identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
5C0C8799FFEE6A0CC3CCF916FC94D3B4.text	5C0C8799FFEE6A0CC3CCF916FC94D3B4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceutorhynchus Germar 1823	<div><p>Genus Ceutorhynchus Germar, 1823</p><p>A key to Ceutorhynchus species associated with Draba nemorosa</p><p>1 (4). Antennal funicle six-segmented. Claws simple, free, fine, short, moderately divergent (at an angle about 45°). Body small, 1.6–1.9 mm (Figs 1, 2). Rostrum, antennae, and at least tibiae and tarsi reddish brown, pronotum and elytra black, often with brownish tint. Dorsal vestiture moderately dense, uniform, subrecumbent to semi-erect, formed by narrow parallel-sided whitish scales. Elytral striae bare, intervals not wider than striae, flat or weakly convex. Hind wings non-functional, strongly reduced. Aedeagus angularly narrowed apically (Figs 11, 12).</p><p>2 (3). Elytra narrow, 1.23–1.27 times as long as wide, slightly rounded at sides, flattened dorsally (Fig. 1). Elytral vestiture subrecumbent. Aedeagus as in Figs 11, 19. Eastern Transcaucasia, Northeastern Turkey .......................... C. gemuricus</p><p>3 (2). Elytra wider, oval, 1.08–1.15 times as long as wide, noticeably rounded laterally and convex dorsally (Figs 2). Elytral vestiture semi-erect. Aedeagus as in Fig. 12, 20. Mountain Caucasus, middle Volga area, Kazakhstan, East Siberia: West Sayan Mts, Buryatia...................... C. kipchak</p><p>4 (1). Antennal funicle seven-segmented. Claws simple or toothed, free or closely approximate at base, thin or rather stout, weakly to moderately divergent. Body on overage larger, 1.5–2.5 mm. Body predominantly black, only tibiae may be varyingly dark brown; if legs, apical margin of pronotum and apex of elytra reddish brown, then body robust, claws finely dentate, elytra with white scutellar spot and mottled vestiture of narrow white and brownish scales, and middle tibia of female finely mucronate, anal ventrite deeply depressed medio-apically. Dorsal vestiture sparse to moderately dense, recumbent to semi-erect, uniform and formed by narrow parallel-sided or acuminate white scales, or predominantly by semi-erect dark setae with an admixture of white setiform scales. Elytral striae bare or with white narrow scales, intervals usually wider than striae, flat or barely convex. Hind wings not conspicuously reduced. Aedeagus of varying structure.</p><p>5 (8). Larger, 1.8–2.5 mm. Dorsal vestiture quite conspicuous, formed by recumbent uniform greyish narrow truncate scales (then claws distinctly dentate, apical margin of 4th abdominal ventrite entire, and male fore tibia finely mucronate) or by semi-erect dark setae with an admixture of white setiform scales (then claws simple, apical margin of 4th abdominal ventrite shallowly excised medially and slightly raised). Elytral striae with conspicuous narrow white scales. Aedeagus angularly narrowing apically, with sides moderately sclerotised.</p><p>6 (7). Dorsal vestiture rather dense, formed by recumbent uniform greyish narrow truncate scales; no trace of white stripes on pronotum or white scutellar spot present (Fig. 3); sides of mesothorax without condensed white scales on apices of mesepimera. Claws distinctly dentate, apical margin of 4th abdominal ventrite entire, male fore tibia finely mucronate; aedeagus sharpened apically (Fig. 13). East Siberia, Northeastern Russia (Magadan Prov., Chukchi Autonomous Area), Mongolia, Northern China ............................ C. seniculus</p><p>7 (6). Dorsal vestitureformedbysubrecumbenttosemierect dark and white very narrow parallel-sided scales; pronotum with narrow median and lateral white stripes; elytra with ill-defined but conspicuous white scutellar spot (Fig. 4); sides of mesothorax with white scales conspicuously condensed on apices of mesepimera. Claws simple; apical margin of male 4th abdominal ventrite shallowly excised medially and slightly raised; male fore tibia non-mucronate; aedeagus narrowly rounded (Fig. 14) or shallowly excised apically. Southern East Siberia northward to Yakutia, eastern half of northern Mongolia ................ C. klementzorum 8 (5). Smaller, 1.5–2.4 mm. Dorsal vestiture fine, sparse and inconspicuous, elytral striae bare, or elytra with ill-defined white scutellar spot and mottled vestiture of narrow white and brownish scales on intervals and striae with conspicuous lanceolate white scales; 4th abdominal ventrite in male with apical margin not excised medially. Male fore tibia without mucro; female middle tibia finely mucronate, or claws very closely approximate and armed with clearly visible appendage in basal half. 9 (10). Claws very closely approximate and armed with a well-developed tooth in basal half. Rostrum slen- der. Body and legs black, legs may be dark brown. Sides of thorax densely clothed with broad-lanceolate scales separated by less than own width (Fig. 5). Middle tibia of female non-mucronate. Elytral striae bare, intervals with sparse narrow, parallel-sided white scales. Aedeagus (Figs 15, 21) with narrowly rounded, weakly attenuate apex. Southern slope of West Sayan, East Tuva Plateau, Central Yakutia (new record)............................................ C. sp. [? C. unguicularis]</p><p>10 (9). Claws weakly diverging, simple or with broad tooth at base [then middle tibia of female finely mucronate, scales on elytral intervals very narrow, pointed apically, legs and partly elytra may be reddish brown (in C. querceti)]. Rostrum wider. Middle tibia of female finely mucronate. Aedeagus with subtruncate or shallowly emarginate apex (Figs 18, 22, 23).</p><p>11 (12). Apical margin of pronotum, legs and at least apical part of elytra reddish brown. Elytra with white scutellar spot and mottled vestiture of narrow white and brownish scales, elytral striae scaled similarly to the intervals (Fig. 6). Claws dentate at base. Anal ventrite of male very deeply depressed medio-posteriorly, apical margin behind the depression glabrous. Aedeagus as in Fig. 22. Holarctic species, mainly on Rorippa palustris, but found in southern Tuva on Draba nemorosa or D. sibirica .................................... C. querceti</p><p>12 (11). Body and legs black, legs occasionally brownish. Elytral vestiture uniform, composed of pointed or parallel-sided very narrow white scales.</p><p>13 (14). Claws with short broad tooth at base. Tibiae not paler than femora. Elytral intervals with sparse hair-like pointed scales, striae with similar but shorter scales. White scales on sides of pronotum, meso- and metathorax sparse, separated by not less than own widths; sides of pronotum without conspicuous white stripes. Elytra wider at base and more strongly narrowing toward apex (Fig. 7). Aedeagus as in Fig. 18. Body length 1.7–2.1 mm. Southern East Siberia east of Lake Baikal, eastern Mongolia, Korean Peninsula. On Draba nemorosa, probably also on Cardamine spp. .... C. dauricus</p><p>14 (13). Claws simple. Tibiae often brown. Elytral intervals with sparse parallel-sided, truncate apically white scales; striae bare or with inconspicuous fine hairs. Sides of pronotum without conspicuous white stripes, meso- and metathorax densely clothed with broad white scales separated by less than own widths. Elytra with humeri less convex, widest usually slightly behind them, less rounded at sides and less strongly narrowing toward apex (Fig. 8). Aedeagus as in Fig. 23. Body length 1.6–2.4 mm. Euro-Siberian species, distributed from West Europe to Irkutsk in the east. Mostly on Cardamine spp., but found on Draba nemorosa in Irkutsk and on Barbarea arquata south-east of Irkutsk ....................................... C. cochleariae</p></div>	https://treatment.plazi.org/id/5C0C8799FFEE6A0CC3CCF916FC94D3B4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Korotyaev, B. A.	Korotyaev, B. A. (2020): A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae). Zoosystematica Rossica (China) 29 (2): 353-367, DOI: 10.31610/zsr/2020.29.2.353, URL: https://doi.org/10.31610/zsr/2020.29.2.353
5C0C8799FFEB6A0CC064FF68FC4DD63F.text	5C0C8799FFEB6A0CC064FF68FC4DD63F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceutorhynchus gemuricus Korotyaev 1997	<div><p>Ceutorhynchus gemuricus Korotyaev, 1997</p><p>(Figs 1, 11, 19)</p><p>Distribution. Armenia, Azerbaijan, Northeastern Turkey (Korotyaev, 1997; Colonnelli, 2004).</p><p>Hosts. Draba nemorosa in the Nakhchivan Autonomous Republic of Azerbaijan (Korotyaev, 1997); in Northeastern Turkey I swept a few specimens in the habitats where no D. nemorosa was noticed.</p></div>	https://treatment.plazi.org/id/5C0C8799FFEB6A0CC064FF68FC4DD63F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Korotyaev, B. A.	Korotyaev, B. A. (2020): A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae). Zoosystematica Rossica (China) 29 (2): 353-367, DOI: 10.31610/zsr/2020.29.2.353, URL: https://doi.org/10.31610/zsr/2020.29.2.353
5C0C8799FFEB6A0CC064FDEEFB04D3B6.text	5C0C8799FFEB6A0CC064FDEEFB04D3B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceutorhynchus kipchak Korotyaev 1996	<div><p>Ceutorhynchus kipchak Korotyaev, 1996</p><p>(Figs 2, 12, 20)</p><p>Distribution. Armenia, eastern European Russia, southern slope of the West Sayan Mts in the south of Krasnoyarsk Terr. and in Tuva, East Tuva Plateau, Buryatia, Kazakhstan: Caspian Lowland (Khruleva et al., 2011, 2012), East Kazakhstan Prov. (Korotyaev, 2019).</p><p>Hosts. Draba nemorosa; in Tuva also on D. sibirica (Korotyaev, 2019) .</p><p>Ceutorhynchus seniculus Brisout de Barneville, 1883</p><p>(Figs 3, 13)</p><p>Distribution. South of East Siberia, Northeastern Yakutia, Magadan Prov. (Korotyaev, 1980; as C. changaicus Schultze, 1898), Chukchi Autonomous Area (Berman et al., 2002), Mongolia [Schultze, 1898 (as C. changaicus); Voss (1967; as C. gobiensis), Korotyaev (1980; as C. changaicus)], Northern China (Alonso-Zarazaga et al., 2017).</p><p>Hosts. Several species of Brassicaceae [often Erysimum flavum and E. cheiranthoides, occasionally Smelowskia alba] in Tuva; Alyssum sp. in Chukotka (Berman et al., 2002). During two short expeditions in Buryatia this species was never found on Erysimum sp. in the Selenga flood plain or on Smelowskia alba and Alyssum lenense in Barguzin Valley and was common only on Draba nemorosa in two flood-plain ruderal localities at the southern boundary of the taiga zone on the Khilok River in the absence of other crucifers around and in the forest-steppe zone at the Chikoy River, also with a poor flora of Brassicaceae (Korotyaev, 2019) .</p></div>	https://treatment.plazi.org/id/5C0C8799FFEB6A0CC064FDEEFB04D3B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Korotyaev, B. A.	Korotyaev, B. A. (2020): A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae). Zoosystematica Rossica (China) 29 (2): 353-367, DOI: 10.31610/zsr/2020.29.2.353, URL: https://doi.org/10.31610/zsr/2020.29.2.353
5C0C8799FFEA6A0DC3CCFF68FBA5D7AE.text	5C0C8799FFEA6A0DC3CCFF68FBA5D7AE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceutorhynchus klementzorum Korotyaev 1980	<div><p>Ceutorhynchus klementzorum Korotyaev, 1980</p><p>(Figs 4, 14)</p><p>Distribution. Irkutsk Prov., Buryatia, Zabaikalskiy Terr., Mongolia (Korotyaev, 1980, 2019).</p><p>Hosts. Draba nemorosa in ruderal flood-plain localities at the Khilok and Chikoy rivers together with C. seniculus (Korotyaev, 2019) .</p><p>Ceutorhynchus sp. [? C. unguicularis C. G. Thomson, 1871]</p><p>(Figs 5, 9, 15, 21)</p><p>Material. Russia, Central Yakutia, Lena-Vilyui interfluve, km 29 of Vilyui Hwy near Yakutsk, “Suntar” Alas, motley-grass meadow, 28.V.2013, S.N. Nogovitsyna leg., 1 male .</p><p>Distribution. Southern East Siberia: southern Krasnoyarsk Terr., Yermakovskii Distr.; Tuva: southern slope of the West Sayan Mts and East Tuva Plateau (Korotyaev, 1992, 2019), Central Yakutia (new record); Eastern Kazakhstan (Korotyaev, 2019).</p><p>Hosts. Draba nemorosa and D. sibirica in southern Krasnoyarsk Terr. and in Tuva (Korotyaev, 2019); both plants occur also in Central Yakutia (Zakharova et al., 2005) where the small (1.5 mm long) male was collected in the period of their flowering.</p><p>Note. The Siberian form is slightly but usually noticeably different from the European specimens of C. unguicularis in the slightly shorter antennal club, shorter rostrum, shorter, more robust and more strongly rounded at sides elytra with more angular preapical prominences (Korotyaev, 1980: 160 – “a female from Tuva of an uncertain identity”). Specimens of C. unguicularis from Krasnodar Terr. and Adygea have broad-lanceolate white scales sparsely scattered over the elytral disc, which is never found in the specimens from Siberia and Kazakhstan. This form differs more strongly in the bionomics; I did not find it on Turritis borealis in southern Krasnoyarsk Terr. in June 1979 when I went there to collect additional material to the single female brought shortly before that by E. Zemlyakova of the Moscow State University but collected in numbers on Draba nemorosa and D. sibirica belonging to a different tribe ( Alysseae) and dissimilar morphologically, with sharply different fruit structure. I also failed to find C. unguicularis in Apsheronsk Distr. of Krasnodar Terr. at an elevation of about 1200 m a.s.l. and collected this species on Turritis borealis only in the piedmont forest near Kaluzhskaya Vill. in Krasnodar Terr. and at an elevation of about 700 m S of Novoprokhladnoye Vill. in the Republic of Adygea. In Europe (Dieckmann, 1972; Alonso-Zarazaga et al., 2017) C. unguicularis is distributed from Ireland, Sweden and Finland in the north to Italy and Greece in the south and is associated mostly with T. borealis (Dieckmann, 1972; as Arabis hirsuta). At the eastern boundary of Europe in the east of the Russian Plain this species lives mostly on Schivereckia hyperborea of the tribe Alysseae (Isaev, 2007; Dedyukhin, 2016), and less often, on T. borealis (Dedyukhin, 2016) . The long series from Bashkortostan taken by S. V. Dedyukhin from Sch. hyperborea, has no broad-lanceolate white scales on the elytral disc except along suture and behind scutellum.</p></div>	https://treatment.plazi.org/id/5C0C8799FFEA6A0DC3CCFF68FBA5D7AE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Korotyaev, B. A.	Korotyaev, B. A. (2020): A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae). Zoosystematica Rossica (China) 29 (2): 353-367, DOI: 10.31610/zsr/2020.29.2.353, URL: https://doi.org/10.31610/zsr/2020.29.2.353
5C0C8799FFEA6A0DC075FC07FC06D28B.text	5C0C8799FFEA6A0DC075FC07FC06D28B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceutorhynchus querceti (Gyllenhal 1813)	<div><p>Ceutorhynchus querceti (Gyllenhal, 1813)</p><p>(Figs 6, 22)</p><p>Distribution. Holarctic species with the Arctic, boreal and northern subboreal distribution (Colonnelli, 2004).</p><p>Hosts. Mostly on Rorippa palustris throughout most of its Holarctic range (Dieckmann, 1972), also in several localities in Tuva [West Sayan Mts: 80 km N of Teli Vill.; southern foothills of West Sayan, town of Turan; Central Tuvinian depression – Chadan River flood plain, Lake Chagytai, Seserlig (25 km NW of Kyzyl), Kok-Tey east of Kyzyl; Shurmakskii Pass on Eastern Tannu-Ola Mt. Range; Sagly and Erzin villages in southern Tuva; Yrban Vill. on East Tuva Plateau; my collections of 1979 and 1980], but beetles were found in southern Tuva in numbers on Draba nemorosa or D. sibirica and did not occur on R. palustris closer to the creek (Korotyaev, 1992), maybe to escape flooding.</p></div>	https://treatment.plazi.org/id/5C0C8799FFEA6A0DC075FC07FC06D28B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Korotyaev, B. A.	Korotyaev, B. A. (2020): A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae). Zoosystematica Rossica (China) 29 (2): 353-367, DOI: 10.31610/zsr/2020.29.2.353, URL: https://doi.org/10.31610/zsr/2020.29.2.353
5C0C8799FFEA6A0EC075F939FDFFD18F.text	5C0C8799FFEA6A0EC075F939FDFFD18F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceutorhynchus dauricus Korotyaev 1997	<div><p>Ceutorhynchus dauricus Korotyaev, 1997</p><p>(Figs 7, 18)</p><p>Distribution. Zabaikalskiy Terr., Amurskaya</p><p>Prov., Primorskiy Terr., Mongolia, North Korea</p><p>B.A. Korotyaev. A review of Ceutorhynchus associated with woodland draba</p><p>(Korotyaev, 1997), and South Korea (Hong et al., 2001; Korotyaev &amp; Hong, 2004).</p><p>Hosts. On the right bank of the Chikoy River near Zhindo Vill. in Zabaikalskiy Terr. two specimens were swept in herbage with Draba nemorosa in early June 2019 (Korotyaev, 2019). In South Korea, Draba nemorosa var. hebecarpa was recorded as a host (Hong et al., 2001).</p></div>	https://treatment.plazi.org/id/5C0C8799FFEA6A0EC075F939FDFFD18F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Korotyaev, B. A.	Korotyaev, B. A. (2020): A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae). Zoosystematica Rossica (China) 29 (2): 353-367, DOI: 10.31610/zsr/2020.29.2.353, URL: https://doi.org/10.31610/zsr/2020.29.2.353
5C0C8799FFE96A0FC3C3FA3DFD62D3B6.text	5C0C8799FFE96A0FC3C3FA3DFD62D3B6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceutorhynchus cochleariae (Gyllenhal 1813)	<div><p>Ceutorhynchus cochleariae (Gyllenhal, 1813)</p><p>(Figs 8, 10, 23)</p><p>Distribution. Sub-boreal Western Palaearctic species distributed from West Europe to Irkutsk Prov. (Korotyaev, 1980) and northern Tuva (Korotyaev, 2019). Dieckmann (1972) and Colonnelli (2004) report it from Anatolia (Asian part of Turkey) but I have not seen material from Turkey. The record from Georgia in Colonnelli (2004) is based apparently on the only known specimen from the City of Poti on the Black Sea coast in Western Georgia (Korotyaev &amp; Cholokava, 1989; see taxonomic notes below).</p><p>Hosts. Usually Cardamine pratensis (Dieckmann, 1972: 70), but species of several other genera of Brassicaceae were also listed in the cited publication by Dieckman of which E. Colonnelli mentions only Dentaria enneaphyllos and Lunaria rediviva (Colonnelli, 2004) . In Tuva the species was found only on Cardamine pratensis out of the 28 investigated in 1979 and 1980 species of crucifers (Korotyaev, 2019). In Irkutsk and its vicinities (Korotyaev, 2019), I have collected beetles from Draba nemorosa and Barbarea arquata far from water bodies in the habitats where no Cardamine sp. was present.</p><p>Taxonomic notes. The taxonomy of C. cochleariae in the Caucasus needs investigation. This species is apparently absent from the largest part of the steppe zone of European Russia (Arzanov, 2015), but a slightly different form with paler (reddish) legs and denser and finer dorsal vestiture is similar to C. cochleariae but with darker legs with black femora and tibiae and rather bright reddish tarsi; it also has sparser dorsal vestiture, and the beetles look darker than northern C. cochleariae . So, three allopatric forms exist along the northsouth transect from the forest zone of European Russia through the Caucasus foothills to the middle forest belt with Picea orientalis and Abies nordmanniana .</p><p>common in April and early May on Cardamine spp. in the Kuban River flood plain in the Republic of Adygea south of Krasnodar City. The single specimen reported by Korotyaev &amp; Cholokava (1989) from Poti is apparently conspecific with this form. In the piedmont and low-hill Northwestern Caucasus this form has not been collected for many years, and only C. filirostris (Reitter, 1888) occurs (usually in small numbers) on Cardamine spp. and Dentaria spp., but more often on Alliaria petiolata . At higher elevations, usually at about 1300 m a.s.l. and above, C. filirostris is replaced (on Cardamine lazica in Karachai-Cherkess Republic: Korotyaev &amp; Cholokava, 1989) by a form very</p></div>	https://treatment.plazi.org/id/5C0C8799FFE96A0FC3C3FA3DFD62D3B6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Korotyaev, B. A.	Korotyaev, B. A. (2020): A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae). Zoosystematica Rossica (China) 29 (2): 353-367, DOI: 10.31610/zsr/2020.29.2.353, URL: https://doi.org/10.31610/zsr/2020.29.2.353
5C0C8799FFE86A02C075FDCBFD59D609.text	5C0C8799FFE86A02C075FDCBFD59D609.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceutorhynchus unguicularis C. G. Thomson 1871	<div><p>Ceutorhynchus unguicularis species group</p><p>Diagnosis. Body small, 1.5–2.3 mm long, robust. Dorsal vestiture sparse, rather uniform. Disc of pronotum with sparse very narrow, parallel-sided, truncate apically white or greyish scales, with short broad-lanceolate white scales along base, usually also with sparse scales along median line and along sides; intervals of elytra with sparse similar scales; in addition, with wider white scales along suture and at bases of 1st and 2nd intervals. Elytral striae bare. Underside with moderately dense white lanceolate scales. Antennal funicle seven-segmented, fine; club weakly to strongly elongate, two to four times as long as wide, terete, matte, with very dense short, fine pubescence. Rostrum moderately long, slender, levelling with frons at base, about as wide as fore tibia in middle part, cylindrical, moderately strongly, evenly curved dorsoventrally, rather finely, moderately densely punctate in basal half, sparsely and very finely punctate and moderately to strongly shining in apical one-third to one-half, with rath- er ill-defined low, obtuse median carina in basal part. Antennae inserted near rostrum mid-length. Pronotum weakly to moderately transverse, with weak but noticeable lateral tubercles, moderately convex, rather shining, with medium-sized, moderately dense punctures. Elytra with weakly to moderately prominent humeri and weakly to moderately rounded sides and moderately convex disc; preapical prominences weakly to moderately pronounced, with fine granules not conspicuously condensed and forming no oblique ridges. Elytral striae rather wide and deep, intervals moderately shining, as wide as, or not much wider than striae, flat or slightly convex, shining, with rather sparse punctures and minute granules. Legs moderately long, femora mutic, tarsi moderately long and wide, claws short, weakly diverging, with well visnot more than triple as long as wide</p><p>(Figs 9, 27). Elytra often less strongly rounded and with more convex humeri</p><p>(Fig. 5). Aedeagus wider, less rounded and wider apically (Figs 15, 21). Body length 1.5–2.3 mm. South of East Siberia, Central Yakutia ..............</p><p>.......... C. sp. [? C. unguicularis]</p><p>3 (2). Antennal club strongly elongate,</p><p>3.5–4.2 times as long as wide (Fig.</p><p>26). Elytra more strongly rounded,</p><p>with less produced humeri (Fig. 24).</p><p>Aedeagus narrower, more strongly rounded and narrower apically (Fig.</p><p>17). Body length 1.7–2.2 mm. South of Figs 19–23. Ceutorhynchus, aedeagus, dorsal view. 19, C. gemuri- the Russian Far East................ cus, Azerbaijan, Nakhchivan Autonomous Republic; 20, C. kipchak,.............. C. kerzhneri sp. nov. Buryatia, Nikol’sk Vill.; 21, C. sp. [? C. unguicularis], Tuva; 22, 4 (1). Disc of pronotum and elytra with C. querceti; 23, C. cochleariae, Irkutsk. Photo by K.V. Makarov. greyish parallel-sided scales clearly con-</p><p>trasting with broad-lanceolate white scales. Elytra wider, with more promiible appendage at base. Fore tibia of male not mu- nent humeri (Fig. 25). Antennal club moderately cronate, middle and hind tibiae finely mucronate. long, 2.4–2.8 times as long as wide (Fig. 28). Aedea- Middle tibia of female not mucronate. Aedeagus gus longer and narrower, evenly rounded apically weakly sclerotised, with obtuse-angular apex, oc- (Figs 16, 25). Body length 2.2–2.3 mm. South of East</p><p>Siberia and the Russian Far East (only Primorskiy casionally slightly attenuate at tip.</p><p>Terr.), eastern Mongolia ...... C. melniki sp. nov. Species included. The group includes C. mohri Dieckmann, 1960 from Spain, C. nigritulus</p><p>Ceutorhynchus kerzhneri sp.nov. Schultze, 1897 from Central and Southern Eu-</p><p>(Figs 17, 24, 26) rope (Alonso-Zarazaga et al., 2017), C. unguicularis from Europe and Anatolia, with obscure form Ceutorhynchus unguicularis: Korotyaev, 1980: 160, from southern East Siberia and Central Yakutia, pro parte (specimens from Primorskiy Terr.), nec and two new species from the Eastern Palaearctic C.G. Thompson, 1871 (misidentification). described below. Holotype, female: Russia, Primorskiy Terr., 20 km Comparison. Species of the C. unguicularis W of Spassk-Dal’niy, Lake Khanka shore, meadows group differ from species of the C. cochleariae with Salix stand, 12.VI.1989, S.A. Belokobylskij leg. group (Korotyaev, 1980) with black elytra and Paratypes. Russia: Primorskiy Terr., Nakhodka legs first of all in the secondary sexual characters, City, waste land, 21.VIII.1959, I.M. Kerzhner leg., namely the non-mucronate middle tibia of female 1 male; Sakhalin Prov., Kunashir I., Sernovodsk Vill.,</p><p>16.VII.1973, I.M. Kerzhner leg., 1 female. and apically sclerotised and convexly rounded</p><p>Description. The new species is very similar aedeagus, which is always emarginate and poorly</p><p>to C. unguicularis and noticeably differs only in sclerotised medially in C. cochleariae, C. dauricus</p><p>the much longer antennal club (Fig. 26) and much and C. ussuricus Korotyaev, 1997 .</p><p>narrower and more strongly rounded at sides aedeagus (Fig. 17). A key to species of the Ceutorhynchus ungui-</p><p>Rostrum of male 1.71 times as long as pronocularis group from Siberia and the Far East</p><p>tum, that of female, 1.77–1.80 times. 1 (4). Body covered dorsally only with white scales. Antennae of male inserted at 0.48 length of Elytra narrower, with less prominent humeri (Figs rostrum from base, those of female, at 0.46–0.48 5, 24). Antennal club varyingly long. length. Club of female 3.54, of male 4.20 times as 2 (3). Antennal club weakly or moderately elongate, long as wide.</p><p>Pronotum of male 1.50, that of female 1.47– 1.50 times as wide as long.</p><p>Body length of male 1.65, of female 2.00 (Lake Khanka)–2.15 (Kunashir I.) mm.</p><p>Etymology. The species is named after the late I.M. Kerzhner, an outstanding Russian entomologist, who made valuable contribution to the knowledge of the insect fauna of every region he ever investigated.</p><p>Distribution. The south of the Russian Far East (Primorskiy Terr. and Kuril Is. – Kunashir).</p></div>	https://treatment.plazi.org/id/5C0C8799FFE86A02C075FDCBFD59D609	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Korotyaev, B. A.	Korotyaev, B. A. (2020): A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae). Zoosystematica Rossica (China) 29 (2): 353-367, DOI: 10.31610/zsr/2020.29.2.353, URL: https://doi.org/10.31610/zsr/2020.29.2.353
5C0C8799FFE56A03C3C3FDB7FB06D3B7.text	5C0C8799FFE56A03C3C3FDB7FB06D3B7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ceutorhynchus melniki Korotyaev 2020	<div><p>Ceutorhynchus melniki sp. nov.</p><p>(Figs 16, 25, 28)</p><p>Ceutorhynchus unguicularis: Korotyaev, 1980: 160, pro parte (female from Mongolia), nec C.G. Thompson, 1871 (misidentification).</p><p>Holotype, male: Russia, Primorskiy Terr., 20 km E of Khasan, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=130.74834&amp;materialsCitation.latitude=42.41417" title="Search Plazi for locations around (long 130.74834/lat 42.41417)">Golubinyy Utyos Rock</a>, 42°24′51″N, 130°44′54″E, 1–8.VII.2000, I. V. Melnik leg.</p><p>Paratypes. Russia: Primorskiy Terr., 5 km NW of Lake Khanka, 22 km SW of Turii Rog Vill., 2.VII.1974, A.B. Egorov leg., 1 female ; Zabaikalskiy Terr., 30 km NE of Borzya Stn, 6.VII.1989, O.N. Kabakov leg., 1 female . Mongolia, East Aimak, 50 km ENE of Mt. Ikh- Chulut-Ula, old Somon Matad, 22.VI.1976, E.L. Gurjeva leg., 1 female .</p><p>Description. Male. Rostrum 1.39 times as long as pronotum, subcylindrical, slightly widening toward base in basal one-third, regularly and rather strongly curved, as wide as middle part of fore tibia. Base of rostrum levelling with frons. Dorsal surface of rostrum in basal one-third almost matt, densely covered with oblong, rather shallow punctures partly merging in fine striae along obtuse low median carina closer to middle third; latter with finer striolate punctation thinning apically; apical one-third shining, with sparse fine punctures. Antennae inserted at 0.49 length of rostrum from base. Scape weakly widening in apical one-third. Funicle seven-segmented, fine and rather long; 1st segment 2.8 times as long as wide, weakly widening apically; 2nd segment 0.75 times as long as 1st, about triple as long as wide; 3rd segment two-thirds as long as 2nd; 4–6th segments of subequal lengths, noticeably shorter than 3rd segment, more than twice as long as wide; 7th segment distinctly shorter than 6th, about as long as wide. Club terete, almost symmetrical, moderately long, 2.4 times as long as wide, narrow at apex, matt, very densely covered with short fine pubescence. Sutures between segments of club not very conspicuous, only the distal to mid-length onebeing quite distinct. Funicle bearing sparse short, fine, semi-erect hairs. Frons flat, moderately widening posteriorly. Head capsule matt, densely covered with medium-sized, moderately deep polygonal punctures, vertex not carinate. Eyes medium-sized, almost round except for rectilinear anterior margin touching base of rostrum laterally. Dorsal margin of eye situated below frons margin in lateral view.</p><p>Pronotum moderately transverse, 1.35 times as wide as long. Base shallowly bisinuate, weakly angularly protruding posteriorly in the middle. Sides rounded, convexly converging to the moderately deep apical constriction separating moderately long apical part (“collar”) narrowing anteriorly. Lateral tubercles small, obtuse, not protruding from pronotum outline. Disc moderately and rather evenly convex, its sides in anterior half moderately obliquely depressed; median sulcus obsolete in the middle, shallow and wide, rounded at base, narrower at apical constriction. Apical margin of pronotum weakly raised, weakly roundly protruding anteriorly, with shallow emargination in the medial third limited by two weak obtuse angulations; emargination somewhat narrower than frons at posterior margin. Surface of disc almost matt, with very dense small, rather deep punctures separated by narrow but flat, shining in a few places intervals.</p><p>Scutellum very small and narrow, convex, shining. Apices of mesepimera clearly visible dorsally.</p><p>Elytra 1.25 times as long as wide, 1.37 times as wide as pronotum, with moderately convex humeral prominences. Sides subparallel in basal half, without depressions behind humeri, moderately and rather smoothly converging in apical half toward ill-defined, almost smoothened preapical prominences bearing rather sparse, small, acute granules not arranged in crests. Disc moderately and rather evenly convex, slightly flattened in basal half but not depressed behind scutellum. Striae moderately wide and deep; dense, almost round punctures in them slightly excising margins of intervals. The latter about 1.5 times as wide as striae, flat, weakly shining, with irregular fine punctures and small rounded, flattened, low granules.</p><p>Legs rather long and slender; femora mutic, weakly S-curved, weekly widened in middle part. Fore tibia non-mucronate, weakly S-curved, weakly widening along most of length and noticeably shortly outcurved and widened at apex; spines in apical comb very fine and dense. Middle and hind tibiae with sharp short mucro, weakly S-curved, moderately roundly and shortly widened at apex, with slightly longer fine setae in apical comb. Tarsi moderately long and rather narrow; 1st segment about 1.5 times as long as wide, 2nd segment in fore tarsus 1.4 times as long as wide, in middle and hind tarsi slightly shorter; 3rd segment 0.8 times as long and almost twice (1.82 times) as wide as 2nd. Claw-segment weakly widening apically, by 0.6 extending beyond lobes of 3rd segment. Claws short, weakly divergent, with well visible subconnate appendages in basal half. Anal ventrite shallowly depressed in medial third along its entire length, sides of the depression weakly, obtusely convex in apical part. Pygidium almost twice as wide as long, weakly convex, neither sulcate nor carinate, matt, densely rugosely punctate. Aedeagus (Figs 16, 25) much longer and narrower than in C. unguicularis, subparallel-sided, shortly narrowly rounded apically, with sclerotised lateral areas narrow and developed only in apical two-thirds.</p><p>Body black, only antennae and 3rd tarsal segment very dark brown. Rostrum with sparse short narrow, posteriorly-pointed recumbent greyish scales along sides in basal quarter. Head capsule with moderately dense narrow subrecumbent, posteriorly-pointed greyish (along eyes margins) and brownish scales. Pronotum and elytra with moderately dense narrow, parallel-sided or somewhat narrowing apically truncate, subrecumbent (on pronotum) or recumbent (on elytral intervals) light brownish scales. In addition, pronotum with a row of small lanceolate white scales along base and along median line; elytra also with a few such scales at bases of 1st and 2nd intervals and with wider scales in mid-length one-third of lateral interval. Underside with almost uniform vestiture of wider short-lanceolate white scales separated mostly by own widths. Depression on anal ventrite in posterior two-thirds densely covered with white scales, especially at sides, and with fine pale erect setae at posterior margin. Pygidium lacking white scales, with moderately dense, long, fine, subrecumbent yellow and white (at sides) hairs.</p><p>Female. Rostrum 1.37–1.48 times as long as pronotum, strongly and evenly curved. Basal part of rostrum moderately shining, finely punctate; mid-length third of rostrum with low, almost strip-like wide median carina and 2 rows of semi-obliterated small punctures along sides; apical half shining, with sparse fine punctures. Frons feebly convex in cross-section and longitudinally. Antennae inserted at 0.46–0.47 length of rostrum from its base, club 2.56–2.84 times as long as wide. Pronotum 1.36 times as wide as long. All tibiae unarmed. First and second ventrites of abdomen rather strongly jointly convex, 2nd rather steeply sloping to 3rd ventrite. Anal ventrite feebly depressed medially at apex.</p><p>Body length 2.25–2.30 mm.</p><p>Comparison. The new species is similar to C. unguicularis, but differs in the slightly more shining, more sparsely punctate in the apical part rostrum; slightly convex and less strongly widening posteriorly, especially in anterior half, frons; less transverse pronotum with more convex disc and noticeably protruding anterior margin shallowly sinuate medially; slightly wider and more strongly narrowing posteriorly in apical half elytra with more protruding humeral prominences; narrower elytral striae and wider, flat, more densely punctate and less shining intervals; slightly wider 3rd tarsal segment and longer claw-segment; slightly longer and more widely diverging claws with finer, not fused medially and more conspicuous inner appendages. The dorsal vestiture is bicolored, so that beetles look more similar to C. typhae (Herbst, 1795) and especially to the Eastern Palaearctic C. asiaticus Korotyaev, 1997 with its long antennal club but may be immediately distinguished by the seven-segmented dark antennal funicle and appendiculate tarsal claws.</p><p>Etymology. The species is named after I. V. Melnik of Moscow, an enthusiastic amateur coleopterist, who has collected the only male of the new species and has donated to the ZIN collection this specimen together with many other interesting beetles.</p><p>Distribution. South of Transbaikalia (Zabaikalskiy Terr.) and the Russian Far East (Primorskiy Terr.), eastern Mongolia.</p></div>	https://treatment.plazi.org/id/5C0C8799FFE56A03C3C3FDB7FB06D3B7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Korotyaev, B. A.	Korotyaev, B. A. (2020): A review of weevils of the genus Ceutorhynchus (Coleoptera: Curculionidae), associated with woodland draba, Draba nemorosa (Brassicaceae). Zoosystematica Rossica (China) 29 (2): 353-367, DOI: 10.31610/zsr/2020.29.2.353, URL: https://doi.org/10.31610/zsr/2020.29.2.353
