identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
268CA781939D523995CBB01ADC8A9234.text	268CA781939D523995CBB01ADC8A9234.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacopsyche abdita Thomson & Ríos-Touma & Holzenthal 2025	<div><p>Rhyacopsyche abdita sp. nov.</p><p>Fig. 1</p><p>Type material.</p><p>Holotype: Ecuador • ♂; Morona-Santiago, trib. to Rio Abanico, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.20238&amp;materialsCitation.latitude=-2.24985" title="Search Plazi for locations around (long -78.20238/lat -2.24985)">Hwy E 46 (via Riobamba-Macas)</a>; 02.24985°S, 078.20238°W; el. 1531 m; 26 January 2015; Holzenthal, Huisman, Ríos-Touma, Amigo leg.; in alcohol; UMSP 000113995; UMSP . Paratypes: Ecuador • 1 ♂; Pichincha, Reserva Rio Bravo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.73287&amp;materialsCitation.latitude=-0.07802" title="Search Plazi for locations around (long -78.73287/lat -0.07802)">Rio Bravo, upstream</a>; 0.07802°S, 078.73287°W; 19 November 2023; el. 1564 m; Holzenthal, Frandsen, Morabowen, Amigo leg.; pan trap; in alcohol; UMSP 000551404; UMSP • 5 ♂; Imbabura, Reserva Los Cedros, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.78229&amp;materialsCitation.latitude=0.30374" title="Search Plazi for locations around (long -78.78229/lat 0.30374)">tributary to Rio Los Cedros</a>; 00.30374°N, 078.78229°W; el. 1312 m; 18–19 October 2011 Holzenthal, Ríos, Encalada, Acosta leg.; in alcohol; UMSP 000145862; MECN • 7 ♂; same as for preceding; UMSP 000145861; UMSP • 8 ♂; Imbabura, Reserva Los Cedros, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.78084&amp;materialsCitation.latitude=0.32495" title="Search Plazi for locations around (long -78.78084/lat 0.32495)">Rio de la Plata</a>; 00.32495°N, 078.78084°W; el. 1587 m; 18 October 2011; Holzenthal, Ríos, Encalada, Acosta leg.; in alcohol; UMSP 000520782; UMSP • 1 ♂; Imbabura, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.7815&amp;materialsCitation.latitude=0.31127" title="Search Plazi for locations around (long -78.7815/lat 0.31127)">Reserva Los Cedros, small stream near station</a>; 00.31127°N 78.78150°W; el. 1460 m; 17 October 2011; Holzenthal, Ríos, Encalada, Acosta leg.; in alcohol; UMSP 000138550; MECN .</p><p>Diagnosis.</p><p>The form and appearance of the dorsolateral lobe, inferior appendage, and phallus of Rhyacopsyche abdita, new species, and R. intraspira Wasmund &amp; Holzenthal, 2007 are similar. However, the prominent hook at the apex of the intermediate appendage of R. abdita, which is lacking in R. intraspira, can be used to distinguish between the two. Additionally, R. abdita has far fewer of the dark, peg-like setae on the dorsolateral lobe than R. intraspira .</p><p>Description.</p><p>Adult (male: n = 23). Forewing length 3.5–3.8 mm (male). Head unmodified, with 3 ocelli; antennae unmodified. Tibial spurs 1, 3, 4. Specimens brown in alcohol. Sternum VII with short mesoventral point. Male genitalia. Abdominal segment IX setose, anterolateral margin convex, posterolateral margin with mesal hump and bearing small digitate process with apical seta on inner surface; dorsolateral lobe of segment IX truncate and setose, in dorsal view deeply divided and with small peg-like setae on inner surface (Fig. 1 A, B). Tergum X membranous, partially retracted inside segment IX, articulating with intermediate appendage (Fig. 1 A). Intermediate appendage elongate, extending past inferior appendage, apex hook-like (Fig. 1 A, B). Inferior appendage setose, dorsal margin enlarged mesally, ventral margin sinuate; apex bearing prominent, peg-like seta (Fig. 1 A). Phallus basally tubular, elongate, thin membranous sheath surrounding middle portion covered with spicules; central tubule with preapical, geniculate projection; lateral process with darkened, hooked apex (Fig. 1 D, E).</p><p>Etymology.</p><p>Abditus, Latin for “ hidden, concealed, ” referring to the digitate projections on the inner surface of abdominal segment IX.</p><p>Distribution.</p><p>Ecuador (Table 1).</p></div>	https://treatment.plazi.org/id/268CA781939D523995CBB01ADC8A9234	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Thomson, Robin E.;Ríos-Touma, Blanca;Holzenthal, Ralph W.	Thomson, Robin E., Ríos-Touma, Blanca, Holzenthal, Ralph W. (2025): Additions to the genus Rhyacopsyche Müller, 1879 (Trichoptera, Hydroptilidae) in Ecuador. ZooKeys 1263: 531-543, DOI: 10.3897/zookeys.1263.148084
7B28FD0220495C37839F624F619A37A0.text	7B28FD0220495C37839F624F619A37A0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacopsyche bellavista Thomson & Ríos-Touma & Holzenthal 2025	<div><p>Rhyacopsyche bellavista sp. nov.</p><p>Fig. 2</p><p>Type material.</p><p>Holotype: Ecuador • ♂; Pichincha, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.68958&amp;materialsCitation.latitude=-0.01212" title="Search Plazi for locations around (long -78.68958/lat -0.01212)">Bellavista Cloud Forest Reserve and Lodge, small stream</a>; 00.01212°S, 078.68958°W; 13 July 2017; el. 2614 m; Andrea Tapia leg.; pan trap; in alcohol; UMSP 000560013; UMSP . Paratypes: Ecuador • 1 ♂ and 1 ♀; same collection data as for holotype; 15 July 2017; in alcohol; UMSP 000560011; MECN .</p><p>Diagnosis.</p><p>Rhyacopsyche bellavista, new species, is most similar to R. rhamphisa Wasmund &amp; Holzenthal, 2007; both species have a slender inferior appendage that curves dorsad and terminates in a strongly sclerotized apex that points mesally. The inferior appendage curves much more strongly in R. bellavista than it does in R. rhamphisa . The general appearance of tergum X in R. bellavista, large and bearing apical spines and spicules apically, can also be used to diagnose R. bellavista .</p><p>Description.</p><p>Adult (male: n = 2, female: n = 1). Forewing length 3.8–3.9 mm (male); 3.9 mm (female). Head unmodified, with 3 ocelli; antennae unmodified. Tibial spurs 1, 3, 4. Specimens brown in alcohol. Sternum VII with short mesoventral point. Male genitalia. Abdominal segment IX setose, anterolateral margin rounded, posterolateral margin short, straight; dorsolateral lobe of segment IX large, straight, directed slightly dorsad, bearing dark, peg-like setae, in dorsal view deeply and broadly divided, slender (Fig. 2 A, B). Tergum X membranous, partially retracted inside segment IX; apex large, extending past both dorsolateral lobe of segment IX and inferior appendage, with spines and spicules apically; in dorsal view, slender (Fig. 2 A, B). Intermediate appendage not apparent. Inferior appendage broadest basally, simple, elongate, slender, curving strongly dorsad; apex strongly sclerotized, pointed, bent mesad (Fig. 2 A, C). Phallus basally tubular, elongate, apex membranous; central tubule pointed; lateral process not apparent (Fig. 2 D).</p><p>Etymology.</p><p>Named for the reserve where the type specimens were collected.</p><p>Distribution.</p><p>Ecuador (Table 1).</p></div>	https://treatment.plazi.org/id/7B28FD0220495C37839F624F619A37A0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Thomson, Robin E.;Ríos-Touma, Blanca;Holzenthal, Ralph W.	Thomson, Robin E., Ríos-Touma, Blanca, Holzenthal, Ralph W. (2025): Additions to the genus Rhyacopsyche Müller, 1879 (Trichoptera, Hydroptilidae) in Ecuador. ZooKeys 1263: 531-543, DOI: 10.3897/zookeys.1263.148084
391D433950BD580D820E72E858B567ED.text	391D433950BD580D820E72E858B567ED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacopsyche decouxi Thomson & Ríos-Touma & Holzenthal 2025	<div><p>Rhyacopsyche decouxi sp. nov.</p><p>Fig. 3</p><p>Type material.</p><p>Holotype: Ecuador • ♂; Imbabura, Reserva Los Cedros, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.78229&amp;materialsCitation.latitude=0.30374" title="Search Plazi for locations around (long -78.78229/lat 0.30374)">tributary to Rio Los Cedros</a>; 00.30374°N, 078.78229°W; el. 1312 m; 18–19 October 2011; Holzenthal, Ríos, Encalada, Acosta leg.; in alcohol; UMSP 000145864; UMSP . Paratypes: Ecuador • 4 ♂; same as for holotype; in alcohol; UMSP 000145864; MECN • 7 ♂ and 2 ♀; same as for holotype; in alcohol; UMSP 000145868, UMSP 00145869; UMSP • 1 ♂; Azuay, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.50786&amp;materialsCitation.latitude=-2.89407" title="Search Plazi for locations around (long -79.50786/lat -2.89407)">Rio Chaucha &amp; waterfall 1.2 km E La Iberia</a>; 02.89407°S, 079.50786°W; el. 700 m; 22 March 2022; Ríos, Pauls, Amigo leg.; in alcohol; UMSP 000418196; MECN • 1 ♂; Azuay, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.41375&amp;materialsCitation.latitude=-2.90736" title="Search Plazi for locations around (long -79.41375/lat -2.90736)">trib. to Rio Chaucha in San Antonio de Chaucha</a>; 02.90736°S, 079.41375°W; el. 1831 m; 23 March 2022; Holzenthal, Ríos, Amigo leg.; in alcohol; UMSP 000418195; UMSP • 1 ♂; Bolivar, Quebrada San Pablo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.15496&amp;materialsCitation.latitude=-1.37978" title="Search Plazi for locations around (long -79.15496/lat -1.37978)">N of Chazojuan</a>; 01.37978°S, 079.15496°W; el. 1150 m; 20 March 2022; Ríos, Holzenthal, Pauls, Thomson, Amigo leg.; in alcohol; UMSP 000418194; UMSP • 1 ♂; Pichincha, Reserva Rio Bravo, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-78.73287&amp;materialsCitation.latitude=-0.07802" title="Search Plazi for locations around (long -78.73287/lat -0.07802)">Rio Bravo, upstream</a>; 00.07802°S, 078.73287°W; 19 November 2023; el. 1564 m; Holzenthal, Frandsen, Morabowen, Amigo leg.; pan trap; in alcohol; UMSP 000551405; UMSP .</p><p>Diagnosis.</p><p>Rhyacopsyche decouxi, new species, is also similar in appearance to R. rhamphisa . Characteristics found on the dorsolateral lobe, inferior appendage, and phallus make them very similar. Key differences, however, make it possible to separate the two. R. decouxi bears a prominent seta on the dorsal margin of the inferior appendage that is lacking in R. rhamphisa; in addition, the shape of the inferior appendage when viewed ventrally is very different.</p><p>Description.</p><p>Adult (male: n = 15, female: n = 2). Forewing length 2.5–2.9 mm (male), 2.9–3.0 mm (female). Head unmodified, with 3 ocelli; antennae unmodified. Tibial spurs 1, 3, 4. Specimens brown in alcohol. Sternum VII with short mesoventral point. Male genitalia. Abdominal segment IX setose, anterolateral margin rounded, posterolateral margin convex; dorsolateral lobe of segment IX broadly truncate, with dark, peg-like setae, in dorsal view deeply divided and with rounded apex (Fig. 3 A, B). Tergum X membranous, retracted inside segment IX, in dorsal view with lateral lobes (Fig. 3 A, B). Intermediate appendage not apparent. Inferior appendage elongate, setose basally, dorsal margin produced basally and with prominent seta mesally; apex digitate, bearing prominent peg-like seta (Fig. 3 A, C). Phallus basally tubular, elongate, membranous apex bearing series of thickened spines; central tubule with pointed apex adjacent to small, sclerotized plate; lateral process lanceolate (Fig. 3 D, E).</p><p>Etymology.</p><p>Named in honor of the late José Decoux, founder and protector of the reserve, who died in 2024, leaving the enormous legacy of Los Cedros Reserve to future generations.</p><p>Distribution.</p><p>Ecuador (Table 1).</p></div>	https://treatment.plazi.org/id/391D433950BD580D820E72E858B567ED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Thomson, Robin E.;Ríos-Touma, Blanca;Holzenthal, Ralph W.	Thomson, Robin E., Ríos-Touma, Blanca, Holzenthal, Ralph W. (2025): Additions to the genus Rhyacopsyche Müller, 1879 (Trichoptera, Hydroptilidae) in Ecuador. ZooKeys 1263: 531-543, DOI: 10.3897/zookeys.1263.148084
8531400331935BE6886F79C48A44C8BD.text	8531400331935BE6886F79C48A44C8BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacopsyche patula Thomson & Ríos-Touma & Holzenthal 2025	<div><p>Rhyacopsyche patula sp. nov.</p><p>Fig. 4</p><p>Type material.</p><p>Holotype: Ecuador • ♂; Pastaza, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-77.79459&amp;materialsCitation.latitude=-1.92251" title="Search Plazi for locations around (long -77.79459/lat -1.92251)">small stream, ca 3.8 km (rd) SE Chuwitayo</a>; 01.92251°S, 077.79459°W; el. 703 m; 20 September 2021; Ríos, Holzenthal, Frandsen, Errigo, Amigo leg.; in alcohol; UMSP 000280912; MECN .</p><p>Diagnosis.</p><p>Rhyacopsyche patula, new species, is most similar to R. holzenthali Harris &amp; Armitage, 2019 . Both species are lacking the deep division often observed in the posterior margin of the dorsolateral lobe when viewed dorsally, and the general appearance of the inferior appendage is very similar between the two when viewed laterally. However, R. patula is lacking both the small emargination and the peg-like setae present on the dorsolateral lobe of R. holzenthali . In addition, the inferior appendage of R. holzenthali is more slender than that of R. patula .</p><p>Description.</p><p>Adult (male: n = 1). Forewing length 2.9 mm (male). Head unmodified, with 3 ocelli; antennae unmodified. Tibial spurs 1, 3, 4. Specimen brown in alcohol. Sternum VII with short mesoventral point. Male genitalia. Abdominal segment IX setose, anterolateral margin rounded, posterolateral margin irregular and with ventral projection; dorsolateral lobe of segment IX setose, rounded, in dorsal view broad and truncate (Fig. 4 A, B). Tergum X membranous, partially retracted inside segment IX, fused with intermediate appendage (Fig. 4 A, B). Intermediate appendage membranous, fused mesally, with lightly sclerotized sections laterally and ventrally (Fig. 4 A, C). Inferior appendage setose, dorsal margin enlarged mesally, ventral margin sinuate; apex bearing prominent, peg-like seta (Fig. 4 A, C). Phallus basally tubular, elongate, apex membranous; central tubule with slight hooked apex in dorsal view; lateral process pointed (Fig. 4 D).</p><p>Etymology.</p><p>Patulus, Latin for “ spread out, broad, ” referring to the broad dorsolateral lobe of abdominal segment IX, without division when viewed dorsally.</p><p>Distribution.</p><p>Ecuador (Table 1).</p></div>	https://treatment.plazi.org/id/8531400331935BE6886F79C48A44C8BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Thomson, Robin E.;Ríos-Touma, Blanca;Holzenthal, Ralph W.	Thomson, Robin E., Ríos-Touma, Blanca, Holzenthal, Ralph W. (2025): Additions to the genus Rhyacopsyche Müller, 1879 (Trichoptera, Hydroptilidae) in Ecuador. ZooKeys 1263: 531-543, DOI: 10.3897/zookeys.1263.148084
9C7C3CF7E91C56EA8DE5C488B0A0697C.text	9C7C3CF7E91C56EA8DE5C488B0A0697C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhyacopsyche tumida Thomson & Ríos-Touma & Holzenthal 2025	<div><p>Rhyacopsyche tumida sp. nov.</p><p>Fig. 5</p><p>Type material. Holotype: Ecuador • ♂; Azuay, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.59665&amp;materialsCitation.latitude=-2.90989" title="Search Plazi for locations around (long -79.59665/lat -2.90989)">Rio Balao Grande in El Progreso [Sta. Isabel]</a>; 02.90989°S, 079.59665°W; el. 213 m; 21 March 2022; Ríos, Holzenthal, Pauls, Frandsen, Thomson, Amigo leg.; in alcohol; UMSP 000280911; UMSP . Paratypes: Ecuador • 1 ♂; Azuay, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-79.50786&amp;materialsCitation.latitude=-2.89407" title="Search Plazi for locations around (long -79.50786/lat -2.89407)">Rio Chaucha &amp; waterfall, 1.2 km E La Iberia</a>; 02.89407°S, 079.50786°W; el. 700 m; 22 March 2022; Ríos, Pauls, Amigo leg.; in alcohol; UMSP 000418191; UMSP • 1 ♂; same as for preceding; UMSP 000418192; MECN .</p><p>Diagnosis.</p><p>This species most closely resembles R. turrialbe Flint, 1971, based on the overall form and appearance of abdominal segment IX and the inferior appendage. It can be most easily distinguished by the inferior appendage, which curves outward at the apex in R. tumida rather than curling inwards as in R. turrialbe .</p><p>Description.</p><p>Adult (male: n = 3). Forewing length 3.3–3.5 mm (male). Head unmodified, with 3 ocelli; antennae unmodified. Tibial spurs 1, 3, 4. Specimens brown in alcohol. Sternum VII with short mesoventral point. Male genitalia. Abdominal segment IX setose, anterolateral margin sinuate, posterolateral margin irregular with ventral projection; dorsolateral lobe of segment IX rounded with dark, peg-like setae, in dorsal view divided and rounded (Fig. 5 A, B). Tergum X membranous, retracted inside segment IX, with basal projections, articulating with intermediate appendage (Fig. 5 A, B). Intermediate appendage elongate, extending posteriad and bent dorsad, in lateral view apex curving slightly outward (Fig. 5 A, B). Inferior appendage wide, setose, divided into dorsal and ventral lobes; dorsal lobe with posterior margin produced into blunt process, dorsal margin bearing large, bulbous, peg-like seta; ventral lobe simple, broadly rounded (Fig. 5 A). Phallus basally tubular, elongate, apex membranous; central tubule with hooked apex; lateral process pointed (Fig. 5 D).</p><p>Etymology.</p><p>Tumidus, Latin for “ swollen, ” referring to the shape of the thick seta on the dorsal lobe of the inferior appendage.</p><p>Distribution.</p><p>Ecuador (Table 1).</p></div>	https://treatment.plazi.org/id/9C7C3CF7E91C56EA8DE5C488B0A0697C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Thomson, Robin E.;Ríos-Touma, Blanca;Holzenthal, Ralph W.	Thomson, Robin E., Ríos-Touma, Blanca, Holzenthal, Ralph W. (2025): Additions to the genus Rhyacopsyche Müller, 1879 (Trichoptera, Hydroptilidae) in Ecuador. ZooKeys 1263: 531-543, DOI: 10.3897/zookeys.1263.148084
