taxonID	type	description	language	source
5E6687C49828FFA78A8EAE01A7E749E2.taxon	diagnosis	Species diagnosis: length 25 − 120 mm, number of segments 101 – 176. Prostomium epilobic. Clitellum from xxvi−xxxiii, tubercula pubertatis from xxix−xxxi; the spermathecal pores are paired in 9 / 10 and 10 / 11 mid-dorsal; body color purple dorsally and faint yellow ventrally.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982FFFA18A8EA9BDA4F34CE0.taxon	materials_examined	Material examined. Holotype: 1 clitellate (471 R 30 _ 01), municipal park (46.6271 ° N, 131.1461 ° E, 141.5 m elev.), Beixiu Park, Shuangyashan City, Heilongjiang Province, 2023 - 07 - 15, coll. Yufeng Zhang. Paratypes: 2 clitellates (464 R _ 01, 04), Qianjin Town (49.1039 ° N, 125.2375 ° E, 281.0 m elev.), Nenjiang County, Heihe City, Heilongjiang Province, 2021 - 08 - 04; 4 clitellates (471 R 30 _ 03, 04, 05, 07), same data as of holotypes; 3 clitellates (507 N _ 01, 03, 507 R 13 _ 05), municipal park (42.3893 ° N, 126.8258 ° E, 575.2 m elev.), Liuhaishan Park, Jingyu County, Tonghua City, Jilin Province, 2023 - 07 - 23, coll. Huifeng Zhao.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982FFFA18A8EA9BDA4F34CE0.taxon	etymology	Etymology. This subspecies is named after the Chinese adjective ‘ dark purple’ translated into English, “ shenzi ” that describes its coloration.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982FFFA18A8EA9BDA4F34CE0.taxon	diagnosis	Diagnosis. Pigmentation of the entire dorsal side is dark-purple except the clitellum (xxv, xxvi – xxxiii), which is gray, ventro-anterior to xv is purple,> xv is pale in alcohol. Body is small-medium size, length 38 – 81 mm diameter 5.32 – 6.5 mm, segments 78 – 156. Prostomium open epilobic. Dorsal pores from 4 / 5. Spermathecal pores paired in 9 / 10 / 11 close to mid-D. Female pores in xiv and male pores in xv lateral. Clitellum within xxv, xxvi – xxxiii, Tubercula pubertatis within ½ xxviii, xxix – xxxi. Spermathecae has two pairs within ix – xi, the shape is spherical near the dorsal midline.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982FFFA18A8EA9BDA4F34CE0.taxon	description	Description: Pigmentation of the entire dorsal side is dark-purple except the clitellum (xxv, xxvi – xxxiii), which is gray, ventro-anterior to xv is purple,> xv is pale in alcohol. Length 38 – 81 mm (n = 10). Diameter 5.32 – 6.5 mm. Segments 78 – 156. Prostomium open epilobic. Dorsal pores from 4 / 5. Setae closely paired. Setal ratio on segment viii in adult specimens: aa: 1.74 – 2.20; ab: 0.23 – 0.39; bc: 1.40 – 1.70; cd: 0.21 – 0.36; dd: 5.22 – 7.00; U (circumference): 7.85 – 13.66; mean: aa: ab: bc: cd: dd = 1.95: 0.34: 1.54: 0.27: 6.30; U: 10.73. Setal ratio on segment xxviii in adult specimens: aa: 2.74 – 3.80; ab: 0.23 – 0.43; bc: 1.17 – 1.53; cd: 0.20 – 0.26; dd: 5.16 – 5.91; U (circumference): 13.19 – 16.64; mean: aa: ab: bc: cd: dd = 3.23: 0.33: 1.39: 0.24: 5.50; U: 15.33. Setal ratio on segment xli in adult specimens: aa: 2.13 – 2.79; ab: 0.24 – 0.36; bc: 1.57 – 1.94; cd: 0.19 – 0.29; dd: 4.71 – 6.00; U (circumference): 12.81 – 14.85; mean: aa: ab: bc: cd: dd = 2.46: 0.29: 1.77: 0.25: 5.31; U: 13.65. Nephropores alternate irregularly between setae b and above d. Spermathecal pores paired in 9 / 10 / 11 close to mid-D. Distance between spermathecal pores 0.037 (average of ten samples). Female pores in xiv lateral of b. Distance between female pores 0.195 (average of ten samples). Male pores in xv lateral of b just wider than female pores. Distance between male pores 0.240 (average of ten samples). One specimen has genital markings on the ventral side: a pair in vii, a pair in viii, one genital marking on the left of x and xi, a pair in xii Lateral: a pair of genital markings in viii – x; one genital marking on the right of xi. Clitellum saddle-shaped; in xxv – xxxiii (N = 4 specimens) or xxvi – xxxiii (N = 6). Tubercula pubertatis in ½ xxviii – xxxi (N = 2) or xxix – xxxi (N = 8). Internally, septa 7 / 8 – 10 / 11 thickened. Spermathecae round, some specimens spermathecae are oval-shaped in ix, white; spermathecal duct short, thin in ix and x. Testis and funnels in x and xi seemingly free. Hearts in vii – xi. Calciferous glands xi and xii. Seminal vesicles four pairs in ix – xii (biggest in xii). Extraoesophageal vessels, the testis sacs and epididymis are absent. Ovaries compact in xiii. Crop in xv – xvi and muscular gizzard in xvii – xviii, some specimens in xvii – xxix. Spermathecae two pairs, 1 pair in each of 9 / 10 and 10 / 11, spermathecal shape spherical positioned near the dorsal midline. Typhlosole is inverted T-shaped from about xxvii.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982FFFA18A8EA9BDA4F34CE0.taxon	discussion	Remarks. Eisenia species that are found in the Palearctic biogeographic region include Eisenia sibirica, E. lagodechiensis, E. nana, E. ventripapillata, E. atlavinyteae, E. angusta, E. balatonica, E. fetida, E. tracta, and E. nordenskioldi. Among these, Eisenia nordenskioldi has a thicker body (3.57 – 6.5 mm) than E. nana (2.5 – 3 mm), E. angusta (2.5 – 3 mm) and E. balatonica (1.5 – 2.5 mm) diameter. The first dorsal pore in E. nordenskioldi is located at 4 / 5, a characteristic shared with E. atlavinitae, E. angusta, and E. Balatonica (Table 4). This positioning differs from E. fetida, which exhibits the first dorsal pore at both 3 / 4 and 5 / 6. The clitellum of E. nordenskioldi starts at segment xxiv, which differs from the starting position of the clitellum in E. sibirica (xxvii – xxxii), E. lagodechiensis (xxv – xxxiv), E. nana (xxvii – xxxiii), E. ventripapillata (xxvi – xxxiii), E. atlavinitae (xxvi – xxxiii), E. tracta (xxvi – xxxiii) and E. angusta (xxvii – ½ xxxiii). Additionally, the clitellum of E. nordenskioldi terminates at segment xxxiv, which is distinct from the ending position of the clitellum observed in E. balatonica (xxiv – xxx) and E. fetida (xxiv – xxxiii). Tubercles in E. nordenskioldi starts in segments xxvii, unlike any other Eisenia species. The distribution of papillae in E. nordenskioldi (ab vii – xiii, xvi – xviii, xxi – xxxv, cd viii – xiii) is more extensive compared to the other species like E. sibirica (ab viii, xxviii – xxxi, cd ix – xi), E. lagodechiensis (ab xxv – xxxviii, cd ix – xi), E. nana (ab viii – xii, cd x – xi), and E. ventripapillata (ab x – xi, xxvi – xxxii, cd x – xi). The calciferous glands in E. nordenskioldi are located on segments xi – xii without diverticula, similar to E. angusta and E. atlavinitae, but differ from E. balatonica and E. fetida. E. balatonica, which have diverticula on segments xi – xii. Eisenia fetida has its calciferous glands on x – xii without diverticula. Among the subspecies, E. n. shenzi ssp. nov. overlaps in size with E. n. nordenskioldi, E. n. polypapillata, E. n. jilinensis, and E. n. chinensis but is relatively shorter than E. manshurica, E. n. mongol and E. n. onon (38 – 81 mm vs. 80 – 144 mm) (Table 5). The new subspecies has a thicker body (5.32 – 6.5) compared with E. n. jilinensis and E. n. chinensis (3.57 – 5.1) (Table 5). The entire dorsal side of E. n. shenzi is dark-purple except the clitellum, which is gray, whereas, the ventro-anterior side up to segment xv is purple, of which after, the rest is pale (Fig. 1). On the other hand, the dorso-anterior side of E. n. jilinensis to segment xv is pinkish and gradually changes to pale throughout the rest of the ventral side. Meanwhile, both the entire dorsal and ventral sides of E. n. chinensis have pale pigmentation. The clitellum of E. n. shenzi starts from segment xxv while the clitellum of the rest of the consubspecifics start from either xxiv, ½ xxv, ½ xxvi, or xxvi. Similar to E. n. polypapillata and some specimens of E. n. chinensis, the TP starts at ½ of segment xxviii. The TP of the other subspecies begins at segment xxvii, xxviii or xxix. The setae ratio in segment viii, xxviii and xli in E. n. shenzi are significantly wider than that of E. n. jilinensis and E. n. chinensis. In addition, specimens of E. n. shenzi have the gizzard from xvii to xix, but the other two subspecies from xvii – xviii.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFA38A8EA9BDA6B64827.taxon	materials_examined	Material examined. Holotype: 1 clitellate (504 R 2 _ 03), Taiwang Town (41.1600 ° N, 126.2500 ° E, 41.1 m elev.), Ji’an County, Jilin Province, 2023 - 07 - 25, coll. Huifeng Zhao. Paratypes: 6 clitellates (504 R 2 _ 01, 02, 04, 05, 07, 08), same data as of holotypes.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFA38A8EA9BDA6B64827.taxon	etymology	Etymology. The subspecies is named after the Province of Jilin in Northeast China, the type locality.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFA38A8EA9BDA6B64827.taxon	diagnosis	Diagnosis. Body is small size, length 40 – 49 mm diameter 3.57 – 4.11 mm, segments 98 – 127. Prostomium open epilobic. Dorsal pores from 4 / 5. Spermathecal pores paired in 9 / 10 / 11 close to mid-D. Female pores in xiv and male pores in xv. Clitellum within ½ xxvi, xxvii – xxxiii, Tubercula pubertatis within xxviii, xxix – xxxi, ½ xxxii. No genital marking. Spermathecae has two pairs within ix – xi, the shape is spherical near the dorsal midline. Crop in xv – xvi, gizzard in xvii – xviii.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFA38A8EA9BDA6B64827.taxon	description	Description: Pigment dorso-anterior to xv pinkish,> xv gradually paling ventral pale in alcohol. Length 40 – 49 mm (n = 7). Diameter 3.57 – 4.11 mm. Segments 98 – 127. Prostomium open epilobic. Dorsal pores from 4 / 5. Setae closely paired. Setal ratio on segment viii in adult specimens: aa: 0.85 – 1.38; ab: 0.17 – 0.23; bc: 0.73 – 1.12; cd: 0.16 – 0.23; dd: 3.36 – 4.40; U (circumference): 7.32 – 9.39; mean: aa: ab: bc: cd: dd = 1.20: 0.19: 0.92: 0.19: 3.98; U: 8.57. Setal ratio on segment xxviii in adult specimens: aa: 1.52 – 2.12; ab: 0.21 – 0.30; bc: 0.92 – 1.54; cd: 0.13 – 0.20; dd: 3.36 – 4.52; U (circumference): 8.67 – 11.18; mean: aa: ab: bc: cd: dd = 1.96: 0.24: 1.29: 0.17: 4.02; U: 10.00. Setal ratio on segment xli in adult specimens: aa: 1.42 – 1.99; ab: 0.20 – 0.32; bc: 1.09 – 1.75; cd: 0.12 – 0.19; dd: 3.69 – 4.20; U (circumference): 10.11 – 11.02; mean: aa: ab: bc: cd: dd = 1.82: 0.26: 1.22: 0.16: 3.97; U: 10.68. Nephropores alternate irregularly between setae b and above d. Spermathecal pores paired in 9 / 10 / 11 close to mid-D. Distance between the spermathecal pores 0.035 (average of seven samples). Female pores in xiv lateral of b. Distance between female pores 0.192 (average of seven samples). Male pores in xv lateral of b just wider than female pores. Distance between male pores 0.251 (average of seven samples). No genital marking. Clitellum saddle-shaped; in ½ xxvi – xxxiii (n = 1), xxvi – xxxii (n = 1) or xxvii – xxxiii (n = 5). Tubercula pubertatis in xxviii – xxxi (n = 1), xxix – ½ xxxii (n = 2) or xxix – xxxi (n = 4). Internally, septa 8 / 9 – 11 / 12 thickened. Spermathecae round, white; spermathecal duct short, thin in ix and x. Testis and funnels in x and xi seemingly free. Hearts in vii – xi. Calciferous glands in xi and xii. Seminal vesicles four pairs in ix – xii (biggest in xii). Extraoesophageal vessels, the testis sacs and epididymis are absent. Ovaries compact in xiii. Crop large in xv – xvi and muscular gizzard in xvii – xviii. Inverted T-shaped typhlosole present.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFA38A8EA9BDA6B64827.taxon	discussion	Remarks. Eisenia nordenskioldi jilinensis overlaps in size with E. n. nordenskioldi, E. n. shenzi and E. n. chinensis but is shorter than E. n. manshurica, E. n. mongol, E. n. polypapillata and E. n. onon (40 – 49 mm vs. 55 – 144 mm) (Table 5). Similar to E. n. chinensis, E. n. jilinensis has a thinner body compared with E. n. shenzi (3.57 – 4.11 mm vs. 5.32 – 6.5 mm). The dorso-anterior side of E. n. jilinensis to segment xv is pinkish and gradually changes to pale throughout the rest of the ventral side while the entire dorsal side of E. n. shenzi is dark-purple except the clitellum, which is gray, whereas, the ventro-anterior side up to segment xv is purple, of which after, the rest is pale (Fig. 1). Meanwhile, both the entire dorsal and ventral sides of E. n. chinensis have pale pigmentation. Similar to some of the specimens of E. n. chinensis, the clitellum of E. n. jilinensis begins at ½ of segment xxvi. Whereas, the clitellum of the other subspecies begins at xxiv, ½ xxv, xxv or xxvi. The setae ratio in segment viii, xxviii and xli in E. n. jilinensis are a little narrower than E. n. chinensis.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFAC8A8EAF71A7674B54.taxon	materials_examined	Material examined. Holotype: 1 clitellate (374 R 1 _ 02), municipal park (43.8570 ° N, 125.3058 ° E, 235.0 m elev.), Nanhu Park, Changchun City, Jinlin Province, 2022 - 07 - 06, coll. Yufeng Zhang. Paratypes: 8 clitellates (374 R 1 _ 07, 08, 09, 374 R 3 _ 14, 16, 19, 29, 374 R 4 _ 01), same data as of holotypes; 1 clitellate (418 N _ 05), Shihe Town (46.0619 ° N, 127.5649 ° E, 132.0 m elev.), Mulan County, Harbin City, Heilongjiang Province, 2021 - 08 - 05, coll. Yufeng Zhang.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFAC8A8EAF71A7674B54.taxon	etymology	Etymology. The subspecies is named after the country of China.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFAC8A8EAF71A7674B54.taxon	diagnosis	Diagnosis. unpigmented dorsally and ventrally in alcohol. Body size small with length 30 – 50 mm, diameter 3.9 – 5.1 mm, segments 92 – 127. Prostomium open epilobic. Dorsal pores from 4 / 5. Spermathecal pores paired in 9 / 10 / 11 close to mid-D. Female pores in xiv and male pores in xv. Clitellum within ½ xxvi, xxvi, xxvii – xxxii, xxxiii, Tubercula pubertatis within ½ xxviii, xxviii, xxix – xxxi, ½ xxxii. Genital markings are random. Spermathecae has two pairs within ix – xi, the shape is spherical near the midline of the dorsal. Crop occupies two segments in xv – xvi, gizzard in xvii – xviii.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFAC8A8EAF71A7674B54.taxon	description	Description: Unpigmented dorsally and ventrally in alcohol. Length 30 – 50 mm (n = 10). Diameter 3.9 – 5.1 mm. Segments 92 – 127. Prostomium open epilobic. Dorsal pores from 4 / 5. Setae closely paired. Setal ratio on segment viii in adult specimens: aa: 1.20 – 1.76; ab: 0.20 – 0.26; bc: 0.83 – 1.37; cd: 0.14 – 0.27; dd: 3.75 – 4.47; U (circumference): 8.60 – 10.11; mean: aa: ab: bc: cd: dd = 1.48: 0.22: 1.13: 0.21: 4.18; U: 8.95. Setal ratio on segment xxviii in adult specimens: aa: 2.05 – 2.67; ab: 0.26 – 0.37; bc: 1.02 – 1.57; cd: 0.15 – 0.23; dd: 3.74 – 5.99; U (circumference): 10.14 – 14.04; mean: aa: ab: bc: cd: dd = 2.33: 0.32: 1.32: 0.20: 4.25; U: 12.32. Setal ratio on segment xli in adult specimens: aa: 1.72 – 2.52; ab: 0.20 – 0.31; bc: 1.27 – 1.68; cd: 0.18 – 0.24; dd: 3.39 – 4.04; U (circumference): 9.36 – 12.56; mean: aa: ab: bc: cd: dd = 2.00: 0.26: 1.47: 0.22: 3.68; U: 10.74. Nephropores alternate irregularly between setae b and above d. Spermathecal pores paired in 9 / 10 / 11 close to mid-D. Distance between spermathecal pores 0.039 (average of ten samples). Female pores in xiv lateral of b. Distance between female pores 0.188 (average of ten samples). Male pores in xv lateral of b just wider than female pores. Distance between male pores 0.246 (average of ten samples). Genital markings on the ventral side: One specimen has a pair in segment xii, with one genital marking on the left and right sides of segments xi and xii. Another specimen has a pair in segment viii and a pair in segment xii, with left genital markings in segments x to xii and a single right genital marking in segments ix to xii. A third specimen has a pair in segment viii and a pair in segment xii, with segments xi and xiii each having one. The left genital markings are found in segments x to xiii, while the right genital marking is in segments x to xii. A fourth specimen shows a pair in segment viii, with segment xii having one genital marking. The lateral genital markings are present on the left and right sides of segments x to xii. A fifth specimen has a pair in segment xii, with lateral genital markings on the left and right sides of segments x to xii. Clitellum saddle-shaped; in ½ xxvi – xxxii (n = 1), xxvi – xxxii (n = 1), xxvi – xxxiii (n = 2) or xxvii – xxxiii (n = 6). Tubercula pubertatis in xxviii – xxxi (n = 1), ½ xxviii – ½ xxxii (n = 2), in xxix – ½ xxxii (n = 2) or xxix – xxxi (n = 5). Internally, nephridial bladders simple, sausage-shaped (in all segments inspected). Septa 7 / 8 strongly strengthened, 9 / 10 / 11 thickened. Spermathecae round, white; spermathecal duct short, thin in ix and x. Testis and funnels in x and xi seemingly free. Hearts in vii – xi. Calciferous glands in xi and xii. Seminal vesicles four pairs in ix – xii (biggest in xii). Extraoesophageal vessels, the testis sacs and epididymis are also absent. Ovaries compact in xiii. Crop large in xv – xvi and muscular gizzard in xvii – xviii. Spermathecae has two pairs, 1 pair each in 9 / 10 and 10 / 11, near the midline of the dorsal. Typhlosole is inverted T-shaped from about xxvii.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
5E6687C4982CFFAC8A8EAF71A7674B54.taxon	discussion	Remarks. Eisenia nordenskioldi chinensis ssp. nov. is more morphologically similar to E. n. jilinensis (Table 5). The difference between the two subspecies are the ratio of their setal distance (as mentioned in the Remarks section of E. n. jilinensis) and their pigmentation pattern wherein the dorso-anterior side of E. n. jilinensis to segment xv is pinkish and gradually turns pale to the rest of the body while both the entire dorsal and ventral sides of E. n. chinensis have pale pigmentation.	en	Qin, Mingyan, Zhao, Huifeng, Zhang, Yufeng, Han, Anne Charis N., Shekhovtsov, Sergei V., Wu, Donghui, Aspe, Nonillon M. (2025): Description of three new subspecies of Eisenia nordenskioldi (Annelida: Crassiclitellata) from Northeastern China. Zootaxa 5642 (3): 225-244, DOI: 10.11646/zootaxa.5642.3.2, URL: https://doi.org/10.11646/zootaxa.5642.3.2
