taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
AE5564907A7C52A8B1C1F60DE0039565.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383974	https://doi.org/10.3897/imafungus.16.154329.figure6	Figure 6. Basidiomata features. A, B. Pluteus albivillus (FJAU 66613). C – G. P. baishanzuensis (C, D. FJAU 66621, E – G. FJAU 66622). H – J. P. velutinus (H. FJAU 66619, I, J. FJAU 66620). K, L. P. longistriatus (FJAU 66596). A, B, K, L. Photos by Zheng-Xiang Qi. C – G. Photos by Rui-Peng Liu. H – J. Photos by Gu Rao. Scale bars: 1 cm.	Figure 6. Basidiomata features. A, B. Pluteus albivillus (FJAU 66613). C – G. P. baishanzuensis (C, D. FJAU 66621, E – G. FJAU 66622). H – J. P. velutinus (H. FJAU 66619, I, J. FJAU 66620). K, L. P. longistriatus (FJAU 66596). A, B, K, L. Photos by Zheng-Xiang Qi. C – G. Photos by Rui-Peng Liu. H – J. Photos by Gu Rao. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
AE5564907A7C52A8B1C1F60DE0039565.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383983	https://doi.org/10.3897/imafungus.16.154329.figure15	Figure 15. A. Macroscopic characteristics of Pluteus albivillus. B. Cheilocystidia. C. Basidiospores. D. Pleurocystidia. E. Basidia and Pleurocystidia. F. Caulocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 5 µm (C); 10 µm (B, D – F); 20 µm (G).	Figure 15. A. Macroscopic characteristics of Pluteus albivillus. B. Cheilocystidia. C. Basidiospores. D. Pleurocystidia. E. Basidia and Pleurocystidia. F. Caulocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 5 µm (C); 10 µm (B, D – F); 20 µm (G).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
AE5564907A7C52A8B1C1F60DE0039565.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383969	https://doi.org/10.3897/imafungus.16.154329.figure1	Figure 1. Phylogenetic tree of the sect. Hispidoderma of the genus Pluteus. The best tree from the ML and BI analysis of the LSU + ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram.	Figure 1. Phylogenetic tree of the sect. Hispidoderma of the genus Pluteus. The best tree from the ML and BI analysis of the LSU + ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
7904921AC94657A28A8558B6A405800F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383974	https://doi.org/10.3897/imafungus.16.154329.figure6	Figure 6. Basidiomata features. A, B. Pluteus albivillus (FJAU 66613). C – G. P. baishanzuensis (C, D. FJAU 66621, E – G. FJAU 66622). H – J. P. velutinus (H. FJAU 66619, I, J. FJAU 66620). K, L. P. longistriatus (FJAU 66596). A, B, K, L. Photos by Zheng-Xiang Qi. C – G. Photos by Rui-Peng Liu. H – J. Photos by Gu Rao. Scale bars: 1 cm.	Figure 6. Basidiomata features. A, B. Pluteus albivillus (FJAU 66613). C – G. P. baishanzuensis (C, D. FJAU 66621, E – G. FJAU 66622). H – J. P. velutinus (H. FJAU 66619, I, J. FJAU 66620). K, L. P. longistriatus (FJAU 66596). A, B, K, L. Photos by Zheng-Xiang Qi. C – G. Photos by Rui-Peng Liu. H – J. Photos by Gu Rao. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
7904921AC94657A28A8558B6A405800F.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383982	https://doi.org/10.3897/imafungus.16.154329.figure14	Figure 14. A. Macroscopic characteristics of Pluteus baishanzuensis. B. Caulocystidia. C. Basidiospores. D. Pleurocystidia. E. Basidia and Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – F); 20 µm (G).	Figure 14. A. Macroscopic characteristics of Pluteus baishanzuensis. B. Caulocystidia. C. Basidiospores. D. Pleurocystidia. E. Basidia and Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – F); 20 µm (G).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
4D37257476025EE0861B126C6A929FE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383986	https://doi.org/10.3897/imafungus.16.154329.figure18	Figure 18. Basidiomata features. A – C. Pluteus costatus (A, B. FJAU 66589, C. FJAU 66618). D, E. P. umbrosus (FJAU 66590). F – H. P. umbrosoides (FJAU 66615). I, J. P. granularis (FJAU 66612). A – C. Photos by Di-Zhe Guo. D, E. Photos by Jia-Jun Hu. C, F – H. Photos by Gu Rao. I, J. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	Figure 18. Basidiomata features. A – C. Pluteus costatus (A, B. FJAU 66589, C. FJAU 66618). D, E. P. umbrosus (FJAU 66590). F – H. P. umbrosoides (FJAU 66615). I, J. P. granularis (FJAU 66612). A – C. Photos by Di-Zhe Guo. D, E. Photos by Jia-Jun Hu. C, F – H. Photos by Gu Rao. I, J. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
4D37257476025EE0861B126C6A929FE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383987	https://doi.org/10.3897/imafungus.16.154329.figure19	Figure 19. A. Macroscopic characteristics of Pluteus costatus. B. Basidiospores. C. Caulocystidia. D. Pleurocystidia. E. Pileipellis elements. F. Cheilocystidia. Scale bars: 1 cm (A); 10 µm (B, D, F); 20 µm (C, E).	Figure 19. A. Macroscopic characteristics of Pluteus costatus. B. Basidiospores. C. Caulocystidia. D. Pleurocystidia. E. Pileipellis elements. F. Cheilocystidia. Scale bars: 1 cm (A); 10 µm (B, D, F); 20 µm (C, E).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
4D37257476025EE0861B126C6A929FE6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383971	https://doi.org/10.3897/imafungus.16.154329.figure3	Figure 3. Phylogenetic tree of the / umbrosus clade and / pantherinus lineage of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 3. Phylogenetic tree of the / umbrosus clade and / pantherinus lineage of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
AB5DA5C2493954F19A5787FE1FC22032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383986	https://doi.org/10.3897/imafungus.16.154329.figure18	Figure 18. Basidiomata features. A – C. Pluteus costatus (A, B. FJAU 66589, C. FJAU 66618). D, E. P. umbrosus (FJAU 66590). F – H. P. umbrosoides (FJAU 66615). I, J. P. granularis (FJAU 66612). A – C. Photos by Di-Zhe Guo. D, E. Photos by Jia-Jun Hu. C, F – H. Photos by Gu Rao. I, J. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	Figure 18. Basidiomata features. A – C. Pluteus costatus (A, B. FJAU 66589, C. FJAU 66618). D, E. P. umbrosus (FJAU 66590). F – H. P. umbrosoides (FJAU 66615). I, J. P. granularis (FJAU 66612). A – C. Photos by Di-Zhe Guo. D, E. Photos by Jia-Jun Hu. C, F – H. Photos by Gu Rao. I, J. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
AB5DA5C2493954F19A5787FE1FC22032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383990	https://doi.org/10.3897/imafungus.16.154329.figure22	Figure 22. A. Macroscopic characteristics of Pluteus granularis. B. Basidiospores. C. Pileipellis elements. D. Cheilocystidia. E. Caulocystidia. F. Basidia and Pleurocystidia. G. Pleurocystidia. Scale bars: 1 cm (A); 5 µm (B); 10 µm (D – G); 20 µm (C).	Figure 22. A. Macroscopic characteristics of Pluteus granularis. B. Basidiospores. C. Pileipellis elements. D. Cheilocystidia. E. Caulocystidia. F. Basidia and Pleurocystidia. G. Pleurocystidia. Scale bars: 1 cm (A); 5 µm (B); 10 µm (D – G); 20 µm (C).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
AB5DA5C2493954F19A5787FE1FC22032.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383971	https://doi.org/10.3897/imafungus.16.154329.figure3	Figure 3. Phylogenetic tree of the / umbrosus clade and / pantherinus lineage of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 3. Phylogenetic tree of the / umbrosus clade and / pantherinus lineage of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
FD3B3022FFBF513999B7E02A4B4293CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383975	https://doi.org/10.3897/imafungus.16.154329.figure7	Figure 7. Basidiomata features. A – C. Pluteus ultraputripiceae (A. FJAU 66611, B, C. FJAU 66594). D, E. P. hinnuleus (FJAU 66614). A – C. Photos by Zheng-Xiang Qi. D, E. Photos by Di-Zhe Guo. Scale bars: 1 cm.	Figure 7. Basidiomata features. A – C. Pluteus ultraputripiceae (A. FJAU 66611, B, C. FJAU 66594). D, E. P. hinnuleus (FJAU 66614). A – C. Photos by Zheng-Xiang Qi. D, E. Photos by Di-Zhe Guo. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
FD3B3022FFBF513999B7E02A4B4293CA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383981	https://doi.org/10.3897/imafungus.16.154329.figure13	Figure 13. A. Macroscopic characteristics of Pluteus hinnuleus. B. Basidiospores. C. Caulocystidia. D. Basidia and Pleurocystidia. E. Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – F); 20 µm (G).	Figure 13. A. Macroscopic characteristics of Pluteus hinnuleus. B. Basidiospores. C. Caulocystidia. D. Basidia and Pleurocystidia. E. Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – F); 20 µm (G).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
DDE8ACF82F015C818DA25E92F33098C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383973	https://doi.org/10.3897/imafungus.16.154329.figure5	Figure 5. Basidiomata features. A – D. Pluteus tenuipileus (A, B. FJAU 66591, C, D. FJAU 66592). E – G. P. jilinensis (FJAU 66616). H – I. P. spaniophyllus (FJAU 66593). J, K. P. aff. semibulbosus (FJAU 66617). A – K. Photos by Gu Rao. Scale bars: 1 cm.	Figure 5. Basidiomata features. A – D. Pluteus tenuipileus (A, B. FJAU 66591, C, D. FJAU 66592). E – G. P. jilinensis (FJAU 66616). H – I. P. spaniophyllus (FJAU 66593). J, K. P. aff. semibulbosus (FJAU 66617). A – K. Photos by Gu Rao. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
DDE8ACF82F015C818DA25E92F33098C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383978	https://doi.org/10.3897/imafungus.16.154329.figure10	Figure 10. A. Macroscopic characteristics of Pluteus jilinensis. B. Caulocystidia. C. Basidiospores. D. Pleurocystidia. E. Basidia and Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 5 µm (C); 10 µm (B, D – F); 20 µm (G).	Figure 10. A. Macroscopic characteristics of Pluteus jilinensis. B. Caulocystidia. C. Basidiospores. D. Pleurocystidia. E. Basidia and Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 5 µm (C); 10 µm (B, D – F); 20 µm (G).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
75A12E12838A5BDFBB69E35B894B16B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383993	https://doi.org/10.3897/imafungus.16.154329.figure25	Figure 25. Basidiomata features. A – E. Pluteus ussuriensis (A, B. FJAU 66576, C, D. FJAU 66577, E. FJAU 66578). F – H. P. leoninus (F. FJAU 66581, G, H. FJAU 66580). A – D. Photos by Gu Rao. E. Photos by Ming-Hao Liu. F. Photos by Di-Zhe Guo. G, H. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	Figure 25. Basidiomata features. A – E. Pluteus ussuriensis (A, B. FJAU 66576, C, D. FJAU 66577, E. FJAU 66578). F – H. P. leoninus (F. FJAU 66581, G, H. FJAU 66580). A – D. Photos by Gu Rao. E. Photos by Ming-Hao Liu. F. Photos by Di-Zhe Guo. G, H. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
75A12E12838A5BDFBB69E35B894B16B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383997	https://doi.org/10.3897/imafungus.16.154329.figure29	Figure 29. A. Macroscopic characteristics of Pluteus leoninus. B. Pileipellis elements. C. Cheilocystidia. D. Basidiospores. E. Pleurocystidia. Scale bars: 1 cm (A); 10 µm (B, C, E); 20 µm (D).	Figure 29. A. Macroscopic characteristics of Pluteus leoninus. B. Pileipellis elements. C. Cheilocystidia. D. Basidiospores. E. Pleurocystidia. Scale bars: 1 cm (A); 10 µm (B, C, E); 20 µm (D).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
75A12E12838A5BDFBB69E35B894B16B3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383972	https://doi.org/10.3897/imafungus.16.154329.figure4	Figure 4. Phylogenetic tree of the / leoninus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 4. Phylogenetic tree of the / leoninus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
A75D4DD24403538C86B0E16639CA1164.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383974	https://doi.org/10.3897/imafungus.16.154329.figure6	Figure 6. Basidiomata features. A, B. Pluteus albivillus (FJAU 66613). C – G. P. baishanzuensis (C, D. FJAU 66621, E – G. FJAU 66622). H – J. P. velutinus (H. FJAU 66619, I, J. FJAU 66620). K, L. P. longistriatus (FJAU 66596). A, B, K, L. Photos by Zheng-Xiang Qi. C – G. Photos by Rui-Peng Liu. H – J. Photos by Gu Rao. Scale bars: 1 cm.	Figure 6. Basidiomata features. A, B. Pluteus albivillus (FJAU 66613). C – G. P. baishanzuensis (C, D. FJAU 66621, E – G. FJAU 66622). H – J. P. velutinus (H. FJAU 66619, I, J. FJAU 66620). K, L. P. longistriatus (FJAU 66596). A, B, K, L. Photos by Zheng-Xiang Qi. C – G. Photos by Rui-Peng Liu. H – J. Photos by Gu Rao. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
A75D4DD24403538C86B0E16639CA1164.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383985	https://doi.org/10.3897/imafungus.16.154329.figure17	Figure 17. A. Macroscopic characteristics of Pluteus longistriatus. B. Caulocystidia. C. Basidiospores. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (C – E); 20 µm (B, F).	Figure 17. A. Macroscopic characteristics of Pluteus longistriatus. B. Caulocystidia. C. Basidiospores. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (C – E); 20 µm (B, F).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
A75D4DD24403538C86B0E16639CA1164.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383970	https://doi.org/10.3897/imafungus.16.154329.figure2	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
7D4C12CF839A5D779D0C02CF8DB52A36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383991	https://doi.org/10.3897/imafungus.16.154329.figure23	Figure 23. Basidiomata features. A – K. Pluteus piceicola (A, B. FJAU 66574, C. FJAU 66569, D, E. FJAU 66575, F, G. FJAU 66571, H, I. FJAU 66573, J, K. FJAU 66572). A – K. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	Figure 23. Basidiomata features. A – K. Pluteus piceicola (A, B. FJAU 66574, C. FJAU 66569, D, E. FJAU 66575, F, G. FJAU 66571, H, I. FJAU 66573, J, K. FJAU 66572). A – K. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
7D4C12CF839A5D779D0C02CF8DB52A36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383992	https://doi.org/10.3897/imafungus.16.154329.figure24	Figure 24. Basidiomata features. A – M. Pluteus piceicola (A, B. FJAU 66603, C, M. FJAU 66607, D, G. FJAU 66605, E, F. FJAU 66606, H, I. FJAU 66608, J. FJAU 66610, K, L. FJAU 66609). A – M. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	Figure 24. Basidiomata features. A – M. Pluteus piceicola (A, B. FJAU 66603, C, M. FJAU 66607, D, G. FJAU 66605, E, F. FJAU 66606, H, I. FJAU 66608, J. FJAU 66610, K, L. FJAU 66609). A – M. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
7D4C12CF839A5D779D0C02CF8DB52A36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383995	https://doi.org/10.3897/imafungus.16.154329.figure27	Figure 27. A. Macroscopic characteristics of Pluteus piceicola. B. Basidiospores. C. Pleurocystidia. D. Basidia and Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – E); 20 µm (F).	Figure 27. A. Macroscopic characteristics of Pluteus piceicola. B. Basidiospores. C. Pleurocystidia. D. Basidia and Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – E); 20 µm (F).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
7D4C12CF839A5D779D0C02CF8DB52A36.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383972	https://doi.org/10.3897/imafungus.16.154329.figure4	Figure 4. Phylogenetic tree of the / leoninus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 4. Phylogenetic tree of the / leoninus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
AE2047D258B65C19B2D85A8C69EA7448.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383973	https://doi.org/10.3897/imafungus.16.154329.figure5	Figure 5. Basidiomata features. A – D. Pluteus tenuipileus (A, B. FJAU 66591, C, D. FJAU 66592). E – G. P. jilinensis (FJAU 66616). H – I. P. spaniophyllus (FJAU 66593). J, K. P. aff. semibulbosus (FJAU 66617). A – K. Photos by Gu Rao. Scale bars: 1 cm.	Figure 5. Basidiomata features. A – D. Pluteus tenuipileus (A, B. FJAU 66591, C, D. FJAU 66592). E – G. P. jilinensis (FJAU 66616). H – I. P. spaniophyllus (FJAU 66593). J, K. P. aff. semibulbosus (FJAU 66617). A – K. Photos by Gu Rao. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
AE2047D258B65C19B2D85A8C69EA7448.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383979	https://doi.org/10.3897/imafungus.16.154329.figure11	Figure 11. A. Macroscopic characteristics of Pluteus aff. semibulbosus. B. Basidiospores. C. Cheilocystidia. D. Basidia and Pleurocystidia. E. Pleurocystidia. F. Caulocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 5 µm (B); 10 µm (C – G).	Figure 11. A. Macroscopic characteristics of Pluteus aff. semibulbosus. B. Basidiospores. C. Cheilocystidia. D. Basidia and Pleurocystidia. E. Pleurocystidia. F. Caulocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 5 µm (B); 10 µm (C – G).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
AE2047D258B65C19B2D85A8C69EA7448.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383970	https://doi.org/10.3897/imafungus.16.154329.figure2	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
B9E5F9BFD43B59F8986A3CE5B4CC9C66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383973	https://doi.org/10.3897/imafungus.16.154329.figure5	Figure 5. Basidiomata features. A – D. Pluteus tenuipileus (A, B. FJAU 66591, C, D. FJAU 66592). E – G. P. jilinensis (FJAU 66616). H – I. P. spaniophyllus (FJAU 66593). J, K. P. aff. semibulbosus (FJAU 66617). A – K. Photos by Gu Rao. Scale bars: 1 cm.	Figure 5. Basidiomata features. A – D. Pluteus tenuipileus (A, B. FJAU 66591, C, D. FJAU 66592). E – G. P. jilinensis (FJAU 66616). H – I. P. spaniophyllus (FJAU 66593). J, K. P. aff. semibulbosus (FJAU 66617). A – K. Photos by Gu Rao. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
B9E5F9BFD43B59F8986A3CE5B4CC9C66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383977	https://doi.org/10.3897/imafungus.16.154329.figure9	Figure 9. A. Macroscopic characteristics of Pluteus spaniophyllus. B. Basidiospores. C. Caulocystidia. D. Basidia and Pleurocystidia. E. Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 0.5 cm (A); 10 µm (B – F); 20 µm (G).	Figure 9. A. Macroscopic characteristics of Pluteus spaniophyllus. B. Basidiospores. C. Caulocystidia. D. Basidia and Pleurocystidia. E. Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 0.5 cm (A); 10 µm (B – F); 20 µm (G).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
B9E5F9BFD43B59F8986A3CE5B4CC9C66.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383970	https://doi.org/10.3897/imafungus.16.154329.figure2	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
174711C0FCBB5BF083A181F42DBFEA0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383973	https://doi.org/10.3897/imafungus.16.154329.figure5	Figure 5. Basidiomata features. A – D. Pluteus tenuipileus (A, B. FJAU 66591, C, D. FJAU 66592). E – G. P. jilinensis (FJAU 66616). H – I. P. spaniophyllus (FJAU 66593). J, K. P. aff. semibulbosus (FJAU 66617). A – K. Photos by Gu Rao. Scale bars: 1 cm.	Figure 5. Basidiomata features. A – D. Pluteus tenuipileus (A, B. FJAU 66591, C, D. FJAU 66592). E – G. P. jilinensis (FJAU 66616). H – I. P. spaniophyllus (FJAU 66593). J, K. P. aff. semibulbosus (FJAU 66617). A – K. Photos by Gu Rao. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
174711C0FCBB5BF083A181F42DBFEA0D.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383976	https://doi.org/10.3897/imafungus.16.154329.figure8	Figure 8. A. Macroscopic characteristics of Pluteus tenuipileus. B. Basidiospores. C. Caulocystidia. D. Cheilocystidia. E. Pileipellis elements. F. Pleurocystidia. Scale bars: 0.5 cm (A); 10 µm (B, F); 20 µm (C, D, E).	Figure 8. A. Macroscopic characteristics of Pluteus tenuipileus. B. Basidiospores. C. Caulocystidia. D. Cheilocystidia. E. Pileipellis elements. F. Pleurocystidia. Scale bars: 0.5 cm (A); 10 µm (B, F); 20 µm (C, D, E).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
8F3C092620B75652B6918F3DE7084A52.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383975	https://doi.org/10.3897/imafungus.16.154329.figure7	Figure 7. Basidiomata features. A – C. Pluteus ultraputripiceae (A. FJAU 66611, B, C. FJAU 66594). D, E. P. hinnuleus (FJAU 66614). A – C. Photos by Zheng-Xiang Qi. D, E. Photos by Di-Zhe Guo. Scale bars: 1 cm.	Figure 7. Basidiomata features. A – C. Pluteus ultraputripiceae (A. FJAU 66611, B, C. FJAU 66594). D, E. P. hinnuleus (FJAU 66614). A – C. Photos by Zheng-Xiang Qi. D, E. Photos by Di-Zhe Guo. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
8F3C092620B75652B6918F3DE7084A52.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383980	https://doi.org/10.3897/imafungus.16.154329.figure12	Figure 12. A. Macroscopic characteristics of Pluteus ultraputripiceae. B. Caulocystidia. C. Basidiospores. D. Basidia. E. Cheilocystidia. F. Pleurocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – G).	Figure 12. A. Macroscopic characteristics of Pluteus ultraputripiceae. B. Caulocystidia. C. Basidiospores. D. Basidia. E. Cheilocystidia. F. Pleurocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – G).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
8F3C092620B75652B6918F3DE7084A52.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383970	https://doi.org/10.3897/imafungus.16.154329.figure2	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
FD56B704813F5616A2FE299299C369EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383986	https://doi.org/10.3897/imafungus.16.154329.figure18	Figure 18. Basidiomata features. A – C. Pluteus costatus (A, B. FJAU 66589, C. FJAU 66618). D, E. P. umbrosus (FJAU 66590). F – H. P. umbrosoides (FJAU 66615). I, J. P. granularis (FJAU 66612). A – C. Photos by Di-Zhe Guo. D, E. Photos by Jia-Jun Hu. C, F – H. Photos by Gu Rao. I, J. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	Figure 18. Basidiomata features. A – C. Pluteus costatus (A, B. FJAU 66589, C. FJAU 66618). D, E. P. umbrosus (FJAU 66590). F – H. P. umbrosoides (FJAU 66615). I, J. P. granularis (FJAU 66612). A – C. Photos by Di-Zhe Guo. D, E. Photos by Jia-Jun Hu. C, F – H. Photos by Gu Rao. I, J. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
FD56B704813F5616A2FE299299C369EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383989	https://doi.org/10.3897/imafungus.16.154329.figure21	Figure 21. A. Macroscopic characteristics of Pluteus umbrosoides. B. Basidiospores. C. Basidia and Pleurocystidia. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – E); 20 µm (F).	Figure 21. A. Macroscopic characteristics of Pluteus umbrosoides. B. Basidiospores. C. Basidia and Pleurocystidia. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – E); 20 µm (F).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
FD56B704813F5616A2FE299299C369EB.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383971	https://doi.org/10.3897/imafungus.16.154329.figure3	Figure 3. Phylogenetic tree of the / umbrosus clade and / pantherinus lineage of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 3. Phylogenetic tree of the / umbrosus clade and / pantherinus lineage of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
B6C6604DB203507D818DD2DA1D675417.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383986	https://doi.org/10.3897/imafungus.16.154329.figure18	Figure 18. Basidiomata features. A – C. Pluteus costatus (A, B. FJAU 66589, C. FJAU 66618). D, E. P. umbrosus (FJAU 66590). F – H. P. umbrosoides (FJAU 66615). I, J. P. granularis (FJAU 66612). A – C. Photos by Di-Zhe Guo. D, E. Photos by Jia-Jun Hu. C, F – H. Photos by Gu Rao. I, J. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	Figure 18. Basidiomata features. A – C. Pluteus costatus (A, B. FJAU 66589, C. FJAU 66618). D, E. P. umbrosus (FJAU 66590). F – H. P. umbrosoides (FJAU 66615). I, J. P. granularis (FJAU 66612). A – C. Photos by Di-Zhe Guo. D, E. Photos by Jia-Jun Hu. C, F – H. Photos by Gu Rao. I, J. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
B6C6604DB203507D818DD2DA1D675417.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383988	https://doi.org/10.3897/imafungus.16.154329.figure20	Figure 20. A. Macroscopic characteristics of Pluteus umbrosus. B. Basidiospores. C. Caulocystidia. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (C, D, E); 20 µm (B, F).	Figure 20. A. Macroscopic characteristics of Pluteus umbrosus. B. Basidiospores. C. Caulocystidia. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (C, D, E); 20 µm (B, F).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
B6C6604DB203507D818DD2DA1D675417.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383971	https://doi.org/10.3897/imafungus.16.154329.figure3	Figure 3. Phylogenetic tree of the / umbrosus clade and / pantherinus lineage of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 3. Phylogenetic tree of the / umbrosus clade and / pantherinus lineage of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
22D64D1053745A6488B08770A4A2E274.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383993	https://doi.org/10.3897/imafungus.16.154329.figure25	Figure 25. Basidiomata features. A – E. Pluteus ussuriensis (A, B. FJAU 66576, C, D. FJAU 66577, E. FJAU 66578). F – H. P. leoninus (F. FJAU 66581, G, H. FJAU 66580). A – D. Photos by Gu Rao. E. Photos by Ming-Hao Liu. F. Photos by Di-Zhe Guo. G, H. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	Figure 25. Basidiomata features. A – E. Pluteus ussuriensis (A, B. FJAU 66576, C, D. FJAU 66577, E. FJAU 66578). F – H. P. leoninus (F. FJAU 66581, G, H. FJAU 66580). A – D. Photos by Gu Rao. E. Photos by Ming-Hao Liu. F. Photos by Di-Zhe Guo. G, H. Photos by Zheng-Xiang Qi. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
22D64D1053745A6488B08770A4A2E274.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383996	https://doi.org/10.3897/imafungus.16.154329.figure28	Figure 28. A. Macroscopic characteristics of Pluteus ussuriensis. B. Basidiospores. C. Basidia and Pleurocystidia. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – E); 20 µm (F).	Figure 28. A. Macroscopic characteristics of Pluteus ussuriensis. B. Basidiospores. C. Basidia and Pleurocystidia. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (B – E); 20 µm (F).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
22D64D1053745A6488B08770A4A2E274.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383972	https://doi.org/10.3897/imafungus.16.154329.figure4	Figure 4. Phylogenetic tree of the / leoninus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 4. Phylogenetic tree of the / leoninus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
736E6338A1BE5868A96CAF3A6F8665C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383994	https://doi.org/10.3897/imafungus.16.154329.figure26	Figure 26. Basidiomata features. A – L. Pluteus variabilicolor (A, B. FJAU 66585, C, D. FJAU 66584, E, F. FJAU 66583, G, H. FJAU 66586, I. FJAU 66588, J – L. FJAU 66623). A – H. Photos by Ya-Jie Liu. I – L. Photos by Li-Bo Wang. Scale bars: 1 cm.	Figure 26. Basidiomata features. A – L. Pluteus variabilicolor (A, B. FJAU 66585, C, D. FJAU 66584, E, F. FJAU 66583, G, H. FJAU 66586, I. FJAU 66588, J – L. FJAU 66623). A – H. Photos by Ya-Jie Liu. I – L. Photos by Li-Bo Wang. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
736E6338A1BE5868A96CAF3A6F8665C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383998	https://doi.org/10.3897/imafungus.16.154329.figure30	Figure 30. A. Macroscopic characteristics of Pluteus variabilicolor. B. Basidiospores. C. Caulocystidia. D. Pleurocystidia. E. Basidia and Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (C – F); 20 µm (B, G).	Figure 30. A. Macroscopic characteristics of Pluteus variabilicolor. B. Basidiospores. C. Caulocystidia. D. Pleurocystidia. E. Basidia and Pleurocystidia. F. Cheilocystidia. G. Pileipellis elements. Scale bars: 1 cm (A); 10 µm (C – F); 20 µm (B, G).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
736E6338A1BE5868A96CAF3A6F8665C3.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383972	https://doi.org/10.3897/imafungus.16.154329.figure4	Figure 4. Phylogenetic tree of the / leoninus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 4. Phylogenetic tree of the / leoninus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
9E5C6E2BB94B585383ECD6706F0E8163.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383974	https://doi.org/10.3897/imafungus.16.154329.figure6	Figure 6. Basidiomata features. A, B. Pluteus albivillus (FJAU 66613). C – G. P. baishanzuensis (C, D. FJAU 66621, E – G. FJAU 66622). H – J. P. velutinus (H. FJAU 66619, I, J. FJAU 66620). K, L. P. longistriatus (FJAU 66596). A, B, K, L. Photos by Zheng-Xiang Qi. C – G. Photos by Rui-Peng Liu. H – J. Photos by Gu Rao. Scale bars: 1 cm.	Figure 6. Basidiomata features. A, B. Pluteus albivillus (FJAU 66613). C – G. P. baishanzuensis (C, D. FJAU 66621, E – G. FJAU 66622). H – J. P. velutinus (H. FJAU 66619, I, J. FJAU 66620). K, L. P. longistriatus (FJAU 66596). A, B, K, L. Photos by Zheng-Xiang Qi. C – G. Photos by Rui-Peng Liu. H – J. Photos by Gu Rao. Scale bars: 1 cm.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
9E5C6E2BB94B585383ECD6706F0E8163.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383984	https://doi.org/10.3897/imafungus.16.154329.figure16	Figure 16. A. Macroscopic characteristics of Pluteus velutinus. B. Caulocystidia. C. Basidiospores. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 5 µm (C); 10 µm (B, D, E); 20 µm (F).	Figure 16. A. Macroscopic characteristics of Pluteus velutinus. B. Caulocystidia. C. Basidiospores. D. Pleurocystidia. E. Cheilocystidia. F. Pileipellis elements. Scale bars: 1 cm (A); 5 µm (C); 10 µm (B, D, E); 20 µm (F).	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
9E5C6E2BB94B585383ECD6706F0E8163.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://binary.pensoft.net/fig/1383970	https://doi.org/10.3897/imafungus.16.154329.figure2	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	Figure 2. Phylogenetic tree of the / plautus clade of the sect. Hispidoderma. The best tree from the ML and BI analysis of the ITS + tef 1 dataset. The two values of internal nodes respectively represent the Maximum Likelihood bootstrap (MLBP) / Bayesian posterior probability (BIPP). Nodes with a bootstrap probability (BP) ≥ 80 and a Bayesian posterior probability (PP) ≥ 0.80 are indicated in the phylogram. This study species is in bold and red font.	2025-07-29	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu		Zenodo	biologists	Qi, Zheng-Xiang;Wang, Li-Bo;Qian, Ke-Qing;Shi, Li-Li;Hu, Jia-Jun;Tuo, Yong-Lan;Rao, Gu;Samwel Jacob, Muharagi;Liu, Rui-Peng;Liu, Ming-Hao;Guo, Di-Zhe;Liu, Ya-Jie;Zhang, Bo;Li, Xiao;Li, Yu			
