but sharply marked and dense, separated by spaces subequal to their diameters; setae sparse, short, suberect. Antennae moderately long and slender, gradually thickened, AnL 0.60 mm, antennomere 1 about as long as broad, 2 strongly elongate, 3 distinctly transverse, 4 – 7 each about as long as broad, 8 – 10 each weakly transverse, 11 as long as 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, broadest near middle and distinctly narrowing posterad; PL 0.35 mm, PW 0.50 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and less strongly posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each slightly closer to lateral than posterior pronotal margin. Punctures on pronotal disc slightly larger and deeper than those on head, distinct, separated by spaces subequal to their diameters; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.68 mm, EW 0.55 mm, EI 1.23; subhumeral lines as long as about 0.2 × EL, developed as sharp borders between elevated humerus and lower mesal region, strongly divergent posterad; basal elytral fovea on each elytron situated slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra similar to those on head, shallower and slightly sparser than those on pronotum; setae similar to those on pronotum. Hind wings not studied. Metaventrite with lateral impressions as long as about half of its length. Legs long and slender, metatibiae with wide subtriangular outer lateral subapical expansion (Fig. 165). Abdomen (Fig. 166) with sternite V bearing long transverse subtriangular median carina. Aedeagus (Figs 167 – 168) of simonis form, AeL 0.14 mm; median lobe in ventral view broadest in subapical region, apical projection oblique in relation to long axis of median lobe, its apex not hidden behind apex of capsular region of median lobe but adpressed against it; parameres long and slender, one nearly reaching apex of aedeagus, each with one short apical seta. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE2091DFF69FD0A69C8DE24.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE2091DFF69FD0A69C8DE24.taxon	etymology	Etymology. The adjective dilatatus refers to the dilated hind tibiae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE2091DFF69FD0A69C8DE24.taxon	discussion	Remarks. Among Chinese species, C. dilatatus can be identified by a combination of the following characters: aedeagus in ventral view broadest in subapical region, apical projection oblique in relation to long axis of median lobe, its apex not hidden behind apex of capsular region of median lobe but adpressed against it. Among the Himalayan species, C. dolakhanus, C. manangensis, C. monolaminatus, and C. thakanus have a modified sternite V and unmodified IV, as in C. dilatatus. Cephennodes dolakhanus and C. thakanus have clearly different shapes of the apical plates of the median lobe. Cephennodes manangensis and C. monolaminatus have the aedeagi most similar to that of C. dilatatus. Cephennodes manangensis has a different pattern of punctures on body parts: on the pronotal disc punctures are much smaller and shallower than those on the head, but denser, and on elytra slightly more distinct than those on pronotum, whereas in C. dilatatus the pronotum is covered with punctures slightly larger, more distinct and similarly dense as those on the frontovertexal region, and elytra with punctures similar to those on the head. Cephennodes monolaminatus is distinctly larger than C. dilatatus, body length respectively 1.45 – 1.58 mm and 1.20 mm, and has the pronotal disc covered with fine and inconspicuous punctures.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE0091DFF69FE4468C2DDAD.taxon	diagnosis	Diagnosis. Head in males with pair of long bristles with thickened apices inserted on vertexal region and directed anteriorly and dorsally; aedeagus of simonis form, with short setae on subapical area of ventral wall; frontoclypeal area in most species modified, with impressions and / or convexities. This group is closely related to the C. impressifrons (head lacking a pair of thickened vertexal setae) and C. bos (head lacking a pair of thickened setae; antennomere 3 modified) groups. This is a moderately large group that includes 12 previously described species (some of them placed in this group in the present study) distributed predominantly in China, but known also from Taiwan, Japan, and Vietnam:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE0091CFF69FCEA68CADDFC.taxon	description	Three species described by Hideto Hoshina, C. akane (Taiwan), C. maya (Taiwan), and C. yume (Japan: Okinawa Is.) cannot be identified based on the original descriptions and illustrations. Judging from highly inadequate figures, C. yume and C. akane can be synonyms. The illustrations, or rather rough sketches of the aedeagus of C. maya are even more problematic. I assign this species to the C. taurus group based on the Hoshina’s (2021) description of the male characters: “ Head impressed in anterior part of vertex and bearing a pair of a long upstanding bristlelike structures on posterior part of vertex. ” Hoshina stated that C. maya is similar to C. hongkongensis, which clearly demonstrates that he was not familiar with the division of Cephennodes into species groups, because C. hongongensis is not a member of the C. taurus group.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE1091AFF69FCFF6EFDDCDF.taxon	description	(Figs 169 – 174)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE1091AFF69FCFF6EFDDCDF.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA [13] - Yunnan, / mt. SE Gejiu, graveyard / with pine, 23 º 18 ’ 27 ’’ N, / 103 º 11 ’ 41 ’’ E, 2400 m, / 20. VIII. 2014, V. Assing ” [white, printed]; “ CEPHENNODES (s. str.) / frontorhinus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNHW). Paratype: 1 ♀, same data as for holotype, and with yellow “ PARATYPUS ” label (cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE1091AFF69FCFF6EFDDCDF.taxon	diagnosis	Diagnosis. Body large, BL around 1.6 – 1.7 mm; antennae gradually thickened; punctures on clypeus, pronotum and elytra small but sharply marked and dense, separated by spaces subequal to their diameters, those on elytra slightly larger than on pronotum, head posterior to frontal ‘ horn’ virtually impunctate; subhumeral lines short, as long as 0.3 × EL. Male: head modified (Figs 170 – 172), with subtriangular median anterior projection (‘ horn’), its anterior margins at each side of apex nearly straight, large median area posterior to ‘ horn’ asetose except for long median longitudinal line composed of minute densely set setiferous punctures, this median area is flanked by densely setose lateral regions with long erect setae directed anterad (Figs 171, 172); aedeagus (Figs 173 – 174) strongly elongate, in ventral view with distal margin of capsular region forming abruptly narrowed elongate sharp-angled subtriangular projection.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE1091AFF69FCFF6EFDDCDF.taxon	description	Description. Body of male (Fig. 169) strongly convex, oval, with barely discernible constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.66 mm. Head (Figs 170 – 172) broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.26 mm, HW 0.43 mm; frontovertexal region modified, with subtriangular median anterior projection, its anterior margins at each side of apex nearly straight, large median area posterior to projection asetose except for long median longitudinal line composed of minute densely set setiferous punctures, this median area is flanked by densely setose lateral regions with long erect setae directed anterad, so that pair of thickened vertexal setae is obscured by long setae. Punctures on clypeus small but sharply marked, separated by spaces subequal to their diameters, frontovertexal region posterior to subtriangular projection virtually impunctate. Antennae long and slender, gradually thickened, AnL 1.00 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 about as long as broad, 4 – 7 each weakly elongate, 8 about as long as broad, 9 and 10 each indistinctly elongate, 11 as long as 9 and 10 combined, about twice as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, broadest slightly behind middle but indistinctly narrowing posterad; PL 0.53 mm, PW 0.78 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and feebly so posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc small but sharply marked, separated by spaces subequal to their diameters, punctures near posterior angles distinctly denser and somewhat coarse. Setae on pronotal disc long and moderately dense, suberect. Elytra as wide as pronotum, together oval, broadest near anterior third; EL 0.88 mm, EW 0.78 mm, EI 1.13; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron about equidistant from lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra slightly larger than those on pronotum and similarly dense; setae slightly longer than those on pronotum, dense and suberect. Hind wings reduced, about as long as elytra and narrow. Metaventrite with diffuse lateral impressions about as long as 1 / 3 of its length. Legs long and slender, unmodified. Aedeagus (Figs 173 – 174) of modified simonis form, AeL 0.40 mm; in ventral view with distal margin of capsular region forming abruptly narrowed elongate sharp-angled subtriangular projection; apical projections about as long as broad, subtriangular; parameres long and slender, not reaching apex of aedeagus, each with one long apical and two similarly long subapical setae. Female. Externally similar to male, except for unmodified head and slightly shorter antennae in relation to BL; wings not studied; BL 1.70 mm; HL 0.25 mm, HW 0.43 mm, AnL 0.88 mm; PL 0.55 mm, PW 0.80 mm; EL 0.90 mm, EW 0.80 mm, EI 1.13.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE1091AFF69FCFF6EFDDCDF.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE1091AFF69FCFF6EFDDCDF.taxon	etymology	Etymology. The adjective frontorhinus refers to the frontal ‘ horn’ on the head in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE1091AFF69FCFF6EFDDCDF.taxon	discussion	Remarks. Among Chinese species of the C. taurus group, C. frontorhinus is unique in having the following combination of characters: the frontal region of head in dorsal view with an anteromedian subtriangular convexity projecting anterad; lateral margins of this convexity nearly straight; and the apex of the capsular region of median lobe subtriangular, sharp-angled and strongly projecting. This species is most similar to C. unicornis described below. In addition to a different shape of the frontal subtriangular projection (with its lateral margins nearly straight in C. frontorhinus, and sinuate in C. unicornis), the apex of the capsular region of the median lobe in ventral view in C. frontorhinus is forming an abruptly narrowing elongate subtriangular sharp-angled projection, whereas in C. unicornis the apex is obtuse-angled and not delimited from lateral and distal margins. See also Remarks for C. unicornis. The same combination of characters differentiates C. frontorhinus from its Vietnamese congeners. Unfortunately, C. akane (Taiwan), C. maya (Taiwan), and C. yume (Japan: Okinawa Is.), all described by Hideto Hoshina, cannot be identified based on the original descriptions and illustrations. However, even accounting for some inaccuracy of Hoshina’s drawings, the aedeagi of C. frontorhinus and C. unicornis are not similar enough to those of C. akane, C. maya or C. yume to pose a risk of conspecificity.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE70918FF69FC006C1FDBE1.taxon	description	(Figs 175 – 180)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE70918FF69FC006C1FDBE1.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Wuding, Lion / Mountain Scenic Area, 2200 m, / 25 º 31 ’ 59 ’’ N, 102 º 22 ’ 36 ’’ E, stream / valley with deciduous forest, moist / litter sifted 17. VIII. 2014, / leg. M. Schülke [CH 14 - 11] ” [white, printed]; “ CEPHENNODES (s. str.) / unicornis m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (7 exx.): 2 ♂♂, 5 ♀♀, same data as for holotype, each with yellow “ PARATYPUS ” label (cPJ, MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE70918FF69FC006C1FDBE1.taxon	diagnosis	Diagnosis. Body large, BL around 1.6 – 1.7 mm; antennae gradually thickened; punctures on clypeus, pronotum and elytra small but sharply marked and dense, separated by spaces subequal to their diameters, those on elytra slightly larger than on pronotum, head posterior to frontal ‘ horn’ virtually impunctate; subhumeral lines short, as long as 0.3 × EL. Male: head modified (Figs 176 – 178), with subtriangular median anterior projection (‘ horn’), its anterior margins at each side of apex sinuate, large median area posterior to ‘ horn’ asetose except for long median longitudinal line composed of minute densely set setiferous punctures, this median area is flanked by densely setose lateral regions with long erect setae directed anterad (Fig. 178); aedeagus (Figs 179 – 180) relatively stout, in ventral view with distal margin of capsular region subtriangular, obtuse-angled.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE70918FF69FC006C1FDBE1.taxon	description	Description. Body of male (Fig. 175) strongly convex, oval, with barely discernible constriction between pronotum and elytra, moderately to strongly dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.55 – 1.63 mm. Head (Figs 176 – 178) broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.25 mm, HW 0.40 – 0.43 mm; frontovertexal region modified, with subtriangular median anterior projection, its anterior margins at each side of apex nearly straight, large median area posterior to projection asetose except for long median longitudinal line composed of minute densely set setiferous punctures, this median area is flanked by densely setose lateral regions with long erect setae directed anterad, so that pair of thickened vertexal setae is obscured by long setae. Punctures on clypeus small but sharply marked, separated by spaces subequal to their diameters, frontovertexal region posterior to subtriangular projection virtually impunctate. Antennae long and slender, gradually thickened, AnL 0.98 – 1.00 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 about as long as broad, 4 – 7 each weakly elongate, 8 about as long as broad, 9 and 10 each indistinctly elongate, 11 as long as 9 and 10 combined, about twice as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near hind corners, broadest slightly behind middle but indistinctly narrowing posterad; PL 0.50 mm, PW 0.70 – 0.78 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and feebly so posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc small but sharply marked, separated by spaces subequal to their diameters, punctures near posterior angles distinctly denser and somewhat coarse. Setae on pronotal disc long and moderately dense, suberect. Elytra as wide as pronotum, together oval, broadest near anterior third; EL 0.80 – 0.88 mm, EW 0.70 – 0.78 mm, EI 1.10 – 1.13; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron about equidistant from lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra slightly larger than those on pronotum and similarly dense; setae slightly longer than those on pronotum, dense and suberect. Hind wings lacking. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 179 – 180) of modified simonis form, AeL 0.40 mm; in ventral view with distal margin of capsular region subtriangular, obtuse-angled; parameres long and slender, not reaching apex of aedeagus, each with one long apical and two similarly long subapical setae. Female. Externally similar to male, except for unmodified head and slightly shorter antennae in relation to BL; wingless; BL 1.63 – 1.65 mm; HL 0.25 mm, HW 0.41 – 0.43 mm, AnL 0.85 – 0.88 mm; PL 0.50 – 0.53 mm, PW 0.73 – 0.83 mm; EL 0.88 – 0.90 mm, EW 0.78 – 0.80 mm, EI 1.09 – 1.16.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE70918FF69FC006C1FDBE1.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE70918FF69FC006C1FDBE1.taxon	etymology	Etymology. The adjective unicornis refers to the frontal ‘ horn’ on the head in males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE70918FF69FC006C1FDBE1.taxon	discussion	Remarks. Cephennodes unicornis is very similar to C. frontorhinus, and I base my decision to treat them as separate species mainly on the strong difference in the apex of the capsular region of median lobe. This hypothesis should be verified by studying a larger sample of both ‘ morphospecies’ to check whether the difference in the aedeagi is stable or forms a morphocline (and in the latter case the species should be merged). See also Remarks for C. frontorhinus.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE50918FF69FB0268CAD80E.taxon	diagnosis	Diagnosis. Male lacking sexual dimorphic characters; aedeagus of modified simonis form, in ventral view capsular region of median lobe symmetrical or weakly asymmetrical, apical projections elongate and asymmetrical, in lateral view median lobe with deep indentation distad basal foramen delimiting subtriangular projection, apex of ventral apical plate curved ventrad. This newly established group includes two Chinese species: the previously known C. yunnanensis Jałoszyński, 2007 a and one newly described species, from the same province. Both are externally unremarkable and very similar to each other, but their aedeagi are oddly shaped and unique, so that it is easy to identify males.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEA0916FF69FF676CDDDAD6.taxon	description	(Figs 181 – 183)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEA0916FF69FF676CDDDAD6.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan [CH 07 - 19], Dehong Dai / Aut. Pref., mountain range 31 km E Luxi, / 2280 m, 24 º 29 ’ 31 ’’ N, 98 º 52 ’ 58 ’’ E, / second. pine forest with old decid. trees, / litter sifted, 3. VI. 2007, M. Schülke ” [white, printed]; “ CEPHENNODES (s. str.) / dehongdaianus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (18 exx.): 8 ♂♂, 3 ♀♀, same data as for holotype; 1 ♂, “ CHINA: Yunnan [CH 07 - 11], Baoshan / Pref., Gaoligong Shan, nr. Xiaoheishan / N. R., 35 km SE Tengchong, 2110 m, / 24 º 50 ’ 16 ’’ N, 98 º 45 ’ 43 ’’ E, decid. forest, / litter, sifted, 30. V. 2007, M. Schülke ”; 1 ♂, “ CHINA: Yunnan [CH 07 - 19], / Dehong Dai Aut. Pref., mountain / range 31 km E Luxi, 2280 m, / 24 º 29 ’ 31 ’’ N, 98 º 52 ’ 58 ’’ E, second. / pine forest with old decid. trees, / litter sifted, 3. VI. 2007, A. Pütz ”; 1 ♂, 1 ♀, “ CHINA: Yunnan [CH 07 - 11], / Baoshan Pref., Gaoligong Shan, / nr. Xiaoheishan N. R., 35 km SE / Tengchong, 2110 m, / 24 º 50 ’ 16 ’’ N, 98 º 45 ’ 43 ’’ E, decid. / forest, litter, sifted, 30. V. 2007, / leg. A. Pütz ”; 2 ♂♂, “ CHINA: Yunnan [CH 07 - 20], / Nujiang Lisu Aut. Pref., creek / valley 3 km SE Gongshan, / 1450 - 1500 m, 27 º 43 ’ 02 ’’ N, / 98 º 41 ’ 27 ’’ E, litter, moss, sifted, / 5. VI. 2007, leg. A. Pütz ”; 1 ♂, “ CHINA (Yunnan) / Baoshan Pref., Gaoligong Shan / nr. Xianheishan N. R., 35 km SE / Tangchong, 2110 m / 24 º 50 ’ 16 ’’ N, 98 º 45 ’ 43 ’’ E / (prim. decid. forest, litter, sifted) / 30. V. / 4. VI. 2007 D. W. Wrase [11] ”. Paratypes each with yellow “ PARATYPUS ” label (cAP, cPJ, MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEA0916FF69FF676CDDDAD6.taxon	diagnosis	Diagnosis. Body small, BL around 1.0 – 1.2 mm; antennae gradually thickened; punctures on clypeus small but sharply marked, on frontovertexal region virtually lacking, on pronotum and elytra small but sharply marked and dense, separated by spaces subequal to their diameters, those on elytra slightly larger than on pronotum; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 182 – 183) in ventral view with capsular region of median lobe conspicuously small, as wide as long and truncate distally, in lateral view dorsal apical projection massive and curved ventrad, and dorsal apical projection also thick but straight, parameres of conspicuously uneven lengths.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEA0916FF69FF676CDDDAD6.taxon	description	Description. Body of male (Fig. 181) moderately strongly convex, oval, with barely discernible constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.05 – 1.19 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.15 mm, HW 0.28 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus small but sharply marked, separated by spaces subequal to their diameters, frontovertexal region virtually impunctate; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.58 – 0.65 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 about as long as broad, 4 – 7 each weakly elongate, 8 transverse, 9 and 10 each about as long as broad, 11 almost as long as 9 and 10 combined, about 1.6 × as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, equally broad between base and anterior third; PL 0.33 – 0.35 mm, PW 0.48 – 0.53 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located much closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc small but sharply marked, separated by spaces subequal to their diameters. Setae on pronotal disc long and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest between middle and anterior third; EL 0.58 – 0.68 mm, EW 0.48 – 0.55 mm, EI 1.14 – 1.29; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron distinctly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra slightly larger than those on pronotum and similarly dense; setae short, moderately dense and suberect. Hind wings in some specimens functional, in others lacking. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 182 – 183) of modified simonis form, AeL 0.20 mm; in ventral view with capsular region of median lobe conspicuously small, as wide as long and truncate distally, in ventral view apical projections strongly elongate and strongly asymmetrical, in lateral view dorsal apical projection massive and curved ventrad, and dorsal apical projection also thick but straight; parameres of conspicuously uneven lengths, each with one moderately long apical and two slightly longer subapical setae. Female. Externally indistinguishable from male; wingless; BL 1.15 – 1.18 mm; HL 0.15 mm, HW 0.28 mm, AnL 0.58 – 0.60 mm; PL 0.35 – 0.38 mm, PW 0.53 – 0.58 mm; EL 0.65 mm, EW 0.55 mm, EI 1.18.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEA0916FF69FF676CDDDAD6.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEA0916FF69FF676CDDDAD6.taxon	etymology	Etymology. The adjective dehongdaianus refers to the Dehong Dai and Jingpo Autonomous Prefecture, where the holotype was found.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEA0916FF69FF676CDDDAD6.taxon	discussion	Remarks. Cephennodes dehongdaianus can be easily identified by its unique aedeagus, impossible to confuse with any other species, but externally adults are very similar to C. yunnanensis (Fig. 184), the latter species having a slightly larger body.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEB0915FF69FA17699ADE5D.taxon	description	(Fig. 184)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEB0915FF69FA17699ADE5D.taxon	materials_examined	Material studied. CHINA (YUNNAN Prov.): 1 ♂, Yunnan, Lincang Pref., Xue Shan, 11 km ENE Lincang, 2510 m, 23 º 55 ’ 01 ’’ N, 100 º 11 ’ 17.5 ’’ E, second. pine forest with Rhododendron, small cleft with water, litter & mushrooms sifted, 10. IX. 2009, leg. M. Schülke (MNB); 1 ♂, Yunnan, Baoshan Pref., Gaoligong Shan, 29 km ESE Tengchong, 2350 m, 24 º 55 ’ 37 ’’ N, 98 º 45 ’ 09 ’’ E, devastated deciduous forest, litter, wood and fungi sifted, 1. VI. 2007, / leg. A. Pütz (cAP).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEB0915FF69FA17699ADE5D.taxon	discussion	Remarks. Cephennodes yunnanensis was described based on specimens collected in Dabei, Gaoligongshan Mts., Tengchong X, at 2430 m. Species of Cephennodes (s. str.) not placed in groups After a reorganization of Cephennodes, only 14 previously described species of Cephennodes (s. str.) will remain not assigned (Jałoszyński, in prep.). Among them there are six species known to occur in China:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE80913FF69F9F96C63DC79.taxon	description	(Figs 185 – 188)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE80913FF69F9F96C63DC79.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Dali Bai Aut. Pref., / mount. range E Weishan, 12 km NE / Weishan, 25 º 17 ’ 02 - 15 ’’ N, 100 º 22 ’ / 23 - 30 ’’ E, 2630 - 2660 m, scrub with / pines and bamboo, litter sifted, 15. IX. / 2009, leg. M. Schülke [CH 09 - 54] ” [white, printed]; “ CEPHENNODES (s. str.) / arcuatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE80913FF69F9F96C63DC79.taxon	diagnosis	Diagnosis. Body small, BL around 1.15 mm; antennae gradually thickened; punctures on head, pronotum and elytra barely discernible, inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: subapical elytral region modified (Fig. 186), with cordiform flattening covered with flattened hyaline setae; aedeagus (Figs 187 – 188) in ventral view with capsular region of median lobe with elongate subtriangular distal portion rounded at apex and slender recurved distal projection with hook-like apex.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE80913FF69F9F96C63DC79.taxon	description	Description. Body of male (Fig. 185) moderately strongly convex, oval, with barely discernible constriction between pronotum and elytra, moderately light brown with slightly lighter tarsi, covered with light brown setae. BL 1.15 mm. Head broadest across moderately large and moderately strongly convex, coarsely faceted eyes, HL 0.13 mm, HW 0.29 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures fine, inconspicuous; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.60 mm, antennomere 1 about as long as broad, 2 strongly elongate, 3 about as long as broad, 4 – 6 each indistinctly elongate, 7 strongly elongate, 8 transverse, 9 and 10 each about as long as broad, 11 as long as 9 and 10 combined, about 1.6 × as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near hind corners, broadest slightly behind middle and weakly narrowing posterad; PL 0.35 mm, PW 0.56 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and weakly so posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located slightly closer to posterior than to lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, suberect. Elytra indistinctly wider than pronotum, together oval, broadest near anterior third; EL 0.68 mm, EW 0.58 mm, EI 1.17; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.3 × EL and distinctly divergent posterad; basal elytral fovea on each elytron slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded; subapical elytral region modified (Fig. 186), with cordiform flattening covered with flattened hyaline setae. Punctures on elytra similar to those on pronotum; setae short, moderately dense and suberect. Hind wings functional. Metaventrite with lateral impressions about as long as half of its length. Legs long and slender, unmodified. Aedeagus (Figs 187 – 188) of modified simonis form, AeL 0.28 mm; in ventral view with capsular region of median lobe with elongate subtriangular distal portion rounded at apex and with slender recurved distal projection with hook-like apex; parameres of uneven lengths, each with one long apical and one similarly long subapical seta. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE80913FF69F9F96C63DC79.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE80913FF69F9F96C63DC79.taxon	etymology	Etymology. The adjective arcuatus refers to the arcuate patch of modified setae on male elytra.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFE80913FF69F9F96C63DC79.taxon	discussion	Remarks. Cephennodes arcuatus can be easily identified by the unique male elytral modification; also the aedeagus of this species is unusually shaped and not possible to be confused with genitalia of any other Cephennodes.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEE0911FF69FC7A69F2DA9A.taxon	description	(Figs 189 – 193)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEE0911FF69FC7A69F2DA9A.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan [CH 07 - 17], / Baoshan Pref., mountain range 25 km S / Tengchong, 1900 m, 24 º 48 ’ 28 ’’ N, / 98 º 32 ’ 03 ’’ E, dev. primary decid. forest, / litter, fungi sifted, 2. VI. 2007, M. Schülke ” [white, printed]; “ CEPHENNODES (s. str.) / deviator m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEE0911FF69FC7A69F2DA9A.taxon	diagnosis	Diagnosis. Body large, BL around 1.6 mm; antennae gradually thickened and extremely long, also legs conspicuously long; punctures on head, pronotum and elytra fine and inconspicuous; subhumeral lines long, as long as 0.5 × EL. Male: head modified, with elongate median oval impression behind supraantennal tubercles; aedeagus (Figs 192 – 193) in ventral view with capsular region of median lobe with complex shape of strongly asymmetrical distal portion, which bears elongate triangular and pointed projection directed to the right, and with elongate and twisted distal projection, parameres of unequal length and width, each with longitudinal row of numerous setae increasing in length toward apices; sternite VI (Fig. 190) posteriorly emarginate; tergite VI (Fig. 191) elongate.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEE0911FF69FC7A69F2DA9A.taxon	description	Description. Body of male (Fig. 189) strongly convex, oval, with barely discernible constriction between pronotum and elytra, moderately light brown with slightly lighter tarsi, covered with light brown setae. BL 1.58 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.20 mm, HW 0.40 mm; frontovertexal region distinctly impressed at middle behind supraantennal tubercles, impression shallow, with flattened bottom, broadest anteriorly and narrowing posterad; supraantennal tubercles weakly elevated. Punctures fine, inconspicuous; setae short, sparse and suberect. Antennae conspicuously long and slender, gradually thickened, AnL 1.03 mm, antennomere 1 about as long as broad, 2 – 7 each weakly elongate, 8 – 10 each strongly elongate, 11 clearly shorter than 9 and 10 combined, about twice as long as broad. Pronotum almost subrectangular, moderately strongly convex at middle and flattened near hind corners, broadest at base; PL 0.50 mm, PW 0.70 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and weakly sinuate posteriorly, with shallow constriction in posterior third, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located slightly closer to posterior than to lateral pronotal margin. Punctures on pronotal disc fine and inconspicuous; setae short and moderately dense, suberect. Elytra distinctly wider than pronotum, together oval, broadest between middle and anterior third; EL 0.88 mm, EW 0.78 mm, EI 1.13; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.5 × EL and weakly divergent posterad; basal elytral fovea on each elytron slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures and setae on elytra similar to those on pronotum. Hind wings not studied. Metaventrite with deep lateral impressions about as long as 3 / 4 of its length. Legs conspicuously long and slender, unmodified. Abdomen with sternite VIII (Fig. 190) posteriorly emarginate, and tergite VIII (Fig. 191) unusually elongate. Aedeagus (Figs 192 – 193) of strongly modified simonis form, AeL 0.38 mm; in ventral view with capsular region of median lobe with complex shape of strongly asymmetrical distal portion, which bears elongate triangular and pointed projection directed to the right, and with elongate and twisted distal projection; in lateral view median lobe with narrow elongate projection directed dorsad and situated in subapical region; basal foramen hidden in strongly twisted, asymmetrical and modified basal region; parameres of unequal length and width, each with longitudinal row of numerous setae increasing in length toward apices. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEE0911FF69FC7A69F2DA9A.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEE0911FF69FC7A69F2DA9A.taxon	etymology	Etymology. The Latin adjective deviator means “ one who leaves the way ” (derived from devio, to go aside, to deviate) and alludes to the odd-shaped aedeagus.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEE0911FF69FC7A69F2DA9A.taxon	discussion	Remarks. Cephennodes deviator externally somewhat resembles members of the genus Hlavaciellus (Jałoszyński 2006, 2011 b, 2021 b) in extremely elongate legs and weakly, gradually thickened and conspicuously long antennae. The general body shape, modification of the head and antennomere 11 distinctly shorter than 9 and 10 combined are sufficient to unambiguously identify this species. The aedeagus of C. deviator is unique, odd-shaped, not possible to confuse with genital organs of any other Cephennodes. Moreover, this is the only known Cephennodes species with the abdominal sternite VIII emarginate posteriorly, and with a distinctly elongate tergite VIII. Subgenus Fusionodes Jałoszyński	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEE0911FF69FC7A69F2DA9A.taxon	description	Some Japanese species described by Hoshina can pose identification problems; this group includes C. koko Hoshina, 2021 (Shikoku), C. sakura Hoshina, 2021 (Kushyu, Shikoku), C. otome Hoshina, 2016 (Honshu), and C. nemu Hoshina, 2021 (Shikoku). Judging from illustrated genitalia, all these species were described based on aedeagi with endophallic structures extruded to various extent, and some of them, or even all, can be identical with C. loebli Kurbatov, 1995 described based on the type material collected in Shikoku. Fortunately, the aedeagus of this group of species (or one species) is not even remotely similar to that of any newly described or previously known Fusionodes from any other country. Species of this subgenus do not have any male sexual dimorphic characters, and externally are rather unremarkable. The only reliable character to identify them is the aedeagus. Because of a usually complex apical region, it is advisable to illustrate the aedeagus not only in ventral and lateral views, but also in dorsal view, otherwise the shape of some important structures (projections and lobes) cannot be assessed.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFEC0911FF69F9DB6D89D803.taxon	diagnosis	Diagnosis. Apical projections of aedeagus asymmetrical in ventral view, with apical lobe (= apical lamina, i. e. the very apex of aedeagus) projecting distad; parameral apices exposed in ventral view and setose. All species except one (C. ascipenis Jałoszyński, 2007 b) also with rounded subapical lateral lobes on median lobe of aedeagus. This group is here redefined to include a larger number of species, some previously treated as not assigned to a group. Seven previously described species are included, distributed in China, Russian Far East, South Korea, and Japan:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFED090EFF69FB576838DB19.taxon	description	(Figs 194 – 197)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFED090EFF69FB576838DB19.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, SE Pingbian, / 22 º 54 ’ 31 ’’ N, 103 º 41 ’ 44 ’’ E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 28. VIII. 2014, / leg. M. Schülke [CH 14 - 22 a] ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / pingbianus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFED090EFF69FB576838DB19.taxon	diagnosis	Diagnosis. Body large, BL around 1.7 mm; antennae gradually thickened; punctures on clypeus virtually absent, on frontovertexal region moderately large, sharply marked and dense, those at middle separated by spaces subequal to their diameters, punctures near eyes slightly denser, punctures on median region of pronotal disc smaller than those on frontovertexal region but also sharply marked and similarly dense, those on elytra fine and inconspicuous; subhumeral lines short, as long as 0.25 × EL. Male: aedeagus (Figs 195 – 197) in ventral view with capsular region of median lobe with broadly rounded distal margin; subapical lateral lobes conspicuously large and semicircular; apical projection trilobate, with two lateral subtriangular lobes and also subtriangular and pointed apical lobe; subapical distal projection in dorsal view subrectangular, wider than long and with subtriangular and projecting distolateral corners, in lateral view apical lobe distally angulate, and subapical dorsal projection recurved; each paramere with four setae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFED090EFF69FB576838DB19.taxon	description	Description. Body of male (Fig. 194) strongly convex, oval, with barely discernible constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.73 mm. Head broadest across large and conspicuously strongly convex, coarsely faceted eyes, HL 0.25 mm, HW 0.40 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus virtually absent, on frontovertexal region moderately large, sharply marked and dense, those at middle separated by spaces subequal to their diameters, punctures near eyes slightly denser; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.88 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 – 6 each weakly transverse, 7 – 10 each about as long as broad, 11 nearly as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, broadest near middle and weakly narrowing posterad; PL 0.53 mm, PW 0.78 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and weakly sinuate posteriorly, just in front of hind corners weakly concave, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each located slightly closer to posterior than to lateral pronotal margin. Punctures on median region of pronotal disc slightly smaller than those on head but also sharply marked and separated by spaces subequal to their diameters, punctures near anterior and posterior corners denser and somewhat coarse; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.95 mm, EW 0.83 mm, EI 1.15; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.25 × EL and weakly divergent posterad; basal elytral fovea on each elytron slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings lacking. Metaventrite with deep lateral impressions as long as 4 / 5 of its length. Legs long and slender, unmodified. Aedeagus (Figs 195 – 197) AeL 0.58 mm; in ventral view capsular region of median lobe with broadly rounded distal margin; subapical lateral lobes conspicuously large and semicircular; apical projection trilobate, with two lateral subtriangular sharp-angled lobes and also subtriangular and pointed apical lobe; in lateral view apical lobe distally angulate; subapical dorsal projection in ventral view subrectangular, wider than long and with subtriangular and projecting distolateral corners, in lateral view subapical dorsal projection sinuate; each parameral apex strongly curved mesad and with four setae. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFED090EFF69FB576838DB19.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFED090EFF69FB576838DB19.taxon	etymology	Etymology. The adjective pingbianus refers to the type locality in Pingbian Miao Autonomous County.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFED090EFF69FB576838DB19.taxon	discussion	Remarks. Cephennodes pingbianus can be identified by its unique aedeagus, which in ventral view has a trilobate apical structure, composed of a pair of subtriangular sharp-angled lateral lobes and also a triangular apical lobe.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF3090DFF69FB5A681BDABF.taxon	description	(Figs 198 – 201)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF3090DFF69FB5A681BDABF.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA (Yunnan) Dali Bai Aut. / Pref., Jizu Shan, summit plateau, / 37 km NE Dali, 3150 m / 25 º 58 ’ 30 ’’ N / 100 º 21 ’ 36 ’’ E, / (mixed forest, sifted from litter, / moss) / 5. IX. 2009 D. W. Wrase [28] ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / jizushanus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (3 exx.): 1 ♂, “ CHINA: Yunnan, Dali Bai Aut. Pref., / Jizu Shan, summit plateau, 37 km / NE Dali, 25 º 58 ’ 30 ’’ N, 100 º 21 ’ 36 ’’ E, / 3150 m, mixed forest, sifted from / litter, moss & mushrooms, 5. IX. 2009. / leg. M. Schülke [CH 09 - 28] ”; 2 ♂♂, “ CHINA: Yunnan, Dali Bai Aut. Pref., / Jizu Shan, path to cable car, 37 km NE / Dali, 25 º 58 ’ N, 100 º 23 ’ E, / 2450 m, mixed forest, sifted from litter, / moss & pine apples, 5. IX. 2009. / leg. M. Schülke [CH 09 - 29] ”. Paratypes with yellow PARATYPUS labels (cPJ, MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF3090DFF69FB5A681BDABF.taxon	diagnosis	Diagnosis. Body large, BL around 1.7 – 1.9 mm; antennae gradually thickened; punctures on clypeus and frontovertexal region forming two lateral longitudinal stripes composed of moderately large, sharply marked punctures separated by spaces slightly shorter than their diameters, median area with distinctly sparser punctures, punctures on median region of pronotal disc smaller than those on frontovertexal region but also sharply marked and separated by spaces subequal to their diameters, those on elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 199 – 201) in ventral view with capsular region of median lobe with subtriangular and rounded distal margin; subapical lateral lobes elongate, subparallel and with subtriangular distal regions; apical projection trilobate, with two lateral subtriangular lobes and flame-like pointed apical lobe; subapical dorsal projection in ventral view divided into two lateral broadly separated elongate processes; in lateral view apical lobe with subtriangular and pointed distal projection, and subapical dorsal projection strongly curved; each paramere with only one thick seta.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF3090DFF69FB5A681BDABF.taxon	description	Description. Body of male (Fig. 198) strongly convex, oval, with barely discernible constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.73 – 1.85 mm. Head broadest across large and conspicuously strongly convex, coarsely faceted eyes, HL 0.25 mm, HW 0.40 – 0.41 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region forming two lateral longitudinal stripes composed of moderately large, sharply marked punctures separated by spaces slightly shorter than their diameters, median area with distinctly sparser punctures; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.85 – 0.90 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 weakly transverse, 4 – 7 each about as long as broad, 8 weakly transverse, 9 and 10 each about as long as broad, 11 nearly as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, equally broad from base to near middle; PL 0.50 – 0.55 mm, PW 0.78 – 0.80 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc slightly smaller than those on head but also sharply marked and separated by spaces subequal to their diameters, punctures near anterior and posterior corners much denser and coarse; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.95 – 1.05 mm, EW 0.80 – 0.85 mm, EI 1.15 – 1.24; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.30 × EL and weakly divergent posterad; basal elytral fovea on each elytron slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite with lateral impressions as long as half of its length. Legs long and slender, unmodified. Aedeagus (Figs 199 – 201) AeL 0.55 mm; in ventral view capsular region of median lobe with subtriangular and rounded distal margin; subapical lateral lobes elongate, subparallel and with subtriangular distal regions; apical projection trilobate, with two lateral subtriangular obtuse-angled lobes and flame-like pointed apical lobe; subapical dorsal projection in ventral view divided into two lateral broadly separated elongate processes, in lateral view each strongly curved; in lateral view apical lobe with subtriangular and pointed distal projection; each parameral apex weakly bent mesad, with only one thick seta. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF3090DFF69FB5A681BDABF.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF3090DFF69FB5A681BDABF.taxon	etymology	Etymology. The adjective jizushanus refers to the type locality, Mount Jizu Shan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF3090DFF69FB5A681BDABF.taxon	discussion	Remarks. Cephennodes jizushanus can be identified by its unique aedeagus, which in ventral view has a trilobate apical structure, composed of a pair of subtriangular obtuse-angled lateral lobes and a flame-like apical lobe, and with an additional pair of elongate dorsal subapical lobes (which do not occur in the similar C. pingbianus).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF0090DFF69F9A06822D85E.taxon	diagnosis	Diagnosis. Aedeagus symmetrical, in ventral view with subtriangular, subtrapezoidal or shield-like apical projection narrowing distad and composed of pair of elongate apical lateral projections; parameral apices exposed in ventral view and setose. This newly established group includes four previously described species distributed in China and Vietnam:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF1090AFF69FC1A6847DCC1.taxon	description	(Figs 202 – 205)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF1090AFF69FC1A6847DCC1.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, SE Pingbian, / 22 º 54 ’ 31 ’’ N, 103 º 41 ’ 44 ’’ E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 28. VIII. 2014, / leg. M. Schülke [CH 14 - 22 a] ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / microsoma m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratype: 1 ♂, same data as for holotype, with yellow PARATYPUS labels (cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF1090AFF69FC1A6847DCC1.taxon	diagnosis	Diagnosis. Body moderately large, BL around 1.4 – 1.5 mm; antennae gradually thickened; punctures on clypeus and frontovertexal region fine but distinct, sharply marked and separated by spaces subequal to their diameters, punctures on median region of pronotal disc similar to those on head, those on elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 203 – 205) in ventral view with capsular region of median lobe with bifurcate distal margin; subapical lateral lobes subtriangular and about as long as broad; apical lateral projections touching at middle only with their apices; in dorsal view proximal subapical dorsal projection with broadly and shallowly emarginate distal margin; in lateral view apical lateral projection vertical and proximal subapical dorsal projection bent apically at right angle; each paramere with two setae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF1090AFF69FC1A6847DCC1.taxon	description	Description. Body of male (Fig. 202) strongly convex, oval, with shallow but distinct constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.43 – 1.45 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.20 mm, HW 0.35 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region fine but distinct, sharply marked and separated by spaces subequal to their diameters; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.68 – 0.75 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 – 10 each transverse, 11 indistinctly longer than 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, broadest near middle and weakly narrowing posterad; PL 0.43 – 0.45 mm, PW 0.60 – 0.63 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior half and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc fine but distinct, sharply marked and separated by spaces subequal to their diameters, punctures near anterior corners much denser and somewhat coarse; setae short and moderately dense, suberect. Elytra distinctly wider than pronotum, together oval, broadest near anterior third; EL 0.80 mm, EW 0.70 – 0.68 mm, EI 1.14 – 1.19; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.30 × EL and strongly divergent posterad; basal elytral fovea on each elytron slightly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 203 – 205) AeL 0.48 mm; in ventral view with capsular region of median lobe with bifurcate distal margin developed as pair of subtriangular lobes separated by U-shaped emargination; subapical lateral lobes subtriangular and each about as long as broad; apical lateral projections with convex lateral margins, except for short subapical concave area, touching at middle only with apices; in dorsal view proximal subapical dorsal projection subtrapezoidal and with broadly and shallowly emarginate distal margin; in lateral view apical lateral projection vertical and distally angulate, proximal subapical dorsal projection bent apically at right angle; parameral apices in ventral view straight and parallel to long axis of aedeagus, each with two setae. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF1090AFF69FC1A6847DCC1.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF1090AFF69FC1A6847DCC1.taxon	etymology	Etymology. The noun microsoma refers to the relatively small body as for a member of a typically large-bodied subgenus.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF1090AFF69FC1A6847DCC1.taxon	discussion	Remarks. Cephennodes microsoma is the only member of the C. hippopotamus species group that has the aedeagus in lateral view with a long proximal subapical dorsal projection distally bent at a right angle.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF70909FF69FC22697CDBE1.taxon	description	(Figs 206 – 209)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF70909FF69FC22697CDBE1.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, mountain W / Gejiu, 23 º 24 ’ 13 ’’ N, 103 º 07 ’ 28 ’’ E, / 1990 m, mixed forest, litter and / various debris sifted, 25. VIII. 2014, / leg. M. Schülke [CH 14 - 20 b] ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / spatulatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (2 exx.): 2 ♀♀, same data as for holotype, each with yellow PARATYPUS labels (cPJ, MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF70909FF69FC22697CDBE1.taxon	diagnosis	Diagnosis. Body moderately large, BL around 1.5 – 1.6 mm; antennae gradually thickened; punctures on clypeus and frontovertexal region fine but distinct, sharply marked and these on lateral regions separated by spaces slightly shorter than their diameters, those on median area distinctly sparser, punctures on median region of pronotal disc small but distinct and separated by spaces subequal to their diameters, those on elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 207 – 209) in ventral view with capsular region of median lobe with weakly concave distal margin; subapical lateral lobes lacking; apical lateral projections not touching at middle; in dorsal view proximal subapical dorsal projection oval, slightly broadening distad with lateral and distal margins rounded, and distal subapical dorsal projection subrectangular with its distal margin straight; in lateral view apical lateral projection vertical and distally rounded; in lateral view distal subapical dorsal projection conspicuously large and broadening distad, and proximal subapical dorsal projection subtriangular; each paramere with one seta.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF70909FF69FC22697CDBE1.taxon	description	Description. Body of male (Fig. 206) strongly convex, oval, with indistinct constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.55 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.20 mm, HW 0.33 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus and frontovertexal region fine but distinct, sharply marked and these on lateral regions separated by spaces slightly shorter than their diameters, those on median area distinctly sparser; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.73 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 about as long as broad, 4 – 10 each transverse, 11 about as long as 9 and 10 combined, about 1.7 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, equally broad between base and slightly anterior to middle; PL 0.48 mm, PW 0.65 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc small but distinct and separated by spaces subequal to their diameters, punctures near anterior and posterior corners much denser and coarse; setae short and moderately dense, suberect. Elytra distinctly wider than pronotum, together oval, broadest near anterior third; EL 0.88 mm, EW 0.73 mm, EI 1.21; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.30 × EL and weakly divergent posterad; basal elytral fovea on each elytron nearly equidistant to lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings not studied. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 207 – 209) AeL 0.40 mm; in ventral view with capsular region of median lobe with weakly concave distal margin; subapical lateral lobes lacking; apical lateral projections not touching at middle, their outer lateral margins strongly sinuate; in dorsal view proximal subapical dorsal projection oval, slightly broadening distad with lateral and distal margins rounded, and distal subapical dorsal projection subrectangular with its distal margin straight; in lateral view apical lateral projection vertical and distally rounded; in lateral view distal subapical dorsal projection conspicuously large and broadening distad, and proximal subapical dorsal projection subtriangular; each paramere with one seta. Female. Externally indistinguishable from male, wings not studied; BL 1.51 – 1.60 mm; HL 0.21 – 0.23 mm, HW 0.33 – 0.35 mm, AnL 0.68 – 0.75 mm; PL 0.48 – 0.50 mm, PW 0.70 mm; EL 0.83 – 0.88 mm, EW 0.73 mm, EI 1.14 – 1.21.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF70909FF69FC22697CDBE1.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF70909FF69FC22697CDBE1.taxon	etymology	Etymology. The adjective spatulatus refers to the spatulate shape of the dorsal subapical projection of the aedeagus in lateral view.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF70909FF69FC22697CDBE1.taxon	discussion	Remarks. Cephennodes spatulatus is the only member of the C. hippopotamus species group that has the aedeagus in lateral view with two conspicuously massive dorsal subapical projections, of which distal one is strongly broadening towards apex, and proximal one is much shorter and distally tapering.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF40907FF69FB026847DF94.taxon	description	(Figs 210 – 213)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF40907FF69FB026847DF94.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA (Yunnan) Pu’er Pref., / Ailao Shan, 37 km NW Jingdong / 24 º 45 ’ 12 ’’ N / 100 º 41 ’ 24.5 ’’ N {sic!} / 2300 m (devastated forest rem- / nant, litter / moss / grass roots sifted / 13. IX. 2009 D. W. Wrase [48] ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / brachycornis m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF40907FF69FB026847DF94.taxon	diagnosis	Diagnosis. Body large, BL around 1.8 mm; antennae gradually thickened; punctures on clypeus virtually absent, on frontovertexal region fine but distinct, sharply marked and these on lateral regions separated by spaces slightly shorter than their diameters, those on median area distinctly sparser, punctures on median region of pronotal disc slightly smaller than on head but distinct and separated by spaces subequal to their diameters, those on elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 211 – 213) in ventral view with capsular region of median lobe with straight distal margin; subapical lateral lobes lacking; apical lateral projections not touching at middle; proximal and distal subapical dorsal projections rudimentary, each visible in lateral view as minute subtriangular convexity; in lateral view apical lateral projection strongly bent dorsad and subtriangular; each paramere with two setae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF40907FF69FB026847DF94.taxon	description	Description. Body of male (Fig. 210) strongly convex, oval, with shallow but distinct constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.78 mm. Head broadest across large and strongly convex, coarsely faceted eyes, HL 0.25 mm, HW 0.40 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus virtually lacking, those on frontovertexal region fine but distinct, sharply marked and on lateral regions separated by spaces slightly shorter than puncture diameters, those on median area distinctly sparser; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.88 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 – 6 each indistinctly transverse, 7 about as long as broad, 8 – 10 each slightly transverse, 11 about as long as 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, broadest near middle but indistinctly narrowing posterad; PL 0.55 mm, PW 0.78 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc small but distinct and separated by spaces subequal to their diameters, punctures near anterior and posterior corners much denser and coarse; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.98 mm, EW 0.83 mm, EI 1.18; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.30 × EL and weakly divergent posterad; basal elytral fovea on each elytron nearly equidistant to lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings vestigial. Metaventrite with lateral impressions as long as 3 / 4 of its length. Legs long and slender, unmodified. Aedeagus (Figs 211 – 213) AeL 0.50 mm; in ventral view capsular region of median lobe nearly parallel-sided, with broadly subtrapezoidal distal portion and distal margin straight at middle; subapical lateral lobes lacking; apical projections in ventral view subtriangular and much narrower than capsular region of median lobe; apical lateral projections not touching at middle; proximal and distal subapical dorsal projections vestigial, each visible in lateral view as minute subtriangular convexity; in lateral view apical lateral projection strongly bent dorsad and subtriangular; each paramere with two setae. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF40907FF69FB026847DF94.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF40907FF69FB026847DF94.taxon	etymology	Etymology. The adjective brachycornis refers to the relatively short antennae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF40907FF69FB026847DF94.taxon	discussion	Remarks. Cephennodes brachycornis is the only member of the C. hippopotamus species group that has the aedeagus in ventral view with the capsular region parallel-sided instead of distinctly narrowing distad.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFA0907FF69FED668CADB3C.taxon	diagnosis	Diagnosis. Aedeagus in lateral view with subcylindrical apical lateral projection and one tongue-like subapical dorsal projection; in ventral view apical structures composed of apical lateral projections and subapical dorsal projection symmetrical and much narrower than basal capsule; parameral apices not discernible and asetose. This group includes two previously described species distributed in Yunnan, C. brachylinguis Jałoszyński, 2007 a and C. longilinguis Jałoszyński, 2007 a. Two more species are described below.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFA0905FF69FB3F6ECDDBE1.taxon	description	(Figs 214 – 217)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFA0905FF69FB3F6ECDDBE1.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan [CH 07 - 19], Dehong Dai / Aut. Pref., mountain range 31 km E Luxi, / 2280 m, 24 º 29 ’ 31 ’’ N, 98 º 52 ’ 58 ’’ E, / second. pine forest with old decid. trees, / litter sifted, 3. VI. 2007, M. Schülke ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / linguatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (4 exx.): 2 ♂♂, 2 ♀♀, same data as for holotype, each with yellow PARATYPUS label (MNB, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFA0905FF69FB3F6ECDDBE1.taxon	diagnosis	Diagnosis. Body moderately large, BL around 1.5 mm; antennae gradually thickened; punctures on median region of clypeus virtually absent, few distinct punctures present on lateral regions of clypeus, punctures on median area of frontovertexal region small but distinct, sharply marked and separated by spaces subequal to their diameters, those on lateral areas of frontovertexal region distinctly denser, separated by spaces slightly shorter than their diameters, punctures on median region of pronotal disc slightly smaller than on head but distinct and separated by spaces subequal to their diameters, those on elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 215 – 217) in ventral view with capsular region of median lobe with shallowly bisinuate distal margin, and its distal margin in lateral view rounded, lacking emargination or indentation; subapical dorsal projection in dorsal view gradually narrowing distad, in lateral view with weakly rounded dorsal margin.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFA0905FF69FB3F6ECDDBE1.taxon	description	Description. Body of male (Fig. 214) strongly convex, oval, with shallow and indistinct constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.43 – 1.58 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.20 – 0.23 mm, HW 0.35 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on median region of clypeus virtually absent, few distinct punctures present on lateral regions of clypeus, punctures on median area of frontovertexal region small but distinct, sharply marked and separated by spaces subequal to their diameters, those on lateral areas of frontovertexal region distinctly denser, separated by spaces slightly shorter than their diameters; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.68 – 0.75 mm, antennomere 1 and 2 each slightly elongate, 3 – 7 each about as long as broad, 8 – 10 each slightly transverse, 11 about as long as 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, broadest at base; PL 0.45 – 0.48 mm, PW 0.65 – 0.70 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc slightly smaller than on head but distinct and separated by spaces subequal to their diameters, punctures near anterior and posterior corners much denser and coarse; setae short and moderately dense, suberect. Elytra slightly wider than pronotum, together oval, broadest near anterior third; EL 0.75 – 0.88 mm, EW 0.65 – 0.75 mm, EI 1.15 – 1.17; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.30 × EL and weakly divergent posterad; basal elytral fovea on each elytron nearly equidistant to lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Of three studied males, two have functional wings and one is wingless. Metaventrite with lateral impressions as long as half of its length. Legs long and slender, unmodified. Aedeagus (Figs 215 – 217) AeL 0.43 mm; in ventral view with capsular region of median lobe with shallowly bisinuate distal margin, and its distal margin in lateral view rounded, lacking emargination or indentation; subapical dorsal projection in dorsal view gradually narrowing distad, in lateral view with weakly rounded dorsal margin; parameres indiscernible. Female. Externally indistinguishable from male, wings not studied. BL 1.49 – 1.50; HL 0.21 – 0.23 mm, HW 0.35 mm, AnL 0.70 – 0.73 mm; PL 0.48 – 0.50 mm, PW 0.68 – 0.70 mm; EL 0.78 – 0.80 mm, EW 0.70 mm, EI 1.11 – 1.14.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFA0905FF69FB3F6ECDDBE1.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFA0905FF69FB3F6ECDDBE1.taxon	etymology	Etymology. As in all four members of the C. longilinguis group, the name is an adjective derived from the Latin lingua, a tongue, referring to the tongue-like subapical dorsal projection of the aedeagus.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFA0905FF69FB3F6ECDDBE1.taxon	discussion	Remarks. Within the C. longilinguis group, C. linguatus can be identified by the following combination of characters: in lateral view, distal margin of the capsular region of median lobe convex and rounded, and dorsal margin of the subapical dorsal projection weakly and evenly convex.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF80904FF69FB026CCADA5D.taxon	description	(Figs 218 – 221)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF80904FF69FB026CCADA5D.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Lincang Pref., / Laobie Shan, Wei Bo Shan pass, / 24 º 08 ’ 16 ’’ N, 99 º 42 ’ 53 ’’ E, 2375 m, / creek valley, devastated secondary / decid. forest, litter & moss sifted, / 8. IX. 2009, leg. M. Schülke [CH 09 - 35] ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / linguatoides m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF80904FF69FB026CCADA5D.taxon	diagnosis	Diagnosis. Body small, BL around 1.3 mm; antennae gradually thickened; punctures on median region of clypeus virtually absent, few distinct punctures present on lateral regions of clypeus, punctures on median area of frontovertexal region small but distinct, sharply marked and separated by spaces slightly wider than their diameters, those on lateral areas of frontovertexal region distinctly denser, separated by spaces subequal to their diameters, punctures on median region of pronotal disc and on elytra barely discernible and separated by spaces subequal to their diameters; subhumeral lines short, as long as 0.25 × EL. Male: aedeagus (Figs 219 – 221) in ventral view with capsular region of median lobe with subtrapezoidal distal margin, its median portion straight, distal margin in lateral view with deep U-shaped emargination; subapical dorsal projection in dorsal view somewhat angulate in submedian region, in lateral view with weakly concave dorsal margin.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF80904FF69FB026CCADA5D.taxon	description	Description. Body of male (Fig. 218) strongly convex, oval, with shallow and indistinct constriction between pronotum and elytra, moderately dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.35 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.18 mm, HW 0.31 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on median region of clypeus virtually absent, few distinct punctures present on lateral regions of clypeus, punctures on median area of frontovertexal region small but distinct, sharply marked and separated by spaces slightly wider than their diameters, those on lateral areas of frontovertexal region distinctly denser, separated by spaces subequal to their diameters; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 0.65 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 – 7 each about as long as broad, 8 – 10 each slightly transverse, 11 about as long as 9 and 10 combined, about 1.8 × as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, equally broad between base and near middle; PL 0.40 mm, PW 0.60 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc barely discernible and separated by spaces subequal to their diameters, punctures near anterior corners much denser and slightly coarse; setae short and moderately dense, suberect. Elytra indistinctly wider than pronotum, together oval, broadest near anterior third; EL 0.78 mm, EW 0.63 mm, EI 1.24; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.25 × EL and weakly divergent posterad; basal elytral fovea on each elytron nearly equidistant to lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra similar to those on median region of pronotum but slightly larger and less distinct; setae similar to those on pronotum. Hind wings functional. Metaventrite lacking lateral impressions. Legs long and slender, unmodified. Aedeagus (Figs 219 – 221) AeL 0.40 mm; in ventral view capsular region of median lobe with subtrapezoidal distal margin, its median portion straight, distal margin in lateral view with deep U-shaped emargination; subapical dorsal projection in dorsal view parallel-sided proximally, somewhat angulate in submedian region, and gradually narrowing distad in distal half, in lateral view with weakly concave dorsal margin; parameres indiscernible. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF80904FF69FB026CCADA5D.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF80904FF69FB026CCADA5D.taxon	etymology	Etymology. The adjective linguatoides is derived from the species name C. linguatus.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF80904FF69FB026CCADA5D.taxon	discussion	Remarks. Within the C. longilinguis group, C. linguatoides can be identified by the following combination of characters: in lateral view, the subapical dorsal projection much longer than apical lateral projection, and distal margin of capsular region of median lobe with deep U-shaped emargination; in ventral view, distal region of aedeagus almost as long as wide.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFF90904FF69FA9F6822D95A.taxon	diagnosis	Diagnosis. Aedeagus symmetrical, in ventral view with shield-like apical structures narrowing distad and not divided into long lateral projections, narrower than basal capsule; and parameral apices setose. This newly established group includes four previously described species distributed in China and Vietnam:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFE0901FF69FD2468CBDD59.taxon	description	(Figs 222 – 225)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFE0901FF69FD2468CBDD59.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, SE Pingbian, / 22 º 54 ’ 31 ’’ N, 103 º 41 ’ 44 ’’ E, 2100 m / primary subtropical broad-leaved / forest, litter sifted / 27. VIII. 2014, / leg. M. Schülke [CH 14 - 22] ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / brachyclavatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFE0901FF69FD2468CBDD59.taxon	diagnosis	Diagnosis. Body small, BL around 1.2 mm; antennae conspicuously short and with strongly thickened tetramerous clubs; punctures on clypeus virtually absent, a few small but sharply marked punctures present on median area of frontovertexal region and unevenly distributed, lateral areas of frontovertexal region with similar but much denser punctures separated by spaces shorter than their diameters, punctures on median region of pronotal disc indistinctly smaller than those on head but also sharply marked, separated by spaces subequal to their diameters, punctures on elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 223 – 225) in ventral view with capsular region of median lobe with broadly and shallowly emarginate distal margin, apex of apical projection shallowly and narrowly emarginate, and parameres not projecting distad beyond capsular region.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFE0901FF69FD2468CBDD59.taxon	description	Description. Body of male (Fig. 222) moderately strongly convex, oval, with barely discernible constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.25 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.15 mm, HW 0.28 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on clypeus virtually absent, a few small but sharply marked punctures present on median area of frontovertexal region and unevenly distributed, lateral areas of frontovertexal region with similar but much denser punctures separated by spaces shorter than their diameters; setae short, sparse and suberect. Antennae conspicuously short and strongly thickened distad, with tetramerous clubs, AnL 0.58 mm, antennomere 1 about as long as broad, 2 distinctly elongate, 3 – 10 each transverse (8 – 10 strongly so), 11 about as long as 9 and 10 combined, about 1.3 × as long as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near anterior and hind corners, equally broad between base and near middle; PL 0.40 mm, PW 0.53 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc indistinctly smaller than those on head but also sharply marked, separated by spaces subequal to their diameters, punctures near anterior and posterior corners much denser and slightly coarse; setae short and moderately dense, suberect. Elytra about as wide as pronotum, together oval, broadest near anterior third; EL 0.70 mm, EW 0.55 mm, EI 1.27; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.3 × EL and weakly divergent posterad; basal elytral fovea on each elytron nearly equidistant to lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings lacking. Metaventrite with lateral impressions as long as 1 / 3 of its length. Legs moderately long and slender, unmodified. Aedeagus (Figs 223 – 225) AeL 0.43 mm; in ventral view with capsular region of median lobe with broadly and shallowly emarginate distal margin; subapical lateral lobes not projecting beyond lateral margins of capsular region; apex of apical projection shallowly and narrowly emarginate, its lateral margins weakly sinuate and narrowing from base of projection to apex; in lateral view apical projection short and broadening distad, subapical dorsal projection about as long as apical projection; parameral apices in ventral view not projecting distad beyond capsular region, slender and each with one long apical seta. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFE0901FF69FD2468CBDD59.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFE0901FF69FD2468CBDD59.taxon	etymology	Etymology. The adjective brachyclavatus refers to the unusually short antennae in relation to the body length.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFE0901FF69FD2468CBDD59.taxon	discussion	Remarks. Cephennodes brachyclavatus and C. microphthalmus are the only members of this group with adults reaching less than 1.4 mm in length. They differ clearly in external morphology. Cephennodes brachyclavatus has conspicuously short and strongly thickened antennae and large eyes, whereas C. microphthalmus has long and slender antennae and strongly reduced eyes; aedeagi of these species also strongly differ, especially in lateral view.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFC0900FF69FD9A69EFDCAD.taxon	description	(Figs 226 – 229)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFC0900FF69FD9A69EFDCAD.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Dali Bai Aut. Pref., / mount. range E Weishan, 12 km NE / Weishan, 25 º 17 ’ 02 - 15 ’’ N, 100 º 22 ’ / 23 - 30 ’’ E, 2630 - 2660 m, scrub with / pines and bamboo, litter sifted, 15. IX. / 2009, leg. M. Schülke [CH 09 - 54] ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / longicornis m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (2 exx.): 2 ♂♂, same data as for holotype (MNB, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFC0900FF69FD9A69EFDCAD.taxon	diagnosis	Diagnosis. Body very large, BL around 1.9 – 2.0 mm; antennae long and slender, gradually thickened; punctures on median region of clypeus virtually absent, punctures on lateral regions of clypeus and on entire frontovertexal region moderately large, sharply marked and separated by spaces shorter than their diameters, punctures on median region of pronotal disc much smaller than those on head but also sharply marked, separated by spaces subequal to their diameters, punctures on elytra fine and inconspicuous; subhumeral lines short, as long as 0.3 × EL. Male: aedeagus (Figs 227 – 229) in ventral view with capsular region of median lobe with strongly projecting distad subtriangular distal margin, apex of apical projection subtriangular and obtuse-angled, and parameres clearly projecting distad beyond capsular region.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFC0900FF69FD9A69EFDCAD.taxon	description	Description. Body of male (Fig. 226) strongly convex, oval, with relatively deep constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.93 – 2.00 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.25 – 0.28 mm, HW 0.43 – 0.44 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on median region of clypeus virtually absent, punctures on lateral regions of clypeus and on entire frontovertexal region moderately large, sharply marked and separated by spaces shorter than their diameters; setae short, sparse and suberect. Antennae long and slender, gradually thickened, AnL 1.03 – 1.05 mm, antennomere 1 slightly elongate, 2 weakly elongate, 3 – 8 each about as long as broad, 9 and 10 each slightly elongate, 11 slightly shorter than 9 and 10 combined, about twice as long as broad. Pronotum semi-oval, strongly convex at middle and flattened near anterior and hind corners, broadest slightly behind middle but indistinctly narrowing posterad; PL 0.60 – 0.63 mm, PW 0.88 – 0.90 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each nearly equidistant to posterior and lateral pronotal margin. Punctures on median region of pronotal disc much smaller than those on head but also sharply marked, separated by spaces subequal to their diameters, punctures near anterior and posterior corners much denser and slightly coarse; setae short and moderately dense, suberect. Elytra indistinctly wider than pronotum, together oval, broadest near anterior third; EL 1.03 – 1.10 mm, EW 0.93 mm, EI 1.11 – 1.19; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.3 × EL and weakly divergent posterad; basal elytral fovea on each elytron clearly closer to lateral margin of mesoscutellum than to subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Hind wings functional. Metaventrite with lateral impressions as long as half of its length. Legs long and slender, unmodified. Aedeagus (Figs 227 – 229) AeL 0.50 mm; in ventral view capsular region of median lobe with subtriangular and sharp-angled distal margin forming plate strongly projecting distad; subapical lateral lobes strongly projecting beyond lateral margins of capsular region; apex of apical projection broadly subtriangular, its lateral margins subtriangular, from base diverging distad and then equally strongly converging towards apex; in lateral view apical projection conspicuously massif, subtriangular and narrowing distad, subapical dorsal projection shorter than apical projection; parameral apices in ventral view projecting distad beyond capsular region, conspicuously long and thick, each with three short apical setae. Female. Unknown.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFC0900FF69FD9A69EFDCAD.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFC0900FF69FD9A69EFDCAD.taxon	etymology	Etymology. The adjective longicornis refers to the long antennae.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFC0900FF69FD9A69EFDCAD.taxon	discussion	Remarks. Cephennodes longicornis is a conspicuously large-bodied species (BL ~ 2 mm), with long and slender antennae, and can be easily identified by its unique aedeagus. Within the C. microphthalmus group, this is the only species with the subapical lateral lobes in ventral view strongly projecting laterally beyond lateral margins of the capsular region of median lobe, a strongly elongate, subtriangular apex of the capsular region, and the apical projection rhomboidal and slightly wider than long. Moreover, the parameral apices are conspicuously long and broad, and projecting distad beyond the apex of the capsular region of the aedeagus. Species of Cephennodes (Fusionodes) not placed in groups After a reorganization of Cephennodes, only four previously described species of Fusionodes will remain not assigned (Jałoszyński, in prep.). Among them there are three species known to occur in China:	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFD093FFF69F99D6F04D8D5.taxon	description	(Figs 230 – 235)	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFD093FFF69F99D6F04D8D5.taxon	materials_examined	Type material. Holotype: CHINA (YUNNAN Prov.): ♂, two labels: “ CHINA: Yunnan, Dali Bai Aut. Pref., / Zhemo Shan, 7 km SW Xiaguan, / 25 º 32 - 33 ’ N, 100 º 10 - 11 ’ E, 2870 - 2970 / m, scrub with bamboo, oaks & / Rhododendr., litter sifted, 18. IX. / 2009, leg. M. Schülke [CH 09 - 60] ” [white, printed]; “ CEPHENNODES / (FUSIONODES) / pseudolinguatus m. / P. Jałoszyński, 2025 / HOLOTYPUS ” [red, printed] (MNB). Paratypes (3 exx.): 1 ♂, 2 ♀♀, same data as for holotype (MNB, cPJ).	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFD093FFF69F99D6F04D8D5.taxon	diagnosis	Diagnosis. Body moderately small, BL around 1.2 – 1.3 mm; antennae moderately long with strongly thickened tetramerous clubs; punctures on median region of clypeus virtually absent, punctures on lateral regions of clypeus and on lateral regions of frontovertexal region small but sharply marked and separated by spaces shorter than their diameters, on median area of frontovertexal region only a few punctures, punctures on median region of pronotal disc similarly small as those on head but also sharply marked, separated by spaces subequal to their diameters, punctures on elytra fine and inconspicuous; subhumeral lines short, as long as 0.2 × EL. Male: aedeagus (Figs 231 – 235) in ventral view capsular region of median lobe with subtrapezoidal and strongly projecting distad distal margin, apex of apical projection deeply emarginate with narrow elongate median process between lateral halves, and parameres indiscernible, asetose.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFD093FFF69F99D6F04D8D5.taxon	description	Description. Body of male (Fig. 230) moderately strongly convex, oval, with distinct constriction between pronotum and elytra, dark brown with slightly lighter tarsi, covered with light brown setae. BL 1.25 mm. Head broadest across large and moderately strongly convex, coarsely faceted eyes, HL 0.15 – 0.18 mm, HW 0.30 mm; frontovertexal region weakly convex at middle; supraantennal tubercles weakly elevated. Punctures on median region of clypeus virtually absent, punctures on lateral regions of clypeus and on lateral regions of frontovertexal region small but sharply marked and separated by spaces shorter than their diameters, on median area of frontovertexal region only a few punctures; setae short, sparse and suberect. Antennae moderately long with strongly thickened tetramerous clubs, AnL 0.55 – 0.60 mm, antennomere 1 about as long as broad, 2 weakly elongate, 3 – 5 each slightly transverse, 6 about as long as broad, 7 weakly transverse, 8 – 10 each weakly transverse, 11 about as long as 9 and 10 combined, about 1.7 × as broad. Pronotum semi-oval, moderately strongly convex at middle and flattened near anterior and hind corners, broadest near middle and weakly narrowing posterad; PL 0.38 – 0.43 mm, PW 0.43 – 0.55 mm. Anterior margin in strictly dorsal view nearly straight; lateral margins strongly rounded in anterior third and nearly straight posteriorly, finely microserrate, posterior corners nearly right-angled; posterior margin shallowly bisinuate; lateral marginal carinae simple (i. e., not doubled); lateral antebasal pits shallow but distinct, each slightly closer to lateral than to posterior pronotal margin. Punctures on median region of pronotal disc similarly small as those on head but also sharply marked, separated by spaces subequal to their diameters, punctures near anterior and posterior corners much denser and slightly coarse; setae short and moderately dense, suberect. Elytra indistinctly wider than pronotum, together oval, broadest near anterior third; EL 0.68 – 0.70 mm, EW 0.55 – 0.58 mm, EI 1.22 – 1.23; subhumeral lines developed as sharp borders between elevated humerus and lower mesal region, as long as 0.2 × EL and weakly divergent posterad; basal elytral fovea on each elytron nearly equidistant to lateral margin of mesoscutellum and subhumeral line; elytral apices separately rounded. Punctures on elytra fine and inconspicuous; setae similar to those on pronotum. Both studied males wingless. Metaventrite with lateral impressions nearly as long as half of its length. Legs long and slender, unmodified. Aedeagus (Figs 231 – 235) AeL 0.43 mm; in ventral view with capsular region of median lobe with subtrapezoidal distal margin forming plate strongly projecting distad; subapical lateral lobes lacking; apex of apical projection deeply emarginate, slender median projection present between lateral halves; in lateral view apical projection broadening distad, subapical dorsal projection about as long as apical projection and narrowing distad; parameral apices indiscernible. Female. Externally indistinguishable from male, wings not studied. BL 1.25 – 1.30 mm; HL 01.8 mm, HW 0.30 mm, AnL 0.55 – 0.60 mm; PL 0.40 mm, PW 0.55 – 0.58 mm; EL 0.68 – 0.73 mm, EW 0.55 – 0.58 mm, EI 1.23 – 1.26.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFD093FFF69F99D6F04D8D5.taxon	distribution	Distribution. S China: Yunnan.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFD093FFF69F99D6F04D8D5.taxon	etymology	Etymology. The adjective pseudolinguatus refers to the similarity of this species to C. linguatus.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
6C303602FFFD093FFF69F99D6F04D8D5.taxon	discussion	Remarks. Cephennodes pseudolinguatus has the aedeagus showing some similarities to the copulatory organs of the C. hippopotamus, C. longilinguis and C. microphthalmus species groups. The parameral apices are indiscernible and asetose, like in the aedeagi of the C. longilinguis group, but the apical projection in ventral view is much larger than that in this group, and has an additional elongate median structure, a rod-like projection, lacking in the longilinguis group. In the C. hippopotamus and C. microphthalmus groups, the parameral apices are well developed and setose. In the two studied males the aedeagus shows small differences in the shape of the apex of the capsular region and in the shape of the apical projection. However, these features, together with lack of any external differences, can be attributed to interspecific variability.	en	Jałoszyński, Paweł (2025): The Cephenniini of China. X. New species and new records of Cephennodes Reitter of Yunnan (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa 5664 (1): 1-101, DOI: 10.11646/zootaxa.5664.1.1, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
