taxonID	type	description	language	source
626F8975FFDEFFA929C4FB16E8A5FCFE.taxon	description	Local Names: — Marasibat, salik-salik (Isneg). Phenology: — Villaria lanceolata has been observed to be flowering and fruiting from November to February, and April to June.	en	Alejandro, Grecebio Jonathan D., Meve, Ulrich (2016): Recollection of former Randia species, recognition of a rheophytic species of the Philippine endemic Villaria (Octotropideae, Rubiaceae). Phytotaxa 253 (2): 171-175, DOI: 10.11646/phytotaxa.253.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.253.2.8
626F8975FFDEFFA929C4FB16E8A5FCFE.taxon	distribution	Distribution and Habitat: — Villaria lanceolata has been found in Luzon, restricted to the provinces of Apayao, Bulacan and Cagayan. Its habit (tough and flexible stems, lanceolate and glossy willow-like leaves) and the streambed habitat characterize this species as a rheophyte sensu Van Steenis (1952). Villaria lanceolata has been typically collected on sandy substrate to stony cliffs along riverbanks and streams subject to overflow during rainy seasons. Species were found also to thrive as well on plant litter and soil deposits on other trees and plants but never epiphytic. Its known altitudinal extent is 30 – 130 m asl.	en	Alejandro, Grecebio Jonathan D., Meve, Ulrich (2016): Recollection of former Randia species, recognition of a rheophytic species of the Philippine endemic Villaria (Octotropideae, Rubiaceae). Phytotaxa 253 (2): 171-175, DOI: 10.11646/phytotaxa.253.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.253.2.8
626F8975FFDEFFA929C4FB16E8A5FCFE.taxon	discussion	Discussion: — The specific epithet “ stenophylla ”, supposedly having priority over “ lanceolata ”, cannot be used because of the earlier homonym Randia stenophylla K. Krause (1909: 140; nowadays regarded as synonym of Rothmannia urcelliformis (Schweinf. ex Hiern) Bullock ex Robyns); thus, making R. stenophylla Merr. illegitimate. The cited type specimens of Randia lanceolata (Merrill 1913: 57) and R. stenophylla (Merrill 1910: 245 – 246) were apparently destroyed in PNH during the World War II. There were no isotypes distributed for R. lanceolata; thus, E. Fenix 28123 collection of a fruiting branch found in US herbarium is designated as the neotype. For R. stenophylla, the only isotype located is available at US, and is here designated as the lectotype. Villaria lanceolata is rather unique from the rest of the Villaria species by its rheophytic habit, narrowly lanceolate leaves, bright red fruits when ripe and the presence of parallel ridges in the fibrous testa of seeds. The locals in the province of Apayao named the plant “ salik-salik ” in reference to salicylic acid. Traditionally, they crushed the leaves of V. lanceolata and applied to body parts with fungal infections. Additional Specimens Examined: — PHILIPPINES: Apayao, Pudtol, Nagan River, 16 November 2009, B. O. C. Lemana 25080 (L, K, US, PNH, USTH), 25081 (L, K, US, PNH, USTH), 25082 (L, US, USTH), 25083 (K, USTH); Apayao, Pudtol, Nagan River, 29 January 2010, B. O. C Lemana 25084 (L, PNH, USTH); Bulacan, Norzagaray, Angat Dam, May 2010, A. Fernandez et al. 25092 (PNH, USTH <2 sheets>), July 2010, J. Osorio et al. 25093 (PNH, USTH <3 sheets>). IUCN Red List category (2012): — Endangered [EN; C 2 ai], population size is estimated to number fewer than 2,500 mature individuals. A continuing decline is projected and no subpopulation is estimated to contain more than 250 mature individuals or even more than 100. Some areas occupied by V. lanceolata are fragmented forests caused by logging activities; others are threatened by human land use such as in Bulacan, Norzagaray and Apayao, Pudtol. The populations in riverbanks may be possibly swipe off during the rainy seasons or several typhoons in a year.	en	Alejandro, Grecebio Jonathan D., Meve, Ulrich (2016): Recollection of former Randia species, recognition of a rheophytic species of the Philippine endemic Villaria (Octotropideae, Rubiaceae). Phytotaxa 253 (2): 171-175, DOI: 10.11646/phytotaxa.253.2.8, URL: http://dx.doi.org/10.11646/phytotaxa.253.2.8
